Turkmenian kulan

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Turkmenian kulan
Equus hemionus kulan.JPG
A Turkmenian kulan at Wildpark Pforzheim.
CITES Appendix II (CITES) [2]
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Perissodactyla
Family: Equidae
Genus: Equus
Species:
Subspecies:
E. h. kulan
Trinomial name
Equus hemionus kulan
Groves & Mazák, 1967
Synonyms

Equus hemionus finschi
(Matschie, 1911)

The Turkmenian kulan (Equus hemionus kulan), also called Transcaspian wild ass, [3] Turkmenistani onager or simply the kulan, is a subspecies of onager (Asiatic wild ass) native to Central Asia. It was declared Endangered in 2016. [1]

Contents

The species's population had recently been in decline in the country while it slowly increases in reintroduction sites. The Turkmenian kulan has been reintroduced to Kazakhstan and Uzbekistan, as well as Israel, where the subspecies are hybridizing with Persian onagers in the wild.

Previously in 2005, the population was estimated at 1,295-1,345 in Turkmenistan. No other data existed on the condition of the Turkmenistan populations, but hope remained that small groups of animals still resided in inaccessible areas around Badkhyz, and were thriving in the West Kopetdagh (Sumbar-Chandyr Valley) and Ustyurt Plateau around Lake Sarakamish. [4] However, certain fragmented populations of Transcaspian wild ass are currently on the rise to even more than 2,000 individuals in the wild. [5] It is also estimated that over 6,000 kulans live in Central Asia. [6] In 2017, there are 3,900 kulans in total roaming in Kazakhstan, of which the largest Kazakh population (3,400) reside in Altyn-Emel National Park. [7]

Physical description

A Turkmenian kulan at Korkeasaari Zoo. Kulaani Korkeasaari.jpg
A Turkmenian kulan at Korkeasaari Zoo.

The Turkmenian kulan is one of the largest subspecies of the onagers. It is 200–250 cm long, 100–140 cm tall at the shoulder, and it weighs 200–240 kg. Male onagers are larger than the females.

The Transcaspian wild ass is characterized by a pale brown coat, a dark stripe down the spine and white patches on the sides, back and belly. It also has shaggy black mane and a tuft at the tail end. During the summer, the Turkmenian kulan's coat is leaner and brown, then it turns into thick grayish-brown winter coat during the cold season.

Evolution

Skull of a giant extinct horse, Equus eisenmannae Equus eisenmannae.JPG
Skull of a giant extinct horse, Equus eisenmannae

The genus Equus, which includes all extant equines, is believed to have evolved from Dinohippus via the intermediate form Plesippus . One of the oldest species is Equus simplicidens , described as zebra-like with a donkey-shaped head. The oldest fossil to date is ~3.5 million years old from Idaho, USA. The genus appears to have spread quickly into the Old World, with the similarly aged Equus livenzovensis documented from western Europe and Russia. [8]

Molecular phylogenies indicate the most recent common ancestor of all modern equids (members of the genus Equus) lived ~5.6 (3.9–7.8) mya. Direct paleogenomic sequencing of a 700,000-year-old middle Pleistocene horse metapodial bone from Canada implies a more recent 4.07 Myr before present date for the most recent common ancestor (MRCA) within the range of 4.0 to 4.5 Myr BP. [9] The oldest divergencies are the Asian hemiones (subgenus E. (Asinus), including the kulan, onager, and kiang), followed by the African zebras (subgenera E. (Dolichohippus), and E. (Hippotigris)). All other modern forms including the domesticated horse (and many fossil Pliocene and Pleistocene forms) belong to the subgenus E. (Equus) which diverged ~4.8 (3.2–6.5) million years ago. [10]

Habitat and range

The Turkmenian kulan lives in Central Asian deltas, hot and cold deserts or semi-deserts, steppes, arid grasslands and shrublands.

The Turkmen specimen used to be the most widespread of the onagers, ranging from northernmost Iran and northern Afghanistan, the Transcaspian Oblast and western China to Ukraine and southern Siberia. It used to live in Saryesik-Atyrau Desert of Kazakhstan. However, it has gone extinct from that location.

Diet and behavior

The Turkmenian kulan are herbivorous mammals. They feed on herbs, shrubbery and plants. Most of their liquid comes from food, although they often travel to water sources, especially when breast-feeding their offspring.

The mare lives with foals in small herds. The dominant stallion defends the surrounding areas to the water sources and tries to mate with any females that come close to drink. The mare, after a gestation period of about a year, gives birth to a single foal that stays with her mother for the first two years of life.

The scorching heat of the Central Asian deserts makes the Transcaspian wild ass mostly active at dawn and dusk when temperatures are milder. The kulans, like most onagers, are one of the fastest land mammals and can run at high speed of 70 km/h.[ citation needed ]

Threats

Like all onagers, the Turkmenian kulan is threatened by poaching, hunting for their meat and their coat, habitat loss and relationships with apex predators such as Persian leopards, striped hyenas and mostly Himalayan wolves. Extinct predators like the Caspian tiger also preyed on the kulan. However, like other onagers, they have anti-predatory protections. Group of stallions can cooperate and chase off the predators.

