Cyathium

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Euphorbia tridentata, cross-section of a cyathium. Cyathium cross2 ies no letters.jpg
Euphorbia tridentata, cross-section of a cyathium.
In euphorbia milii-large Stamen of euphorbia milii.jpg
In euphorbia milii-large

A cyathium (pl.: cyathia) is one of the specialised pseudanthia ("false flowers") forming the inflorescence of plants in the genus Euphorbia (Euphorbiaceae). A cyathium consists of:

The flower-like characteristics of the cyathia are underlined by brightly coloured nectar glands and often by petal-like appendages to the nectar glands, or brightly coloured, petal-like bracts positioned under the cyathia. The paired petal-like bracts of Euphorbia section Goniostema are called cyathophylls. here female to male flower ratio is 1:α

The cyathia are sometimes solitary, but are usually in cymes, inflorescences of the second order, in pseudumbels, on dichotomously branched stalks or in so-called simple cymes which consist of one central and two lateral cyathia.

In one group of Madagascan species in the subfamily Euphorbia section Goniostema (E. aueoviridiflora, E. capmanambatoensis, E. iharanae, E. leuconeura, E. neohumbertii, E. viguieri) there is a tendency for a further pseudanthium to grow from the cyme. Probably as an adaptation to pollination by birds, the cyathia have become specialised: Most cyathia have upright cyathophylls which surround them protectively, but render the nectar glands inaccessible. To compensate, between them are naked sterile cyathia whose only job is to produce nectar.

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Euphorbia revoluta is a species of euphorb known by the common name threadstem sandmat. It is native to Mexico and the southwestern United States from California to the Rocky Mountains. It is an annual herb producing thin, erect stems with pairs of linear leaves, each leaf up to 2.6 centimeters long. The inflorescence is a cyathium with rounded nectar glands surrounding one female flower and several male flowers. There are sometimes white petal-like appendages as well. The Navajo used this plant as a skin lotion.

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