The shell of Stenotrema florida, a land snail. The periostracum is an organic layer of protein which, in this species, is developed into minute hairs, giving the snail a velvety look and feel
Conchiolins (sometimes referred to as conchins) are complex proteins which are secreted by a mollusc's outer epithelium (the mantle).
These proteins are part of a matrix of organic macromolecules, mainly proteins and polysaccharides, that assembled together form the microenvironment where crystals nucleate and grow. This organic matrix also holds and binds to the crystals of aragonite which give such shells their stiffness.
The ions necessary to form calcium carbonate are also secreted by the mantle, but it is the tailored environment created by the organic matrix which causes aragonite (rather than calcite) crystals to nucleate, in much the same way that collagen nucleates hydroxyapatite crystals.
Conchiolin serves as a relatively flexible, crack-deflecting matrix for the mineral aggregate particles; its strength and the strong bonding of perlucin can in some cases (such as in the formation of nacre) give the finished material an impressive level of toughness.
As well as providing a matrix into which the hard calcium carbonate part of the shell is precipitated, many species of mollusks (such as the land snail shown above) also have an outer shell layer called the periostracum which is composed of the protein conchiolin. Some land snails (especially the taxa that have become adapted to living on acidic soils) have very thin, transparent, tan-colored shells, even as adults, and those shells are composed entirely of conchiolin.
Limestone is a type of carbonate sedimentary rock which is the main source of the material lime. It is composed mostly of the minerals calcite and aragonite, which are different crystal forms of CaCO3. Limestone forms when these minerals precipitate out of water containing dissolved calcium. This can take place through both biological and nonbiological processes, though biological processes, such as the accumulation of corals and shells in the sea, have likely been more important for the last 540 million years. Limestone often contains fossils which provide scientists with information on ancient environments and on the evolution of life.
A pearl is a hard, glistening object produced within the soft tissue of a living shelled mollusk or another animal, such as fossil conulariids. Just like the shell of a mollusk, a pearl is composed of calcium carbonate in minute crystalline form, which has deposited in concentric layers. The ideal pearl is perfectly round and smooth, but many other shapes, known as baroque pearls, can occur. The finest quality of natural pearls have been highly valued as gemstones and objects of beauty for many centuries. Because of this, pearl has become a metaphor for something rare, fine, admirable and valuable.
Calcium carbonate is a chemical compound with the chemical formula CaCO3. It is a common substance found in rocks as the minerals calcite and aragonite, most notably in chalk and limestone, eggshells, gastropod shells, shellfish skeletons and pearls. Materials containing much calcium carbonate or resembling it are described as calcareous. Calcium carbonate is the active ingredient in agricultural lime and is produced when calcium ions in hard water react with carbonate ions to form limescale. It has medical use as a calcium supplement or as an antacid, but excessive consumption can be hazardous and cause hypercalcemia and digestive issues.
An exoskeleton is a skeleton that is on the exterior of an animal to both support the body shape and protect the internal organs, in contrast to an internal endoskeleton which is enclosed underneath other soft tissues. Some large, hard protective exoskeletons are known as "shells".
Aragonite is a carbonate mineral and one of the three most common naturally occurring crystal forms of calcium carbonate, the others being calcite and vaterite. It is formed by biological and physical processes, including precipitation from marine and freshwater environments.
Nacre, also known as mother of pearl, is an organic–inorganic composite material produced by some molluscs as an inner shell layer. It is also the material of which pearls are composed. It is strong, resilient, and iridescent.
Cultured pearls are pearls which are formed within a cultured pearl sac with human intervention in the interior of productive living molluscs in a variety of conditions depending upon the mollusc and the goals. Having the same material as natural pearls, cultured pearls can be cultivated in seawater or freshwater bodies. Nowadays, over 95% of the pearls available on the market would be cultured pearls.
The mantle is a significant part of the anatomy of molluscs: it is the dorsal body wall which covers the visceral mass and usually protrudes in the form of flaps well beyond the visceral mass itself.
Biomineralization, also written biomineralisation, is the process by which living organisms produce minerals, often resulting in hardened or stiffened mineralized tissues. It is an extremely widespread phenomenon: all six taxonomic kingdoms contain members that are able to form minerals, and over 60 different minerals have been identified in organisms. Examples include silicates in algae and diatoms, carbonates in invertebrates, and calcium phosphates and carbonates in vertebrates. These minerals often form structural features such as sea shells and the bone in mammals and birds.
