A trace fossil, also known as an ichnofossil, is a fossil record of biological activity but not the preserved remains of the plant or animal itself. Trace fossils contrast with body fossils, which are the fossilized remains of parts of organisms' bodies, usually altered by later chemical activity or mineralization. The study of such trace fossils is ichnology and is the work of ichnologists.
Sauropoda, whose members are known as sauropods, is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads, and four thick, pillar-like legs. They are notable for the enormous sizes attained by some species, and the group includes the largest animals to have ever lived on land. Well-known genera include Brachiosaurus, Diplodocus, Apatosaurus and Brontosaurus.
A fossil track or ichnite is a fossilized footprint. This is a type of trace fossil. A fossil trackway is a sequence of fossil tracks left by a single organism. Over the years, many ichnites have been found, around the world, giving important clues about the behaviour of the animals that made them. For instance, multiple ichnites of a single species, close together, suggest 'herd' or 'pack' behaviour of that species.
Europasaurus is a basal macronarian sauropod, a form of quadrupedal herbivorous dinosaur. It lived during the Late Jurassic of northern Germany, and has been identified as an example of insular dwarfism resulting from the isolation of a sauropod population on an island within the Lower Saxony basin.
Footprints are the impressions or images left behind by a person walking or running. Hoofprints and pawprints are those left by animals with hooves or paws rather than feet, while "shoeprints" is the specific term for prints made by shoes. They may either be indentations in the ground or something placed onto the surface that was stuck to the bottom of the foot. A "trackway" is a set of footprints in soft earth left by a life-form; animal tracks are the footprints, hoofprints, or pawprints of an animal.
Dinosaur Footprints in Holyoke, Massachusetts, USA is an 8-acre (3 ha) wilderness reservation purchased for the public in 1935 by The Trustees of Reservations. The Reservation is currently being managed with the assistance from the Massachusetts Department of Conservation and Recreation (DCR). The fossil and plant resources on the adjacent Holyoke Gas and Electric (HG&E) riverfront property are being managed cooperatively by The Trustees, Mass DCR, and HG&E.
Dinosaur Ridge is a segment of the Dakota Hogback in the Morrison Fossil Area National Natural Landmark located in Jefferson County, Colorado, near the town of Morrison and just west of Denver.
Paleontology in Maryland refers to paleontological research occurring within or conducted by people from the U.S. state of Maryland. The invertebrate fossils of Maryland are similar to those of neighboring Delaware. For most of the early Paleozoic era, Maryland was covered by a shallow sea, although it was above sea level for portions of the Ordovician and Devonian. The ancient marine life of Maryland included brachiopods and bryozoans while horsetails and scale trees grew on land. By the end of the era, the sea had left the state completely. In the early Mesozoic, Pangaea was splitting up. The same geologic forces that divided the supercontinent formed massive lakes. Dinosaur footprints were preserved along their shores. During the Cretaceous, the state was home to dinosaurs. During the early part of the Cenozoic era, the state was alternatingly submerged by sea water or exposed. During the Ice Age, mastodons lived in the state.
Paleontology in Pennsylvania refers to paleontological research occurring within or conducted by people from the U.S. state of Pennsylvania. The geologic column of Pennsylvania spans from the Precambrian to Quaternary. During the early part of the Paleozoic, Pennsylvania was submerged by a warm, shallow sea. This sea would come to be inhabited by creatures like brachiopods, bryozoans, crinoids, graptolites, and trilobites. The armored fish Palaeaspis appeared during the Silurian. By the Devonian the state was home to other kinds of fishes. On land, some of the world's oldest tetrapods left behind footprints that would later fossilize. Some of Pennsylvania's most important fossil finds were made in the state's Devonian rocks. Carboniferous Pennsylvania was a swampy environment covered by a wide variety of plants. The latter half of the period was called the Pennsylvanian in honor of the state's rich contemporary rock record. By the end of the Paleozoic the state was no longer so swampy. During the Mesozoic the state was home to dinosaurs and other kinds of reptiles, who left behind fossil footprints. Little is known about the early to mid Cenozoic of Pennsylvania, but during the Ice Age it seemed to have a tundra-like environment. Local Delaware people used to smoke mixtures of fossil bones and tobacco for good luck and to have wishes granted. By the late 1800s Pennsylvania was the site of formal scientific investigation of fossils. Around this time Hadrosaurus foulkii of neighboring New Jersey became the first mounted dinosaur skeleton exhibit at the Academy of Natural Sciences in Philadelphia. The Devonian trilobite Phacops rana is the Pennsylvania state fossil.
