Cloven hoof

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Cloven hooves of roe deer (Capreolus capreolus) Capreolus capreolus cloven hoof.jpg
Cloven hooves of roe deer ( Capreolus capreolus )

A cloven hoof, cleft hoof, divided hoof, or split hoof is a hoof split into two toes. Members of the mammalian order Artiodactyla that possess this type of hoof include cattle, deer, pigs, antelopes, gazelles, goats, and sheep. [1]

Contents

The two digits of cloven-hooved animals are homologous to the third and fourth fingers of the hand. They are called claws and are named for their relative location on the foot: the outer, or lateral, claw and the inner, or medial claw. The space between the two claws is called the interdigital cleft; the area of skin is called the interdigital skin. The hard outer covering of the hoof is called the hoof wall or horn. It is a hard surface, similar to the human fingernail. [2]

The almost finger-like dexterity available to cloven-hooved mammals such as mountain goats and wild sheep combined with a hard outer shell and soft and flexible inner pads provides excellent traction in their precarious habitats. [3] [4] [5]

Evolution

It is speculated by paleontologists that during the Eocene period hooved marsh dwellers carried their body weight mainly on two of the middle toes, which grew to equal size, becoming the Artiodactyla or even-toed hooved animals. Before the end of the Eocene period the side toes of some had dwindled and practically disappeared (mainly in the form of a dewclaw) while the basal pieces or metapodium of the pair of supporting toes became fused together, thus producing the appearance of a cloven hoof. [6]

The mammal with a cloven hoof is an even-toed ungulate of order Artiodactyla as opposed to the odd-toed ungulates of Perissidactyla, such as the horse, which have one toe, or the rhinoceros, which has three toes. The five-toed ancestors of the earliest Eocene had already developed feet that suggest odd-toed and even-toed descendants to the modern viewer. Even Phenacodus, the most generalized of the early mammals, has a foot in which the central toe is somewhat larger than the others and could be placed in the division of odd-toed ungulates, Perissidactyla. [7]

Jewish dietary laws

The distinction between cloven and uncloven hooves is highly relevant for dietary laws of Judaism ( kashrut ), as set forth in the Torah [8] and the Talmud. [9] [10] Animals that both chew their cud (ruminate, i.e. regurgitate partly digested food from a specialised multi-chambered stomach back to the mouth to be chewed for a second time as part of their ordinary digestive process) and have split true cloven hooves (a hoof being hard or rubbery sole and a hard wall formed by a thick nail) are allowed ( kosher , lit. "fit") for Jewish consumption. Those animals that have neither of these two characteristics, or only one of the characteristics, are considered unclean animals ( treif , not fit for Jewish consumption) and Jews are forbidden to eat them.

This rule thus excludes the camel from the list of kosher animals because although the camel does ruminate it does not possess true "hooves" it walks on soft toes which have little more than a nail merely giving an appearance of a "hoof". Similarly the pig, although it has cloven true hooves, does not ruminate. [11] [12] Other animals such as dogs and cats exhibit neither characteristic and are thus also forbidden for Jewish consumption.

In folklore and popular culture, a cloven hoof has long been associated with the Devil.[ citation needed ]

Related Research Articles

Kashrut is a set of dietary laws dealing with the foods that Jewish people are permitted to eat and how those foods must be prepared according to Jewish law. Food that may be consumed is deemed kosher, from the Ashkenazi pronunciation of the term that in Sephardic or Modern Hebrew is pronounced kashér, meaning "fit".

<span class="mw-page-title-main">Perissodactyla</span> Order of hoofed mammals

Perissodactyla is an order of ungulates. The order includes about 17 living species divided into three families: Equidae, Rhinocerotidae (rhinoceroses), and Tapiridae (tapirs). They typically have reduced the weight-bearing toes to three or one of the five original toes, though tapirs retain four toes on their front feet. The nonweight-bearing toes are either present, absent, vestigial, or positioned posteriorly. By contrast, artiodactyls bear most of their weight equally on four or two of the five toes: their third and fourth toes. Another difference between the two is that odd-toed ungulates digest plant cellulose in their intestines, rather than in one or more stomach chambers as even-toed ungulates, with the exception of Suina, do.

