Carinatae is the group of all birds and their extinct relatives to possess a keel, or "carina", on the underside of the breastbone used to anchor large flight muscles.
Carinatae is the group of all birds and their extinct relatives to possess a keel, or "carina", on the underside of the breastbone used to anchor large flight muscles.
Traditionally, Carinatae were defined as all birds whose sternum (breast bone) has a keel (carina). The keel is a strong median ridge running down the length of the sternum. This is an important area for the attachment of flight muscles. Thus, all flying birds have a pronounced keel. Ratites, all of which are flightless, lack a strong keel. Thus, living birds were divided into carinatae (keeled) and ratites (from ratis, "raft", referring to the flatness of the sternum). The difficulty with this scheme phylogenetically was that some flightless birds, without strong keels, are descended directly from ordinary flying birds possessing one. Examples include the kākāpō, a flightless parrot, and the dodo, a columbiform (the pigeon family). Neither of these birds are a ratite. Thus, this supposedly distinctive feature was easy to use, but had nothing to do with actual phylogenetic relationship.[ citation needed ]
Beginning in the 1980s, Carinatae was given several phylogenetic definitions. The first was as a node-based clade uniting Ichthyornis with modern birds. [1] However, in many analyses, this definition would be synonymous with the more widely used name Ornithurae. An alternate definition was provided in 2001, naming Carinatae an apomorphy-based clade defined by the presence of a keeled sternum. [2]
The most primitive known bird relative with a keeled breastbone is Confuciusornis . While some specimens of this stem-bird have flat breastbones, some show a small ridge that could have supported a cartilaginous keel. [3]
A ratite is any of a group of mostly large, long-necked, and long-legged birds of the infraclass Palaeognathae. Kiwi, the exception, are much smaller and shorter-legged and are the only nocturnal extant ratites. All extant ratites are flightless.
Flightless birds are birds that, through evolution, lost the ability to fly. There are over 60 extant species, including the well-known ratites and penguins. The smallest flightless bird is the Inaccessible Island rail. The largest flightless bird, which is also the largest living bird in general, is the ostrich.
In phylogenetics, the crown group or crown assemblage is a collection of species composed of the living representatives of the collection, the most recent common ancestor of the collection, and all descendants of the most recent common ancestor. It is thus a way of defining a clade, a group consisting of a species and all its extant or extinct descendants. For example, Neornithes (birds) can be defined as a crown group, which includes the most recent common ancestor of all modern birds, and all of its extant or extinct descendants.
Alvarezsauridae is a family of small, long-legged dinosaurs. Although originally thought to represent the earliest known flightless birds, they are now thought to be an early diverging branch of maniraptoran theropods. Alvarezsaurids were highly specialized. They had tiny but stout forelimbs, with compact, bird-like hands. Their skeletons suggest that they had massive breast and arm muscles, possibly adapted for digging or tearing. They had long, tube-shaped snouts filled with tiny teeth. They have been interpreted as myrmecophagous, adapted to prey on colonial insects such as termites, with the short arms acting as effective digging instruments to break into nests.
A keel or carina in bird anatomy is an extension of the sternum (breastbone) which runs axially along the midline of the sternum and extends outward, perpendicular to the plane of the ribs. The keel provides an anchor to which a bird's wing muscles attach, thereby providing adequate leverage for flight. Not all birds have keels; in particular, some flightless birds lack a keel structure. Some flightless birds have a keel, such as the penguin; but in the penguin's case, its wings are too small for its body, so flight would require flapping its wings too fast to be practical.
Dromornithidae, known as mihirungs and informally as thunder birds or demon ducks, were a clade of large, flightless Australian birds of the Oligocene through Pleistocene Epochs. All are now extinct. They were long classified in Struthioniformes, but are now usually classified as galloanseres. Dromornithids were part of the Australian megafauna. One species, Dromornis stirtoni, was 3 m tall. Only a single species, Genyornis newtoni survived into the Late Pleistocene. They are thought to have been herbivorous.
Palaeognathae is an infraclass of birds, called paleognaths or palaeognaths, within the class Aves of the clade Archosauria. It is one of the two extant infraclasses of birds, the other being Neognathae, both of which form Neornithes. Palaeognathae contains five extant branches of flightless lineages, termed ratites, and one flying lineage, the Neotropic tinamous. There are 47 species of tinamous, five of kiwis (Apteryx), three of cassowaries (Casuarius), one of emus (Dromaius), two of rheas (Rhea) and two of ostriches (Struthio). Recent research has indicated that paleognaths are monophyletic but the traditional taxonomic split between flightless and flighted forms is incorrect; tinamous are within the ratite radiation, meaning flightlessness arose independently multiple times via parallel evolution.
The evolution of birds began in the Jurassic Period, with the earliest birds derived from a clade of theropod dinosaurs named Paraves. Birds are categorized as a biological class, Aves. For more than a century, the small theropod dinosaur Archaeopteryx lithographica from the Late Jurassic period was considered to have been the earliest bird. Modern phylogenies place birds in the dinosaur clade Theropoda. According to the current consensus, Aves and a sister group, the order Crocodilia, together are the sole living members of an unranked reptile clade, the Archosauria. Four distinct lineages of bird survived the Cretaceous–Paleogene extinction event 66 million years ago, giving rise to ostriches and relatives (Palaeognathae), waterfowl (Anseriformes), ground-living fowl (Galliformes), and "modern birds" (Neoaves).