Conservation

A Turkmenian kulan at Grodno Zoo, Belarus. Turkmenskii kulan Grodno.JPG
A Turkmenian kulan at Grodno Zoo, Belarus.

In early 1919, the Soviet Union placed the Turkmenian kulan under protection. However, the measures enacted in their favor have not saved them from local extinctions, such as in Kazakhstan, where the subspecies disappeared in 1935. The main kulan population has suffered a dramatic decline in recent years. Badkhyz was the main stronghold and the last habitat of the Transcaspian wild ass which contained a few thousands of individuals, until it was reintroduced in other protected areas in Turkmenistan and other Central Asian countries that was built purposely for the kulans, such as the Gaplaňgyr Nature Reserve, the Çemenebit Sanctuary and the Mäne-Çäçe Sanctuary.

As of 1996-1997 the total population of Turkmenian kulan were estimated at between 500 and 700 kulans in Kazakhstan and about 6,000 kulans in Turkmenistan. Of the latter about 5,000 to 6,000 lived in Badkhyz Zapovednik alone and about 1,000 in artificially restored populations elsewhere in Turkmenistan. In Badkhyz, the Turkmenian kulan population has decreased from 6,000 individuals as of 1993 to 2,400 in 1998 and to 650 in 2002. Currently, there are 900 kulans in Badkhyz. [4] Previously in 2005, there were more than 1,300 Turkmenian kulans in Turkmenistan (850-900 in Badhyz State Nature Reserve and another 445 in seven different reintroduction sites).

Reintroduction projects

After their local extinction in Kazakhstan in the 1930s, the kulan was reintroduced in four localities of Kazakhstan (where live about 900 specimens), and in Uzbekistan afterwards (34 specimens).

The first reintroduction site of Turkmenian kulans in Kazakhstan was the Barsa-Kelmes Nature Reserve. Another 35 were reintroduced to the Aktau-Buzachinsky Sanctuary in 1991, which have grown to a population of more than 100 individuals on the Mangishlak Peninsula. Between 1986 and 1990, 105 Turmenian kulans were reintroduced to the Andasai Sanctuary, and have since increased to 200.

In 1984, 32 kulans were reintroduced to the Kapchagai Game Area, which subsequently became the Altyn-Emel National Park (over 50,000,000 hectares in area), [4] from southeastern Kazakhstan which consists of desert, between the Ili River and the Ak-Tau mountain range, near Lake Kapchagai. The previous census revealed that more than 700 individuals lived at Altyn-Emel. The population of Turkmenian kulan are on the rise, as that said small populations of Turkmenian kulan at Altyn-Emel increased to 2,000 as of 2012. [5] The population has further increased to 3,400 kulans in Altyn-Emel as of 2017. [7]

In Kazakhstan, BirdLife partner, the ACBK (Association for the Conservation of Biodiversity of Kazakhstan) has recently started work on a project to establish a new kulan population in Central Asian steppe. On 24 October 2017, nine kulans were taken from Altyn-Emel and released to Altyn Dala protected area of the central Kazakh steppes. The reintroduction project aims to further move 30 or 40 kulans from Altyn Emel to the central steppes during the next 3 or 4 years. [11] [12] They are also released in Irgiz-Turgai. [7]

The Turkmenian kulans, along with the Przewalski's horse, have been reintroduced to the Askania-Nova Biosphere Reserve in southern Ukraine. In 2020, a herd of 20 kulans sourced from the Askania-Nova Biosphere Reserve were reintroduced to the Tarutyns'kyj steppe in southwestern Ukraine. [13] A July 2023 Wired story reported that at least two kulan foals had been born at Tarutino Steppe preserve, and despite the disruptions caused by nearby military activity, the herd has adapted well. [14]

In captivity

The Turkmenian kulans are breeding in captivity in zoos, breeding centers and wildlife parks part of the American (SSP, AZA) and Eurasian (EEP, EAZA) captive breeding programs. There are over 1,000 Turkmenian kulans registered in the international studbook of the World Association of Zoos and Aquariums (WAZA). [6]

Related Research Articles

<span class="mw-page-title-main">Equidae</span> Family of hoofed mammals

Equidae is the taxonomic family of horses and related animals, including the extant horses, asses, and zebras, and many other species known only from fossils. The family evolved around 50 million years ago from a small, multi-toed ungulate into larger, single-toed animals. All extant species are in the genus Equus, which originated in North America. Equidae belongs to the order Perissodactyla, which includes the extant tapirs and rhinoceros, and several extinct families.

<span class="mw-page-title-main">Kyzylkum Desert</span> Desert in Central Asia between the Amu Darya and Syr Darya

The Kyzylkum Desert is the 15th largest desert in the world. Its name means Red Sand in Turkic languages. It is located in Central Asia, in the land between the confluent rivers Amu Darya and Syr Darya, a region historically known as Transoxania. Today it is divided among Kazakhstan, Turkmenistan and Uzbekistan. It covers about 298,000 km2 (115,000 sq mi).