<span class="mw-page-title-main">Ungulate</span> Group of animals that use the tips of their toes or hooves to walk on

Ungulates are members of the diverse clade Euungulata which primarily consists of large mammals with hooves. Once part of the clade "Ungulata" along with the clade Paenungulata, "Ungulata" has since been determined to be a polyphyletic and thereby invalid clade based on molecular data. As a result, true ungulates had since been reclassified to the newer clade Euungulata in 2001 within the clade Laurasiatheria while Paenungulata has been reclassified to a distant clade Afrotheria. Living ungulates are divided into two orders: Perissodactyla including equines, rhinoceroses, and tapirs; and Artiodactyla including cattle, antelope, pigs, giraffes, camels, sheep, deer, and hippopotamuses, among others. Cetaceans such as whales, dolphins, and porpoises are also classified as artiodactyls, although they do not have hooves. Most terrestrial ungulates use the hoofed tips of their toes to support their body weight while standing or moving. Two other orders of ungulates, Notoungulata and Litopterna, both native to South America, became extinct at the end of the Pleistocene, around 12,000 years ago.

<span class="mw-page-title-main">Artiodactyl</span> Order of mammals

Artiodactyls are mammals belonging to the order Artiodactyla. Typically, they are ungulates which bear weight equally on two of their five toes: the third and fourth, often in the form of a hoof. The other three toes are either present, absent, vestigial, or pointing posteriorly. By contrast, odd-toed ungulates bear weight on an odd number of the five toes. Another difference between the two is that many even-toed ungulates digest plant cellulose in one or more stomach chambers rather than in their intestine as the odd-toed ungulates do. The advent of molecular biology, along with new fossil discoveries, found that cetaceans fall within this taxonomic branch, being most closely related to hippopotamuses. Some modern taxonomists thus apply the name Cetartiodactyla to this group, while others opt to include cetaceans within the existing name of Artiodactyla. Some researchers use "even-toed ungulates" to exclude cetaceans and only include terrestrial artiodactyls, making the term paraphyletic in nature.

<span class="mw-page-title-main">Claw</span> Curved, pointed appendage at the end of a digit of a mammal or reptile

A claw is a curved, pointed appendage found at the end of a toe or finger in most amniotes. Some invertebrates such as beetles and spiders have somewhat similar fine, hooked structures at the end of the leg or tarsus for gripping a surface as they walk. The pincers of crabs, lobsters and scorpions, more formally known as their chelae, are sometimes called claws.

<span class="mw-page-title-main">Hoof</span> Tip of a toe of an ungulate mammal, strengthened by a thick and horny keratin covering

The hoof is the tip of a toe of an ungulate mammal, which is covered and strengthened with a thick and horny keratin covering. Artiodactyls are even-toed ungulates, species whose feet have an even number of digits; the ruminants with two digits are the most numerous, e.g. giraffe, deer, bison, cattle, goat, and sheep. The feet of perissodactyl mammals have an odd number of toes, e.g. the horse, the rhinoceros, and the tapir. Although hooves are limb structures primarily found in placental mammals, hadrosaurs such as Edmontosaurus possessed hoofed forelimbs. The marsupial Chaeropus also had hooves.

<span class="mw-page-title-main">Mountain goat</span> Species of mammal

The mountain goat, also known as the Rocky Mountain goat, is a cloven-footed mammal that is endemic to the remote and rugged mountainous areas of western North America. A subalpine to truly alpine species, it is a sure-footed climber commonly seen on sheer rock faces, near-vertical cliffs and icy passages. Mountain goats generally avoid venturing down into lower elevations—except during seasonal food shortages or during particularly bad weather—as the extreme elevation which they inhabit is their primary defense against predators such as black and brown bears, pumas and wolves.