Avemetatarsalia is a clade of diapsid reptiles containing all archosaurs more closely related to birds than to crocodilians. The two most successful groups of avemetatarsalians were the dinosaurs and pterosaurs. Dinosaurs were the largest terrestrial animals for much of the Mesozoic Era, and one group of small feathered dinosaurs has survived up to the present day. Pterosaurs were the first flying vertebrates and persisted through the Mesozoic before dying out at the Cretaceous-Paleogene (K-Pg) extinction event. Both dinosaurs and pterosaurs appeared in the Triassic Period, shortly after avemetatarsalians as a whole. The name Avemetatarsalia was first established by British palaeontologist Michael Benton in 1999. An alternate name is Pan-Aves, or "all birds", in reference to its definition containing all animals, living or extinct, which are more closely related to birds than to crocodilians.
Yandangornis is a genus of theropods from the Late Cretaceous Tangshang Formation of China. It lived 81.5 million years ago in what is now China. The type species, Y. longicaudus, was formally described by Cai and Zhou in 1999.
Yungavolucris is a genus of enantiornithean birds. It contains the single species Yungavolucris brevipedalis, which lived in the Late Cretaceous. The fossil bones were found in the Lecho Formation at estancia El Brete, Argentina."Yungavolucris brevipedalis" means "Short-footed Yungas bird". The generic name, Yungavolucris is after the Yungas region + the Latin volucris, which translates to "bird". The specific name brevipedalis is from the Latin brevis, which means "short", + pedalis, from the Latin pes, meaning "foot".
Maniraptoriformes is a clade of dinosaurs with pennaceous feathers and wings that contains ornithomimosaurs and maniraptorans. This group was named by Thomas Holtz, who defined it as "the most recent common ancestor of Ornithomimus and birds, and all descendants of that common ancestor."
Avialae is a clade containing the only living dinosaurs, the birds. It is usually defined as all theropod dinosaurs more closely related to birds (Aves) than to deinonychosaurs, though alternative definitions are occasionally used.
Confuciusornithidae is an extinct family of pygostylian avialans known from the Early Cretaceous, found in northern China. They are commonly placed as a sister group to Ornithothoraces, a group that contains all extant birds along with their closest extinct relatives. Confuciusornithidae contains four genera, possessing both shafted and non-shafted (downy) feathers. They are also noted for their distinctive pair of ribbon-like tail feathers of disputed function.
Archaeorhynchus is a genus of beaked avialan stem-birds from the early Cretaceous period. A fossil of its only known species, Archaeorhynchus spathula, was first reported in 2005 by Zhou & Zhang to have been found in Yixian Formation rocks at Yixian, Liaoning province, China, showing a well-preserved and essentially complete skeleton. Two more complete specimens were found in Lower Cretaceous deposits of Jianchang, Liaoning, northeastern China, preserving new anatomical information. These deposits are 120 million years old, whereas the original specimen was 125 million years old, meaning the age range for this species is 125-120Ma.
Sauriurae is a now-deprecated subclass of birds created by Ernst Haeckel in 1866. It was intended to include Archaeopteryx and distinguish it from all other birds then known, which he grouped in the sister-group Ornithurae. The distinction Haeckel referred to in this name is that Archaeopteryx possesses a long, reptile-like tail, while all other birds known to him had short tails with few vertebrae, fused at the end into a pygostyle. The unit was not much referred to, and when Hans Friedrich Gadow in 1893 erected Archaeornithes for basically the same fossils, this became the common name for the early reptile-like grade of birds.
Ornithothoraces is a group of avialans that includes all enantiornithes and the euornithes, which includes modern birds and their closest ancestors. The name Ornithothoraces means "bird thoraxes". This refers to the modern, highly advanced anatomy of the thorax that gave the ornithothoracines superior flight capability compared with more primitive avialans. This anatomy includes a large, keeled breastbone, elongated coracoids and a modified glenoid joint in the shoulder, and a semi-rigid rib cage. In spite of this at least the sternum seems to have developed convergently rather than being a true homology.
Paraphysornis is an extinct genus of giant flightless terror birds that inhabited Brazil during Late Oligocene or Early Miocene epochs. Although not the tallest phorusrhacid, Paraphysornis was a notably robust bird, having short and robust tarsal bones not suited for pursuit hunting.
Notopalaeognathae is a clade that contains the order Rheiformes (rheas), the clade Novaeratitae, and the clade Dinocrypturi. The exact relationships of this group, including its recently extinct members, have only recently been uncovered. The two lineages endemic to New Zealand, the kiwis and the extinct moas, are not each other's closest relatives: the moas are most closely related to the Neotropical tinamous, and the kiwis are sister to the extinct elephant birds of Madagascar, with kiwis and elephant birds together sister to the cassowaries and emu of New Guinea and Australia. The South American rheas are either sister to all other notopalaeognaths or sister to Novaeratitae. The sister group to Notopalaeognathae is Struthionidae.
Paracrax is a genus of extinct North American flightless birds, possibly related to modern seriemas and the extinct terror birds. Part of Bathornithidae, it is a specialised member of this group, being cursorial carnivores much like their South American cousins, some species attaining massive sizes.
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