Color or colour is the visual perception based on the electromagnetic spectrum. Though color is not an inherent property of matter, color perception is related to an object's light absorption, reflection, emission spectra and interference. For most humans, colors are perceived in the visible light spectrum with three types of cone cells (trichromacy). Other animals may have a different number of cone cell types or have eyes sensitive to different wavelength, such as bees that can distinguish ultraviolet, and thus have a different color sensitivity range. Animal perception of color originates from different light wavelength or spectral sensitivity in cone cell types, which is then processed by the brain.
The retina is the innermost, light-sensitive layer of tissue of the eye of most vertebrates and some molluscs. The optics of the eye create a focused two-dimensional image of the visual world on the retina, which then processes that image within the retina and sends nerve impulses along the optic nerve to the visual cortex to create visual perception. The retina serves a function which is in many ways analogous to that of the film or image sensor in a camera.
A luminous efficiency function or luminosity function represents the average spectral sensitivity of human visual perception of light. It is based on subjective judgements of which of a pair of different-colored lights is brighter, to describe relative sensitivity to light of different wavelengths. It is not an absolute reference to any particular individual, but is a standard observer representation of visual sensitivity of theoretical human eye. It is valuable as a baseline for experimental purposes, and in colorimetry. Different luminous efficiency functions apply under different lighting conditions, varying from photopic in brightly lit conditions through mesopic to scotopic under low lighting conditions. When not specified, the luminous efficiency function generally refers to the photopic luminous efficiency function.
A photoreceptor cell is a specialized type of neuroepithelial cell found in the retina that is capable of visual phototransduction. The great biological importance of photoreceptors is that they convert light into signals that can stimulate biological processes. To be more specific, photoreceptor proteins in the cell absorb photons, triggering a change in the cell's membrane potential.
Photometry is a branch of science that deals with the measurement of light in terms of its perceived brightness to the human eye. It is concerned with quantifying the amount of light that is emitted, transmitted, or received by an object or a system.
Tetrachromacy is the condition of possessing four independent channels for conveying color information, or possessing four types of cone cell in the eye. Organisms with tetrachromacy are called tetrachromats.
Rod cells are photoreceptor cells in the retina of the eye that can function in lower light better than the other type of visual photoreceptor, cone cells. Rods are usually found concentrated at the outer edges of the retina and are used in peripheral vision. On average, there are approximately 92 million rod cells in the human retina. Rod cells are more sensitive than cone cells and are almost entirely responsible for night vision. However, rods have little role in color vision, which is the main reason why colors are much less apparent in dim light.
Cone cells or cones are photoreceptor cells in the retinas of vertebrates' eyes. They respond differently to light of different wavelengths, and the combination of their responses is responsible for color vision. Cones function best in relatively bright light, called the photopic region, as opposed to rod cells, which work better in dim light, or the scotopic region. Cone cells are densely packed in the fovea centralis, a 0.3 mm diameter rod-free area with very thin, densely packed cones which quickly reduce in number towards the periphery of the retina. Conversely, they are absent from the optic disc, contributing to the blind spot. There are about six to seven million cones in a human eye, with the highest concentration being towards the macula.
In visual physiology, adaptation is the ability of the retina of the eye to adjust to various levels of light. Natural night vision, or scotopic vision, is the ability to see under low-light conditions. In humans, rod cells are exclusively responsible for night vision as cone cells are only able to function at higher illumination levels. Night vision is of lower quality than day vision because it is limited in resolution and colors cannot be discerned; only shades of gray are seen. In order for humans to transition from day to night vision they must undergo a dark adaptation period of up to two hours in which each eye adjusts from a high to a low luminescence "setting", increasing sensitivity hugely, by many orders of magnitude. This adaptation period is different between rod and cone cells and results from the regeneration of photopigments to increase retinal sensitivity. Light adaptation, in contrast, works very quickly, within seconds.
As a part of the retina, bipolar cells exist between photoreceptors and ganglion cells. They act, directly or indirectly, to transmit signals from the photoreceptors to the ganglion cells.
The Purkinje effect or Purkinje phenomenon is the tendency for the peak luminance sensitivity of the eye to shift toward the blue end of the color spectrum at low illumination levels as part of dark adaptation. In consequence, reds will appear darker relative to other colors as light levels decrease. The effect is named after the Czech anatomist Jan Evangelista Purkyně. While the effect is often described from the perspective of the human eye, it is well established in a number of animals under the same name to describe the general shifting of spectral sensitivity due to pooling of rod and cone output signals as a part of dark/light adaptation.
Photopic vision is the vision of the eye under well-lit conditions (luminance levels from 10 to 108 cd/m2). In humans and many other animals, photopic vision allows color perception, mediated by cone cells, and a significantly higher visual acuity and temporal resolution than available with scotopic vision.
Visual phototransduction is the sensory transduction process of the visual system by which light is detected by photoreceptor cells in the vertebrate retina. A photon is absorbed by a retinal chromophore, which initiates a signal cascade through several intermediate cells, then through the retinal ganglion cells (RGCs) comprising the optic nerve.
In the anatomy of the eye, amacrine cells are interneurons in the retina. They are named from Greek a– 'non', makr– 'long', and in– 'fiber', because of their short neuronal processes. Amacrine cells are inhibitory neurons, and they project their dendritic arbors onto the inner plexiform layer (IPL), they interact with retinal ganglion cells, and bipolar cells or both of these.
Intrinsically photosensitive retinal ganglion cells (ipRGCs), also called photosensitive retinal ganglion cells (pRGC), or melanopsin-containing retinal ganglion cells (mRGCs), are a type of neuron in the retina of the mammalian eye. The presence of something like ipRGCs was first suspected in 1927 when rodless, coneless mice still responded to a light stimulus through pupil constriction; this implied that rods and cones are not the only light-sensitive neurons in the retina. Yet research on these cells did not advance until the 1980s. Recent research has shown that these retinal ganglion cells, unlike other retinal ganglion cells, are intrinsically photosensitive due to the presence of melanopsin, a light-sensitive protein. Therefore, they constitute a third class of photoreceptors, in addition to rod and cone cells.
A duplex retina is a retina consisting of both rod cells and cone cells, which are the photoreceptor cells for two parallel but mostly separate visual systems. The rods enable the scotopic visual system, which is active in dim light. The cones enable the photopic visual system, which is active in bright light. While one is active, the other is generally inactive; either the rods are photobleached, or oversaturated, in bright light, or the cones are not sensitive enough to hyperpolarize, or instigate the phototrasduction cascade, in dim light. However, at mesopic (twilight) conditions, both visual systems are active. In this region of overlap, both systems are active and combine to contribute to mesopic vision.
Mesopic vision, sometimes also called twilight vision, is a combination of photopic and scotopic vision under low-light conditions. Mesopic levels range approximately from 0.01 to 3.0 cd/m2 in luminance. Most nighttime outdoor and street lighting conditions are in the mesopic range.
The Stiles–Crawford effect is a property of the human eye that refers to the directional sensitivity of the cone photoreceptors.
Spectral sensitivity is the relative efficiency of detection, of light or other signal, as a function of the frequency or wavelength of the signal.
Visual perception is the ability to interpret the surrounding environment through photopic vision, color vision, scotopic vision, and mesopic vision, using light in the visible spectrum reflected by objects in the environment. This is different from visual acuity, which refers to how clearly a person sees. A person can have problems with visual perceptual processing even if they have 20/20 vision.