<span class="mw-page-title-main">Unclean animal</span> Animal whose consumption or handling is taboo

In some religions, an unclean animal is an animal whose consumption or handling is taboo. According to these religions, persons who handle such animals may need to ritually purify themselves to get rid of their uncleanliness.

Kosher foods are foods that conform to the Jewish dietary regulations of kashrut. The laws of kashrut apply to food derived from living creatures and kosher foods are restricted to certain types of mammals, birds and fish meeting specific criteria; the flesh of any animals that do not meet these criteria is forbidden by the dietary laws. Furthermore, kosher mammals and birds must be slaughtered according to a process known as shechita and their blood may never be consumed and must be removed from the meat by a process of salting and soaking in water for the meat to be permissible for use. All plant-based products, including fruits, vegetables, grains, herbs and spices, are intrinsically kosher, although certain produce grown in the Land of Israel is subjected to other requirements, such as tithing, before it may be consumed.

<span class="mw-page-title-main">Condylarth</span> Grouping of extinct mammals

Condylarthra is an informal group – previously considered an order – of extinct placental mammals, known primarily from the Paleocene and Eocene epochs. They are considered early, primitive ungulates. It is now largely considered to be a wastebasket taxon, having served as a dumping ground for classifying ungulates which had not been clearly established as part of either Perissodactyla or Artiodactyla, being composed thus of several unrelated lineages.

<span class="mw-page-title-main">Mesonychia</span> Extinct taxon of carnivorous ungulates

Mesonychia is an extinct taxon of small- to large-sized carnivorous ungulates related to artiodactyls. Mesonychians first appeared in the early Paleocene, went into a sharp decline at the end of the Eocene, and died out entirely when the last genus, Mongolestes, became extinct in the early Oligocene. In Asia, the record of their history suggests they grew gradually larger and more predatory over time, then shifted to scavenging and bone-crushing lifestyles before the group became extinct.

Some people do not eat various specific foods and beverages in conformity with various religious, cultural, legal or other societal prohibitions. Many of these prohibitions constitute taboos. Many food taboos and other prohibitions forbid the meat of a particular animal, including mammals, rodents, reptiles, amphibians, fish, molluscs, crustaceans and insects, which may relate to a disgust response being more often associated with meats than plant-based foods. Some prohibitions are specific to a particular part or excretion of an animal, while others forgo the consumption of plants or fungi.

<i>Phenacodus</i> Genus of mammals (fossil)

Phenacodus is an extinct genus of mammals from the late Paleocene through middle Eocene, about 55 million years ago. It is one of the earliest and most primitive of the ungulates, typifying the family Phenacodontidae and the order Perissodactyla.

<span class="mw-page-title-main">Panperissodactyla</span> Clade of mammals

Panperissodactyla is a clade of ungulates containing living order Perissodactyla and all extinct ungulates more closely related to Perissodactyla than to Artiodactyla.

<i>Mesonyx</i> Extinct genus of mammals

Mesonyx is a genus of extinct mesonychid mesonychian mammal: fossils of the various species are found in Early to Late Eocene-age strata in the United States and Early Eocene-aged strata in China, 51.8—51.7 Ma (AEO).

The Islamic dietary laws (halal) and the Jewish dietary laws are both quite detailed, and contain both points of similarity and discord. Both are the dietary laws and described in distinct religious texts: an explanation of the Islamic code of law found in the Quran and Sunnah and the Jewish code of laws found in the Torah, Talmud and Shulchan Aruch.

