Freshwater molluscs are those members of the Phylum Mollusca which live in freshwater habitats, both lotic (flowing water) such as rivers, streams, canals, springs, and cave streams (stygobite species) and lentic (still water) such as lakes, ponds (including temporary or vernal ponds), and ditches.
This article is about freshwater Mollusca in general; for information on one particular family of freshwater molluscs, please follow the appropriate links in the lists below.
The two major classes of molluscs have representatives in freshwater: the gastropods (snails) and the bivalves (freshwater mussels and clams.) It appears that the other classes within the Phylum Mollusca -the cephalopods, scaphopods, polyplacophorans, etc. - never made the transition from a fully marine environment to a freshwater environment.
A few species of freshwater molluscs are among the most notorious invasive species. In contrast, numerous others have become threatened or have become extinct in the face of anthropogenic change.
Typical freshwater species (such as many river mussel species in the family Unionidae) have a range which may consist of a series of adjacent river systems, a series of adjacent tributaries, or part of a single large river system. Large rivers and small tributary creeks typically share few species, and distribution patterns suggest large lowland rivers represent substantial barriers to the dispersal of species adapted to small upland streams. Endemism is common in some families, and species may be endemic to a single creek or spring. In contrast, some of the tiny pill clams have a nearly worldwide distribution (Burch, 1972) [1]
Challenges in the natural environment faced by freshwater Mollusca include floods, droughts, siltation, extreme temperature variations, predation, and the constant unidirectional flow characteristic of river habitats. Osmoregulation, or the maintenance of constant salinity within body tissue and fluids, is another challenge faced by freshwater Mollusca. Dillon (2000) [2] indicates that they have characteristically low tissue salinities relative to other freshwater animals, and unionid mussels have some of the lowest tissue salinities of any animal.
Families of freshwater bivalves occur within the orders Unionida and Venerida.
Ten families of prosobranchiate snails (gilled operculate snails) and five pulmonate families (lunged snails, distantly related to common landsnails) inhabit freshwater environments in many parts of the world. Some freshwater snail species serve as hosts for human and animal parasites.
Shellfish is a colloquial and fisheries term for exoskeleton-bearing aquatic invertebrates used as food, including various species of molluscs, crustaceans, and echinoderms. Although most kinds of shellfish are harvested from saltwater environments, some are found in freshwater. In addition, a few species of land crabs are eaten, for example Cardisoma guanhumi in the Caribbean. Shellfish are among the most common food allergens.
Mussel is the common name used for members of several families of bivalve molluscs, from saltwater and freshwater habitats. These groups have in common a shell whose outline is elongated and asymmetrical compared with other edible clams, which are often more or less rounded or oval.
Bivalvia, in previous centuries referred to as the Lamellibranchiata and Pelecypoda, is a class of marine and freshwater molluscs that have laterally compressed bodies enclosed by a shell consisting of two hinged parts. Bivalves as a group have no head and they lack some usual molluscan organs like the radula and the odontophore. They include the clams, oysters, cockles, mussels, scallops, and numerous other families that live in saltwater, as well as a number of families that live in freshwater. The majority are filter feeders. The gills have evolved into ctenidia, specialised organs for feeding and breathing. Most bivalves bury themselves in sediment where they are relatively safe from predation. Others lie on the sea floor or attach themselves to rocks or other hard surfaces. Some bivalves, such as the scallops and file shells, can swim. The shipworms bore into wood, clay, or stone and live inside these substances.
A siphon is an anatomical structure which is part of the body of aquatic molluscs in three classes: Gastropoda, Bivalvia and Cephalopoda.
The Unionidae are a family of freshwater mussels, the largest in the order Unionida, the bivalve molluscs sometimes known as river mussels, or simply as unionids.
The periostracum is a thin organic coating or "skin" which is the outermost layer of the shell of many shelled animals, including molluscs and brachiopods. Among molluscs it is primarily seen in snails and clams, i.e. in gastropods and bivalves, but it is also found in cephalopods such as Allonautilus scrobiculatus. Periostracum is an integral part of the shell, and it forms as the shell forms, along with the other shell layers.
A molluscivore is a carnivorous animal that specialises in feeding on molluscs such as gastropods, bivalves, brachiopods and cephalopods. Known molluscivores include numerous predatory molluscs,, arthropods such as crabs and firefly larvae, and, vertebrates such as fish, birds and mammals. Molluscivory is performed in a variety ways with some animals highly adapted to this method of feeding behaviour. A similar behaviour, durophagy, describes the feeding of animals that consume hard-shelled or exoskeleton bearing organisms, such as corals, shelled molluscs, or crabs.
Unio crassus, the thick shelled river mussel, is a species of freshwater mussel, an aquatic bivalve mollusk in the family Unionidae, the river mussels.
The freshwater pearl mussel is an endangered species of freshwater mussel, an aquatic bivalve mollusc in the family Margaritiferidae.
Physella acuta is a species of small, left-handed or sinistral, air-breathing freshwater snail, an aquatic gastropod mollusk in the family Physidae. Common names include European physa, tadpole snail, bladder snail, and acute bladder snail. In addition, Physa acuta, Physa heterostropha and Physa integra are synonyms of Physella acuta.
Sinanodonta woodiana, the Chinese pond mussel, Eastern Asiatic freshwater clam or swan-mussel, is a species of freshwater mussel, an aquatic bivalve mollusk in the family Unionidae, the river mussels.
Unionida is a monophyletic order of freshwater mussels, aquatic bivalve molluscs. The order includes most of the larger freshwater mussels, including the freshwater pearl mussels. The most common families are the Unionidae and the Margaritiferidae. All have in common a larval stage that is temporarily parasitic on fish, nacreous shells, high in organic matter, that may crack upon drying out, and siphons too short to permit the animal to live deeply buried in sediment.
The triangular kidneyshell is a species of freshwater mussel, in the family Unionidae, the river mussels. It is endemic to Alabama in the United States, where it is known from several rivers and streams in the Mobile River Basin. It is a federally listed endangered species of the United States.
Freshwater bivalves are one kind of freshwater molluscs, along with freshwater snails. They are bivalves which live in freshwater, as opposed to saltwater, the main habitat type for bivalves.
Marine invertebrates are the invertebrates that live in marine habitats. Invertebrate is a blanket term that includes all animals apart from the vertebrate members of the chordate phylum. Invertebrates lack a vertebral column, and some have evolved a shell or a hard exoskeleton. As on land and in the air, marine invertebrates have a large variety of body plans, and have been categorised into over 30 phyla. They make up most of the macroscopic life in the oceans.
Mollusca is the second-largest phylum of invertebrate animals after the arthropoda. The members are known as molluscs or mollusks. Around 85,000 extant species of molluscs are recognized. The number of fossil species is estimated between 60,000 and 100,000 additional species. The proportion of undescribed species is very high. Many taxa remain poorly studied.
Villosa arkansasensis, the Ouachita creekshell, is a species of freshwater mussel, an aquatic bivalve mollusc in the family Unionidae, the river mussels. It is endemic to certain rivers and streams in the Ouachita Mountains, Arkansas, and has a complex life history including its larvae being parasitic on a fish host.
Trichomya is a monotypic genus of marine bivalve molluscs in the family Mytilidae, the mussels. The only species is Trichomya hirsuta which is endemic to southern and eastern Australia. Its common names include the hairy mussel, the greenling and the kelp greenling.