Cyathea acanthophora | |
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Scientific classification | |
Kingdom: | Plantae |
Division: | Pteridophyta |
Class: | Pteridopsida |
Order: | Cyatheales |
Family: | Cyatheaceae |
Genus: | Cyathea |
Subgenus: | Cyathea |
Section: | Alsophila |
Species: | C. acanthophora |
Binomial name | |
Cyathea acanthophora Holttum, 1962 | |
Synonyms | |
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Cyathea acanthophora is a species of tree fern endemic to Kinabalu National Park in Borneo, where it grows in montane rain forest at an altitude of 1250–2000 m. The trunk is erect and can grow as tall as 5 m or higher. Fronds are 2–3 m in length and bi- or tripinnate. The stipe is dark towards the base and has slender spines. It is covered with scattered scales, which are glossy brown and have a narrow paler margin. Sori occur near the midvein of fertile pinnules. They are covered in thin, fragile indusia, which resemble small saucers.
The order Cyatheales, which includes the tree ferns, is a taxonomic division of the fern class, Polypodiopsida. No clear morphological features characterize all of the Cyatheales, but DNA sequence data indicate the order is monophyletic. Some species in the Cyatheales have tree-like growth forms, but others have rhizomes.
Borneo is the third-largest island in the world and the largest in Asia. At the geographic centre of Maritime Southeast Asia, in relation to major Indonesian islands, it is located north of Java, west of Sulawesi, and east of Sumatra.
A frond is a large, divided leaf. In both common usage and botanical nomenclature, the leaves of ferns are referred to as fronds and some botanists restrict the term to this group. Other botanists allow the term frond to also apply to the large leaves of cycads and palms (Arecaceae). "Frond" is commonly used to identify a large, compound leaf, but if the term is used botanically to refer to the leaves of ferns and algae it may be applied to smaller and undivided leaves.
Cyathea cooperi, also known as the Australian tree fern, lacy tree fern, scaly tree fern, or Cooper's tree fern, is a tree fern native to Australia, in New South Wales and Queensland.
Cyathea alpicola is a species of tree fern native to central Sumatra, where it grows in montane rain forest at an altitude of 2000–2750 m. The erect trunk can reach 5 m or more in height. Fronds are bi- or tripinnate and usually 2–3 m long. They have a tendency to persist on the plant after withering, forming an irregular skirt around the trunk. This species has a spiny stipe, which is covered with a woolly layer of scales. There appear to be two forms with different scales; either dark glossy brown with a broad paler margin and fragile edges, or small, brown and finely fringed. Sori are covered by thin, fragile indusia and occur near the midvein of fertile pinnules. The closest relative of C. alpicola is thought to be Cyathea polycarpa, which differs by lacking spines altogether. Cyathea macropoda and Cyathea magnifolia may also be closely allied with this species.
Cyathea apiculata is a species of tree fern native to Sumatra, where it grows in montane rain forest at an altitude of about 1800 m. The trunk is erect and usually 5 m tall or more. Fronds are bi- or tripinnate and 2–3 m long. The stipe is pale, becoming darker towards the base. It is covered with dark scales with pale, fragile edges. Sori occur near the midvein of fertile pinnules and are covered by thin, very fragile indusia.
Cyathea baileyana, also known as the wig tree fern, is a species of tree fern native to northeastern Queensland in Australia, where it grows in wet gullies and forest at an altitude of 850–1200 m. It is a rare species that is seldom found in the wild. The erect trunk is 4–5 m tall, approximately 10 cm in diameter and may be covered in stipe bases in the upper regions. C. baileyana is notable for being able to develop offshoots from the base of the trunk. Fronds are bi- or tripinnate and may be exceptionally long, up to 7 m, though they are usually around 2–3 m. The rachis and stipe are dark to darkish red, scaly and may be warty, but lack spines. Scales on the rachis and stipe are purplish brown to black and have a long hair-like apex. Characteristically of this species, the last pair of pinnae are separated from the others along the rachis and may form a clump around the trunk apex. Sori are circular and occur in one to three rows along the pinnule midvein. They lack indusia.
Cyathea brooksii is a species of tree fern native to Cuba, Hispaniola and Puerto Rico, where it grows on serpentine soils in shaded ravines, along streams, and on forested slopese at an altitude of 250–950 m. The trunk is prostrate and only about 6 cm in diameter. Fronds are pinnate or bipinnate and up to 2 m long. The base of the rachis is covered with blackish scales that have a paler margin. Sori occur in two rows, one along each side of the pinnule midvein.
Cyathea callosa is a species of tree fern endemic to Luzon in the Philippines, where it grows in midmontane forest. The trunk is erect and about 3 m tall or more. Fronds are bi- or tripinnate and 1–2 m long. The stipe is spiny and covered with scattered scales that are dark and have pale, fragile edges. Sori occur near the midvein of fertile pinnules and are covered by thin, pale indusia.