Foot rot, also known as foul-in-the-foot, interdigital necrobacillosis or infectious pododermatitis, is a hoof infection commonly found in sheep, goats, and cattle. As the name suggests, it rots away the foot of the animal, more specifically the area between the two toes of the affected animal. It is extremely painful and contagious. It can be treated with a series of medications, but if not treated, the whole herd can become infected. The cause of the infection in cattle is two species of anaerobic bacteria, Fusobacterium necrophorum and Bacteroides melaninogenicus. Both bacteria are common to the environment in which cattle live, and Fusobacterium is present in the rumen and fecal matter of the cattle. In sheep, F. necrophorum first invades the interdigital skin following damage to the skin, and causes interdigital lesions and slight inflammation. The second stage of the disease is marked by the invasion of the foot by the foot rot bacterium Dichelobacter nodosus, a Gram-negative anaerobe. Usually, an injury to the skin between the hooves allows the bacteria to infect the animal. Another cause of foot rot may be high temperatures or humidity, causing the skin between the hooves to crack and let the bacteria infect the foot. This is one of the reasons foot rot is such a major problem in the summer. Foot rot is easily identifiable by its appearance and foul odor. Treatment is usually with an antibiotic medication, and preventing injury to the feet is the best way to prevent foot rot.

<span class="mw-page-title-main">Kosher animals</span> Animals that comply with Jewish regulations for consumption

Kosher animals are animals that comply with the regulations of kashrut and are considered kosher foods. These dietary laws ultimately derive from various passages in the Torah with various modifications, additions and clarifications added to these rules by halakha. Various other animal-related rules are contained in the 613 commandments.

<i>Agriochoerus</i> Extinct genus of mammals

Agriochoerus is an extinct genus of scansorial herbivore of the tylopod family Agriochoeridae, endemic to North America. Agriochoerus and other agriochoerids possessed claws, which is rare within Artiodactyla, as well as likely being scansorial. Agriochoerus was first described in 1869.

References

  1. American Museum of Natural History (1892). Visitors' Guide to the Geological and Palaeontological Collections. Original from the University of Michigan. p.  59. Cloven hoof -wikipedia Artiodactyla.{{cite book}}: CS1 maint: location missing publisher (link)
  2. Kate Hepworth, Animal Sciences Student; Michael Neary, Extension Animal Scientist; Simon Kenyon, Extension Veterinarian (October 2004). "Hoof Anatomy, Care and Management in Livestock" (PDF). Purdue University. Archived from the original (PDF) on 2008-04-10. Retrieved 2007-12-02.
  3. Jackson, Brenda (1998). North American Wildlife (Revised and Updated). Reader's Digest. p. 68. ISBN   0-7621-0020-6.
  4. Streubel, Donald (2000). "Oreamnos americanus (Mountain Goat)" (Web). Idaho Museum of Natural History. Retrieved 2007-12-02.
  5. Plekon, Hannah J. (April 2007). "Oreamnos americanus; General description". Davidson College. Archived from the original (Web) on 2007-12-30. Retrieved 2007-12-03.
  6. British Museum (Natural History) (1906). British Museum Guides: Vertebrates. Original from the University of Michigan. p. 28.{{cite book}}: CS1 maint: location missing publisher (link)
  7. Cleland, Herdman Fitzgerald (1916). Geology, Physical and Historical. Original from the University of Michigan: American Book Company. p.  599. Cloven hoof -wikipedia Artiodactyla.
  8. Leviticus 11:3–8
  9. Shulchan Aruch, Yoreh De'ah 79
  10. Slifkin, Natan (March 1, 2004). "6". Shafan– The Hyrax (PDF). pp. 99–125. ISBN   1-56871-312-6. Archived from the original (PDF) on February 11, 2012. Retrieved May 10, 2016.{{cite book}}: |work= ignored (help) ISBN   978-1-56871-312-0.
  11. Glover, Alfred Kingsley (1900). Jewish Laws and Customs: Some of the Laws and Usages of the Children of the Ghetto. Original from Harvard University: W.A. Hammond. p.  157. kosher cloven.
  12. Eisenberg, Ronald L. (2005). The 613 Mitzvot: A Contemporary Guide to the Commandments of Judaism. Schreiber Publishing, Incorporated. p. 251. ISBN   0-88400-303-5.