<span class="mw-page-title-main">Onager</span> Species of mammal

The onager, also known as hemione or Asiatic wild ass, is a species of the family Equidae native to Asia. A member of the subgenus Asinus, the onager was described and given its binomial name by German zoologist Peter Simon Pallas in 1775. Six subspecies have been recognized, two of which are extinct.

Turkmen, Türkmen, Turkoman, or Turkman may refer to:

<span class="mw-page-title-main">Kiang</span> Tibetan wild ass

The kiang is the largest of the Asinus subgenus. It is native to the Tibetan Plateau in Ladakh, northern Pakistan, Tajikistan, China and northern Nepal. It inhabits montane grasslands and shrublands. Other common names for this species include Tibetan wild ass, khyang and gorkhar.

The wild asses (Asinus) are a subgenus of single toed grazing ungulates. Its species are:

<span class="mw-page-title-main">Mongolian wild ass</span> Subspecies of onager

The Mongolian wild ass, also known as Mongolian khulan, is the nominate subspecies of the onager. It is found in southern Mongolia and northern China. It was previously found in eastern Kazakhstan and southern Siberia before being extirpated there through hunting. As of 2015, the Mongolian wild ass is listed as Near Threatened by the IUCN. Current population estimates are approximately 42,000 individuals in Mongolia and around 5,000 individuals in Northern China.

<span class="mw-page-title-main">African wild ass</span> Species of wild ass

The African wild ass or African wild donkey is a wild member of the horse family, Equidae. This species is thought to be the ancestor of the domestic donkey, which is sometimes placed within the same species. They live in the deserts and other arid areas of the Horn of Africa, in Eritrea, Ethiopia and Somalia. It formerly had a wider range north and west into Sudan, Egypt, and Libya. It is Critically Endangered, with about 570 existing in the wild.

<i>Asinus</i> Subgenus of mammals

Asinus is a subgenus of Equus that encompasses several subspecies of the Equidae commonly known as wild asses, characterized by long ears, a lean, straight-backed build, lack of a true withers, a coarse mane and tail, and a reputation for considerable toughness and endurance.

<span class="mw-page-title-main">Yotvata Hai-Bar Nature Reserve</span> Nature reserve in Israel

The Yotvata Hai-Bar Nature Reserve is a 3,000-acre (12 km2) breeding and reacclimation center administered by the Israel Nature Reserves & National Parks Authority, situated in the Southern Arava near Yotvata.

<span class="mw-page-title-main">Syrian wild ass</span> Extinct subspecies of onager

The Syrian wild ass, less commonly known as a hemippe, an achdari, or a Mesopotamian or Syrian onager, is an extinct subspecies of onager native to the Arabian peninsula and surrounding areas. It ranged across present-day Iraq, Palestine, Israel, Iran, Jordan, Saudi Arabia, Syria, and Turkey.

<span class="mw-page-title-main">Indian wild ass</span> Subspecies of onager

The Indian wild ass, also called the Indian onager or, in the local Gujarati language, Ghudkhur and Khur, is a subspecies of the onager native to South Asia.

<span class="mw-page-title-main">Altyn-Emel National Park</span>

Altyn-Emel National Park is a national park in Kazakhstan. It was created in 1996. The park covers about 4,600 km2 (1,800 sq mi) between the Ili River and the Ak-Tau mountain range, near Lake Kapchagai, and consists mostly of desert and rocky terrain.

The kulan, is a wild ass.

<span class="mw-page-title-main">Badhyz State Nature Reserve</span> Nature reserve in Turkmenistan

The Badhyz State Nature Reserve is a protected area (zapovednik) in south-western Turkmenistan that was established in 1941 and extends over 877 km2 (339 sq mi) in the Mary and Akhal Provinces. It is located south of the Karakum Desert, and the Tejen River forms its western border.

Çemenebit Sanctuary is a sanctuary (zakaznik) of Turkmenistan.

Şasenem Sanctuary is a sanctuary (zakaznik) of Turkmenistan.

<span class="mw-page-title-main">Persian onager</span> Subspecies of onager

The Persian onager, also called the Persian wild ass or Persian zebra, is a subspecies of onager native to Iran (Persia). It is listed as Endangered, with no more than 600 individuals left in the wild and only 30 individuals living within North American institutions.

<span class="mw-page-title-main">Badghyz and Karabil semi-desert</span> Ecoregion in Asia

The Badghyz and Karabil semi-desert ecoregion covers the hills north of the central mountain ranges of Afghanistan, southeast Turkmenistan, and portions of Uzbekistan and Tajikistan. The landscape has been described as "savannah-like" and reminiscent of Africa, with desert sedges (Carex) and stands of wild pistachio trees. The area supports high biodiversity and a number of rare and endemic species, such as the endangered Turkmenian kulan (Equus hemionus kulan).

The Altyn Dala Conservation Initiative (ADCI) is a longstanding collaborative partnership of national and international NGOs and the Government of Kazakhstan dedicated to safeguarding and restoring the diverse steppe, desert, and wetland ecosystems of Kazakhstan, for the benefit of both wildlife and people. Geographically, the focus is on the parts of the historical range of the Saiga antelope which are still re-occupiable.

References

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