Cyathea cooperi, also known as the Australian tree fern, lacy tree fern, scaly tree fern, or Cooper's tree fern, is a tree fern native to Australia, in New South Wales and Queensland.
The Cyatheaceae are the scaly tree fern family and include the world's tallest tree ferns, which reach heights up to 20 m. They are also very ancient plants, appearing in the fossil record in the late Jurassic, though the modern genera likely appeared in the Cenozoic. Cyatheaceae are the largest family of tree ferns, including about 500 species. Cyatheaceae and Dicksoniaceae, together with Metaxyaceae and Cibotiaceae, are a monophyletic group and constitute the "core tree ferns". Cyatheaceae are leptosporangiate ferns, the most familiar group of monilophytes.
Cyathea medullaris, popularly known as the black tree fern, is a large tree fern up to 20 m tall. It is distributed across the south-west Pacific from Fiji to Pitcairn and New Zealand. It is called mamaku, katātā, kōrau, or pītau in the Māori language.
Cyathea affinis is a variable species of tree fern native to Fiji, Samoa, the Cook Islands, Austral Islands, Tahiti, and the Marquesas Islands. The trunk of this plant is erect and 2–6 m tall. Fronds are bipinnate and 2–3 m in length. The rachis and stipe are pale to brown in colour, or flushed with red towards the pinnule rachis. The stipe is sparsely covered in narrow basal scales, which are pale to dark and have broad fragile edges. Characteristically of this species, the lowest one or two pairs of pinnae may be slightly reduced and occur towards the base of the stipe. Sori are located near the pinnule midvein and are partially or fully covered by indusia, which open towards the pinnule margin.
Cyathea australis, also known as the Rough Tree Fern, is a species of tree fern native to southeastern Queensland, New South Wales and southern Victoria in Australia, as well as Tasmania and Norfolk Island.
Cyathea brevipinna is a species of tree fern endemic to the higher parts of Mount Gower on Lord Howe Island, where it grows in exposed areas at an altitude of about 790 m. The trunk is erect and may reach 3 m in height. It is often covered with reddish brown scales and stipe bases. This species may produce stolons at ground level. Fronds are tripinnate, densely crowded, and up to about 3 m long. The stipe is brown and sometimes warty after scales fall off. The scales are long, glossy dark brown, with a distinctly narrow apex and fragile paler edges. Sori are attached to deeply divided fertile pinnules that may uncurl over the sori. Indusia are firm and large. C. brevipinna is a stunted plant with short pinnae.
Cyathea brooksii is a species of tree fern native to Cuba, Hispaniola and Puerto Rico, where it grows on serpentine soils in shaded ravines, along streams, and on forested slopese at an altitude of 250–950 m. The trunk is prostrate and only about 6 cm in diameter. Fronds are pinnate or bipinnate and up to 2 m long. The base of the rachis is covered with blackish scales that have a paler margin. Sori occur in two rows, one along each side of the pinnule midvein.
Cyathea capensis is a regionally widespread and highly variable species of tree fern. It is indigenous to Southern Africa and South America.
Cyathea costulisora is a species of tree fern native to Sumatra. The trunk is erect and usually 1–4 m tall. Fronds are bi- or tripinnate and 1–2 m long. The stipe is covered with warts and scales. The scales are either pale and glossy or dark and flat. Sori occur near the fertile pinnule midvein and are covered by large, firm, brown indusia.
Cyathea dregei is a widespread species of tree fern in southern Africa.
Cyathea gigantea is a species of tree fern native to northeastern to southern India, Sri Lanka, Nepal to Myanmar, Thailand, Laos, Vietnam, the Malay Peninsula, as well as central Sumatra and western Java. It grows in moist open areas at an altitude of 600–1000 m. The trunk of this species is erect and may be as tall as 5 m or more. Fronds are bi- or tripinnate and usually 2–3 m long. The rachis is long, dark to black in colouration and rough in appearance after the fall of scales. These scales are dark brown, glossy and have a narrow paler margin and fragile edges. Sori are round and indusia absent.
Cyathea glabra is a species of tree fern native to Borneo, western Java, Sumatra and the Malay Peninsula, where it grows in lowland swamp forest and montane forest at an elevation of up to 1500 m. The trunk of this plant is erect and 2–4 m tall. Fronds are bi- or tripinnate and 1–2 m in length. Characteristically of this species, the lowest pinnae may be significantly reduced. The stipe is very dark and bears basal scales. These scales are dark, glossy and have a paler margin and fragile edges. Sori are produced in groups of one to three on fertile pinnule veins. They lack indusia.
Cyathea hancockii is a species of tree fern native to the Ryukyu Islands, Japan, Taiwan and Hong Kong. The specific epithet hancockii commemorates William Hancock (1847-1914), who collected numerous plants in Japan, China and Southeast Asia. It grows in forest, on stream banks, and in forest margins at an elevation of about 600 m or higher.
Cyathea henryi is a species of tree fern native to India and Yunnan in China, where it grows submontane and montane forest at an altitude of 600–1200 m. The trunk of this plant is erect and may be 5–7 m tall or more. Fronds are bi- or tripinnate and usually 2–3 m in length. The rachis is smooth and dark, but occasionally has a few scattered scales. The stipe also has these scales. The scales are either small and pale with irregular fringed edges, or large and dark with a paler margin. Sori are borne on minor veins and lack indusia.
Cyathea heterochlamydea is a little-known species of tree fern native to the islands of Luzon, Panay, Negros and Mindanao in the Philippines, where it grows in montane forest. The trunk of this plant is erect and usually up to 4 m tall or more. Fronds may be bi- or tripinnate and 1–2 m in length. The stipe is warty and/or bears short spines and scales. These scales are dark, glossy and have a narrow pale margin. Sori are borne near the fertile pinnule midvein and are protected by firm, brown indusia.
Cyathea × marcescens, commonly known as the skirted tree fern, is a tree fern endemic to the Cape Otway ranges in Victoria and Tasmania, Australia. It is a natural hybrid, apparently Cyathea australis × Cyathea cunninghamii. Braggins and Large (2004) note that it has characteristics midway between these two species. The spores of C. × marcescens are usually malformed although sterile. The trunk of this plant is erect and up to 10 m tall. Fronds may be bi- or tripinnate and 3–4 m in length. Dead fronds often persist, forming a characteristic skirt around the trunk. The stipe is thick, black and warty. The rachis and trunk are covered in shiny, dark brown scales. Sori are borne near the fertile pinnule midvein and are protected by thin indusia that are saucer-like in appearance.
Cyathea imbricata is a species of tree fern endemic to Western New Guinea, where it grows in open forest at an altitude of 3240 m. The trunk of this plant is erect and approximately 2 m tall. Fronds may be bi- or tripinnate and are usually less than 1 m in length. The stipe is dark, spiny, and covered with caducous scales. These scales are glossy brown in colouration and have a paler margin and fragile edges. Sori are borne in groups of one to four per pinnule lobe. They are protected by firm indusia.
Cyathea javanica is a species of tree fern native to western Java and Sumatra, in Indonesia.
Cyathea latebrosa is a common and widespread species of tree fern native to Indochina. Its natural range covers Cambodia and Thailand, and stretches from the Malay Peninsula to Indonesia, where it is present on the islands of Borneo and Sumatra. Plants reported from India and Sri Lanka have thinner indusia and may represent a separate, as-yet undescribed, species. C. latebrosa grows in a wide range of habitats, including forest, secondary forest, and plantations, from sea level up to an elevation of about 1500 m.
Cyathea smithii, commonly known as the soft tree fern or kātote, is a species of tree fern from New Zealand.