Paleontology in New Jersey refers to paleontological research in the US state of New Jersey. The state is especially rich in marine deposits.
Paleontology in Connecticut refers to paleontological research occurring within or conducted by people from the U.S. state of Connecticut. Apart from its famous dinosaur tracks, the fossil record in Connecticut is relatively sparse. The oldest known fossils in Connecticut date back to the Triassic period. At the time, Pangaea was beginning to divide and local rift valleys became massive lakes. A wide variety of vegetation, invertebrates and reptiles are known from Triassic Connecticut. During the Early Jurassic local dinosaurs left behind an abundance of footprints that would later fossilize.
Paleontology in Arkansas refers to paleontological research occurring within or conducted by people from the U.S. state of Arkansas. The fossil record of Arkansas spans from the Ordovician to the Eocene. Nearly all of the state's fossils have come from ancient invertebrate life. During the early Paleozoic, much of Arkansas was covered by seawater. This sea would come to be home to creatures including Archimedes, brachiopods, and conodonts. This sea would begin its withdrawal during the Carboniferous, and by the Permian the entire state was dry land. Terrestrial conditions continued into the Triassic, but during the Jurassic, another sea encroached into the state's southern half. During the Cretaceous the state was still covered by seawater and home to marine invertebrates such as Belemnitella. On land the state was home to long necked sauropod dinosaurs, who left behind footprints and ostrich dinosaurs such as Arkansaurus.
Paleontology in Missouri refers to paleontological research occurring within or conducted by people from the U.S. state of Missouri. The geologic column of Missouri spans all of geologic history from the Precambrian to present with the exception of the Permian, Triassic, and Jurassic. Brachiopods are probably the most common fossils in Missouri.
Paleontology in Texas refers to paleontological research occurring within or conducted by people from the U.S. state of Texas. Author Marian Murray has remarked that "Texas is as big for fossils as it is for everything else." Some of the most important fossil finds in United States history have come from Texas. Fossils can be found throughout most of the state. The fossil record of Texas spans almost the entire geologic column from Precambrian to Pleistocene. Shark teeth are probably the state's most common fossil. During the early Paleozoic era Texas was covered by a sea that would later be home to creatures like brachiopods, cephalopods, graptolites, and trilobites. Little is known about the state's Devonian and early Carboniferous life. However, evidence indicates that during the late Carboniferous the state was home to marine life, land plants and early reptiles. During the Permian, the seas largely shrank away, but nevertheless coral reefs formed in the state. The rest of Texas was a coastal plain inhabited by early relatives of mammals like Dimetrodon and Edaphosaurus. During the Triassic, a great river system formed in the state that was inhabited by crocodile-like phytosaurs. Little is known about Jurassic Texas, but there are fossil aquatic invertebrates of this age like ammonites in the state. During the Early Cretaceous local large sauropods and theropods left a great abundance of footprints. Later in the Cretaceous, the state was covered by the Western Interior Seaway and home to creatures like mosasaurs, plesiosaurs, and few icthyosaurs. Early Cenozoic Texas still contained areas covered in seawater where invertebrates and sharks lived. On land the state would come to be home to creatures like glyptodonts, mammoths, mastodons, saber-toothed cats, giant ground sloths, titanotheres, uintatheres, and dire wolves. Archaeological evidence suggests that local Native Americans knew about local fossils. Formally trained scientists were already investigating the state's fossils by the late 1800s. In 1938, a major dinosaur footprint find occurred near Glen Rose. Pleurocoelus was the Texas state dinosaur from 1997 to 2009, when it was replaced by Paluxysaurus jonesi after the Texan fossils once referred to the former species were reclassified to a new genus.
Paleontology in Wyoming includes research into the prehistoric life of the U.S. state of Wyoming as well as investigations conducted by Wyomingite researchers and institutions into ancient life occurring elsewhere. The fossil record of the US state of Wyoming spans from the Precambrian to recent deposits. Many fossil sites are spread throughout the state. Wyoming is such a spectacular source of fossils that author Marian Murray noted in 1974 that "[e]ven today, it is the expected thing that any great museum will send its representatives to Wyoming as often as possible." Murray has also written that nearly every major vertebrate paleontologist in United States history has collected fossils in Wyoming. Wyoming is a major source of dinosaur fossils. Wyoming's dinosaur fossils are curated by museums located all over the planet.
