Angarella | |
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Scientific classification ![]() | |
Kingdom: | Animalia |
Clade: | Lophophorata |
Phylum: | Brachiopoda (?) |
Order: | † Kirengellida |
Family: | † Kirengellidae |
Genus: | † Angarella |
Angarella is an early palaeozoic genus of problematic fossils, long held to belong to the molluscs based on supposed similarities in its musculature to that of hyoliths (now known to be brachiopod relatives [1] ) and mobergellids, but now considered to represent a brachiopod based on its calcitic mineralogy and paired symmetrical valves. [2]
Its dorsal valve is low and conical, whereas its ventral is flat and follows the contours of the substrate, indicating a cementing habit. [2] The dorsal valve bears a series of symmetrical muscle impressions, forming a ring concentric with the edge of the shell. The ventral valve has a pair of internal depressions. [2]
Hyoliths are animals with small conical shells, known as fossils from the Palaeozoic Era. They are lophophorates, a group which includes the brachiopods.
The lophophore is a characteristic feeding organ possessed by four major groups of animals: the Brachiopoda, Bryozoa, Hyolitha, and Phoronida, which collectively constitute the protostome group Lophophorata. All lophophores are found in aquatic organisms.
Inarticulata was historically defined as one of the two classes of the phylum Brachiopoda and referred to those having no hinge. The other class was Articulata, meaning articulated — having a hinge between the dorsal and ventral valves. These classifications have now been superseded, see brachiopod classification.
Haplophrentis is a genus of tiny shelled hyolithid which lived in the Cambrian Period. Its shell was long and conical, with the open end protected by an operculum, from which two fleshy arms called helens protruded at the sides. These arms served to elevate the opening of the shells above the sea floor, acting like stilts.
Lingula is a genus of brachiopods within the class Lingulata. Lingula or forms very close in appearance have existed possibly since the Cambrian. Like its relatives, it has two unadorned organo-phosphatic valves and a long fleshy stalk. Lingula lives in burrows in barren sandy coastal seafloor and feeds by filtering detritus from the water. It can be detected by a short row of three openings through which it takes in water (sides) and expels it again (middle).
The Mesozoic marine revolution is the rapid adaption to shell-crushing (durophagous) and boring predation in benthic organisms throughout the Mesozoic era. The term was first coined by Geerat J. Vermeij, who based his work on that of Steven M. Stanley. While initially restricted to the Late Cretaceous, more recent studies have suggested that the beginning of this ecological arms race extends back into the Norian. It is an important transition between the Palaeozoic evolutionary fauna and the Cenozoic evolutionary fauna that occurred throughout the Mesozoic.
Strophomenida is a large, extinct order of articulate brachiopods in the extinct class Strophomenata that existed from the lower Ordovician to the lower Jurassic period. It was the largest known order of brachiopods, encompassing over 400 genera, including the largest and heaviest of known brachiopod shells. The strophomenids lost the ability to attach by the stalk in adult specimens, so they either lay free, attached the ventral valve at the umbo to a firm substrate, or balanced with their spines sunken into a soft substrate. Typically the dorsal valve was either concave or flat, though occasionally it was convex; the ventral valve was convex. Typically, a member of this order was wider than it was long. In juveniles, there was a tiny hole at the animal's umbo for a stalk to emerge from. Tiny bumps cover the interior of the valves.
Brachiopods, phylum Brachiopoda, are a group of lophotrochozoan animals that have hard "valves" (shells) on the upper and lower surfaces, unlike the left and right arrangement in bivalve molluscs. Brachiopod valves are hinged at the rear end, while the front can be opened for feeding or closed for protection. Two major groups are recognized, articulate and inarticulate. The word "articulate" is used to describe the tooth-and-groove features of the valve-hinge which is present in the articulate group, and absent from the inarticulate group. This is the leading diagnostic feature (fossilizable), by which the two main groups can be readily distinguished. Articulate brachiopods have toothed hinges and simple opening and closing muscles, while inarticulate brachiopods have untoothed hinges and a more complex system of muscles used to keep the two valves aligned. In a typical brachiopod a stalk-like pedicle projects from an opening in one of the valves near the hinges, known as the pedicle valve, keeping the animal anchored to the seabed but clear of silt that would obstruct the opening.
The origin of the brachiopods is uncertain; they either arose from reduction of a multi-plated tubular organism, or from the folding of a slug-like organism with a protective shell on either end. Since their Cambrian origin, the phylum rose to a Palaeozoic dominance, but dwindled during the Mesozoic.
The cephalopods have a long geological history, with the first nautiloids found in late Cambrian strata, and purported stem-group representatives present in the earliest Cambrian lagerstätten.
Stenothecoida is a taxon of bivalved fossils from the Early to middle Cambrian period. They look a bit like brachiopods or bivalve molluscs.
Turcutheca is a Tommotian genus of shelly fossil whose affinities are uncertain, generally considered as an orthothecid hyolith (which would make it a brachiopod, but also resembling the ellesmeroceratids.
Mickwitziids are a Cambrian group of shelly fossils with originally phosphatic valves, belonging to the Brachiopod stem group, and exemplified by the genus Mickwitzia – the other genera are Heliomedusa and Setatella. The family Mickwitziidae is conceivably paraphyletic with respect to certain crown-group brachiopods.
Sharpirhynchia sharpi is a species of extinct, small-sized lampshell, named after Samuel Sharp (1814–1882), an early fossil collector, who discovered the limited time span of some fossils, now known as index fossils. S. sharpi is a marine rhynchonellate brachiopod in the family Rhynchonellidae. It is roughly ½ inch (1.25 cm) measured along the axis, with a slender beak, the brachial valve more convex than the pedunculate valve, and it has 21–31 ribs fanning out from the hinge.
Gigantoproductus giganteus is an extinct species of brachiopods in the family Monticuliferidae, known only from its fossil remains. It was a marine invertebrate found on the seabed in shallow seas. It probably evolved during the Devonian period and it is believed to be the largest brachiopod that has ever existed.
Argyrotheca is a genus of very small to minute lampshells. All species share a large pedicel opening, one ridge on the inside of the pedunculate valve, pits in a diamond pattern on the inside of both valves, and without radial ridges that end in tubercles. It occurs in depths between 6 and 1300 m. It is known since the latest Cretaceous.
Craniscus is a genus of small brachiopods. The shell is approximately round to square with one side rounded. The ventral valve is approximately flat and attached to the underground by the entire surface. The outside of the dorsal valve is more or less conical, the inside is divided by three ridges that join in the center.
Valdiviathyris quenstedti is a small species of brachiopods with a maximum size of about 0.3 inches or 7.6 millimetres wide.
The camenellans, consisting of the genera Camenalla, Dailyatia, Kennardia, Kelanella and Lapworthella, are a group of Tommotiid invertebrates from the Cambrian period, reconstructed as sister to all others. They are known from isolated sclerites, but are believed to have a scleritomous, Halkieria-like construction.