Cyathea capensis is a regionally widespread and highly variable species of tree fern. It is indigenous to Southern Africa and South America.
Cyathea caudata is a species of tree fern native to the islands of Luzon and Mindanao in the Philippines, where it grows in montane forest. The trunk is erect and up to 4 m tall or more. Fronds are bi- or tripinnate and 1–2 m long. The stipe is warty and covered with dark, narrow, glossy scales. Sori are borne near the midvein of fertile pinnules and are protected by firm, brown indusia.
Cyathea christii is a species of tree fern endemic to Mindanao in the Philippines, where it grows in forest at an altitude of 900–1800 m. The trunk is erect and may be 5 m tall or more. Fronds are bi- or tripinnate and 2–3 m long. The stipe is covered with some warts and narrow, brown scales. Sori occur near the midvein of fertile pinnules and are covered by thin, fragile indusia.
Cyathea cinerea is a species of tree fern endemic to Mindanao in the Philippines, where it grows in forest at an altitude of about 1000 m. The trunk is erect, up to 5 m tall and about 12 cm in diameter. Fronds are bi- or tripinnate and 1–2 m long. The stipe has long spines and scattered scales, which are either narrow and pale or wide and brown. Sori occur near the midvein of fertile pinnules and are covered by brown indusia.
Cyathea cunninghamii, also known as the gully tree fern and slender tree fern, is a species of tree fern indigenous to New Zealand including North Island, South Island and Chatham Islands; also to Victoria, possibly New South Wales, southeastern Queensland and Tasmania in Australia. It grows in damp forest, often emerging from stream gullies and riverbanks. Brownsey noted that it has a lower tolerance for drought than other species of Cyathea. The erect trunk may be 20 m tall and is usually 6–15 cm in diameter, occasionally as much as 20 cm. Fronds are tri- to tetrapinnate and 3 m or more in length. The rachis and stipe are slender, black brown, warty and covered with brown scales. Sori occur along each side of the pinnule midvein and are covered by hood-like indusia. C. cunninghamii is an uncommon and slow-growing tree fern.
Cyathea dregei is a widespread species of tree fern in southern Africa.
Cyathea edanoi is a species of tree fern endemic to Luzon in the Philippines, where it grows in montane forest at an altitude of about 1300 m. The trunk is erect and 1–2 m tall. Fronds are bi- or tripinnate and approximately 1 m long. The stipe is covered in dark, glossy scales that have narrow, fragile edges. Sori occur near the midvein of fertile pinnules and are covered, sometimes half-covered, by large, firm, brown indusia.
Cyathea squamulata is a species of tree fern native to the Malay Peninsula, Sumatra, Java, Borneo and the southern Philippines, including the Sulu Archipelago, where it grows in forest from the lowlands to an altitude of about 1500 m. The trunk is erect and up to 2 m tall. Fronds are pinnate or bipinnate and approximately 1.5 m long. The stipe is covered in densely packed firm, medium brown scales. Sori occur near the fertile pinnule midvein and lack indusia.
Cyathea × marcescens, commonly known as the skirted tree fern, is a tree fern endemic to the Cape Otway ranges in Victoria and Tasmania, Australia. It is a natural hybrid, apparently Cyathea australis × Cyathea cunninghamii. Braggins and Large (2004) note that it has characteristics midway between these two species. The spores of C. × marcescens are usually malformed although sterile. The trunk of this plant is erect and up to 10 m tall. Fronds may be bi- or tripinnate and 3–4 m in length. Dead fronds often persist, forming a characteristic skirt around the trunk. The stipe is thick, black and warty. The rachis and trunk are covered in shiny, dark brown scales. Sori are borne near the fertile pinnule midvein and are protected by thin indusia that are saucer-like in appearance.
Cyathea hotteana is a species of tree fern endemic to Haiti, where it grows in cloud forest at an altitude of 1000–1200 m. The trunk of this plant is erect, up to 2 m tall, and approximately 8 cm in diameter. Fronds are pinnate and may reach 2 m in length. Brown basal scales cover the rachis and stipe, which are also brown. Sori are borne in two rows, one on each side of the pinnule midvein. They are covered by shallow, plate-like indusia, which bear one or two clefts at the margin.
Alsophila is a section in the subgenus Cyathea. The name was originally applied to a genus of tree ferns which is now considered to be synonymous with Cyathea.
Cyathea glauca is a species of tree fern endemic to Réunion. Little is known about this species. C. glauca is not to be confused with Cyathea glauca used as a synonym of Cyathea mexicana.
Cyathea smithii, commonly known as the soft tree fern or kātote, is a species of tree fern from New Zealand.
The International Standard Book Number (ISBN) is a numeric commercial book identifier which is intended to be unique. Publishers purchase ISBNs from an affiliate of the International ISBN Agency.
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