Paleontology in Colorado refers to paleontological research occurring within or conducted by people from the U.S. state of Colorado. The geologic column of Colorado spans about one third of Earth's history. Fossils can be found almost everywhere in the state but are not evenly distributed among all the ages of the state's rocks. During the early Paleozoic, Colorado was covered by a warm shallow sea that would come to be home to creatures like brachiopods, conodonts, ostracoderms, sharks and trilobites. This sea withdrew from the state between the Silurian and early Devonian leaving a gap in the local rock record. It returned during the Carboniferous. Areas of the state not submerged were richly vegetated and inhabited by amphibians that left behind footprints that would later fossilize. During the Permian, the sea withdrew and alluvial fans and sand dunes spread across the state. Many trace fossils are known from these deposits.
Paleontology in New Mexico refers to paleontological research occurring within or conducted by people from the U.S. state of New Mexico. The fossil record of New Mexico is exceptionally complete and spans almost the entire stratigraphic column. More than 3,300 different kinds of fossil organisms have been found in the state. Of these more than 700 of these were new to science and more than 100 of those were type species for new genera. During the early Paleozoic, southern and western New Mexico were submerged by a warm shallow sea that would come to be home to creatures including brachiopods, bryozoans, cartilaginous fishes, corals, graptolites, nautiloids, placoderms, and trilobites. During the Ordovician the state was home to algal reefs up to 300 feet high. During the Carboniferous, a richly vegetated island chain emerged from the local sea. Coral reefs formed in the state's seas while terrestrial regions of the state dried and were home to sand dunes. Local wildlife included Edaphosaurus, Ophiacodon, and Sphenacodon.
Paleontology in Utah refers to paleontological research occurring within or conducted by people from the U.S. state of Utah. Utah has a rich fossil record spanning almost all of the geologic column. During the Precambrian, the area of northeastern Utah now occupied by the Uinta Mountains was a shallow sea which was home to simple microorganisms. During the early Paleozoic Utah was still largely covered in seawater. The state's Paleozoic seas would come to be home to creatures like brachiopods, fishes, and trilobites. During the Permian the state came to resemble the Sahara desert and was home to amphibians, early relatives of mammals, and reptiles. During the Triassic about half of the state was covered by a sea home to creatures like the cephalopod Meekoceras, while dinosaurs whose footprints would later fossilize roamed the forests on land. Sand dunes returned during the Early Jurassic. During the Cretaceous the state was covered by the sea for the last time. The sea gave way to a complex of lakes during the Cenozoic era. Later, these lakes dissipated and the state was home to short-faced bears, bison, musk oxen, saber teeth, and giant ground sloths. Local Native Americans devised myths to explain fossils. Formally trained scientists have been aware of local fossils since at least the late 19th century. Major local finds include the bonebeds of Dinosaur National Monument. The Jurassic dinosaur Allosaurus fragilis is the Utah state fossil.
Paleontology in Arizona refers to paleontological research occurring within or conducted by people from the U.S. state of Arizona. The fossil record of Arizona dates to the Precambrian. During the Precambrian, Arizona was home to a shallow sea which was home to jellyfish and stromatolite-forming bacteria. This sea was still in place during the Cambrian period of the Paleozoic era and was home to brachiopods and trilobites, but it withdrew during the Ordovician and Silurian. The sea returned during the Devonian and was home to brachiopods, corals, and fishes. Sea levels began to rise and fall during the Carboniferous, leaving most of the state a richly vegetated coastal plain during the low spells. During the Permian, Arizona was richly vegetated but was submerged by seawater late in the period.
The 20th century in ichnology refers to advances made between the years 1900 and 1999 in the scientific study of trace fossils, the preserved record of the behavior and physiological processes of ancient life forms, especially fossil footprints. Significant fossil trackway discoveries began almost immediately after the start of the 20th century with the 1900 discovery at Ipolytarnoc, Hungary of a wide variety of bird and mammal footprints left behind during the early Miocene. Not long after, fossil Iguanodon footprints were discovered in Sussex, England, a discovery that probably served as the inspiration for Sir Arthur Conan Doyle's The Lost World.