An eggshell is the outer covering of a hard-shelled egg and of some forms of eggs with soft outer coats.
The periostracum is a thin, organic coating that is the outermost layer of the shell of many shelled animals, including molluscs and brachiopods. Among molluscs, it is primarily seen in snails and clams, i.e. in gastropods and bivalves, but it is also found in cephalopods such as Allonautilus scrobiculatus. The periostracum is an integral part of the shell, and it forms as the shell forms, along with the other shell layers. The periostracum is used to protect the organism from corrosion.
The gastropod shell is part of the body of a gastropod or snail, a kind of mollusc. The shell is an exoskeleton, which protects from predators, mechanical damage, and dehydration, but also serves for muscle attachment and calcium storage. Some gastropods appear shell-less (slugs) but may have a remnant within the mantle, or in some cases the shell is reduced such that the body cannot be retracted within it (semi-slug). Some snails also possess an operculum that seals the opening of the shell, known as the aperture, which provides further protection. The study of mollusc shells is known as conchology. The biological study of gastropods, and other molluscs in general, is malacology. Shell morphology terms vary by species group.
A land snail is any of the numerous species of snail that live on land, as opposed to the sea snails and freshwater snails. Land snail is the common name for terrestrial gastropod mollusks that have shells. However, it is not always easy to say which species are terrestrial, because some are more or less amphibious between land and fresh water, and others are relatively amphibious between land and salt water.
A bivalve shell is part of the body, the exoskeleton or shell, of a bivalve mollusk. In life, the shell of this class of mollusks is composed of two hinged parts or valves. Bivalves are very common in essentially all aquatic locales, including saltwater, brackish water, and freshwater. The shells of bivalves commonly wash up on beaches and along the edges of lakes, rivers, and streams. Bivalves by definition possess two shells or valves, a "right valve" and a "left valve", that are joined by a ligament. The two valves usually articulate with one another using structures known as "teeth" which are situated along the hinge line. In many bivalve shells, the two valves are symmetrical along the hinge line—when truly symmetrical, such an animal is said to be equivalved; if the valves vary from each other in size or shape, inequivalved. If symmetrical front-to-back, the valves are said to be equilateral, and are otherwise considered inequilateral.
The molluscshell is typically a calcareous exoskeleton which encloses, supports and protects the soft parts of an animal in the phylum Mollusca, which includes snails, clams, tusk shells, and several other classes. Not all shelled molluscs live in the sea; many live on the land and in freshwater.
Mollusca is the second-largest phylum of invertebrate animals, after Arthropoda; members are known as molluscs or mollusks. Around 76,000 extant species of molluscs are recognized. The number of fossil species is estimated between 60,000 and 100,000 additional species. The proportion of undescribed species is very high. Many taxa remain poorly studied.
Mineralized tissues are biological tissues that incorporate minerals into soft matrices. Typically these tissues form a protective shield or structural support. Bone, mollusc shells, deep sea sponge Euplectella species, radiolarians, diatoms, antler bone, tendon, cartilage, tooth enamel and dentin are some examples of mineralized tissues.
Shell growth in estuaries is an aspect of marine biology that has attracted a number of scientific research studies. Many groups of marine organisms produce calcified exoskeletons, commonly known as shells, hard calcium carbonate structures which the organisms rely on for various specialized structural and defensive purposes. The rate at which these shells form is greatly influenced by physical and chemical characteristics of the water in which these organisms live. Estuaries are dynamic habitats which expose their inhabitants to a wide array of rapidly changing physical conditions, exaggerating the differences in physical and chemical properties of the water.
Marine biogenic calcification is the process by which marine organisms such as oysters and clams form calcium carbonate. Seawater is full of dissolved compounds, ions and nutrients that organisms can use for energy and, in the case of calcification, to build shells and outer structures. Calcifying organisms in the ocean include molluscs, foraminifera, coccolithophores, crustaceans, echinoderms such as sea urchins, and corals. The shells and skeletons produced from calcification have important functions for the physiology and ecology of the organisms that create them.
Foraminiferal tests are the tests of Foraminifera.
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