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Ungulates are members of the diverse clade Ungulata which primarily consists of large mammals with hooves. Living ungulates are divided into two orders: the odd-toed ungulates (Perissodactyla) including horses, rhinoceroses, and tapirs; and even-toed ungulates (Artiodactyla) such as cattle, pigs, giraffes, camels, sheep, deer, and hippopotamuses. Cetaceans such as whales, dolphins, and porpoises are also classified as even-toed ungulates, although they do not have hooves. Most terrestrial ungulates use the hoofed tips of their toes to support their body weight while standing or moving. Two other orders of ungulates, Notoungulata, and Litopterna, native to South America, became extinct at the end of the Pleistocene, around 12,000 years ago.
Horse teeth refers to the dentition of equine species, including horses and donkeys. Equines are both heterodontous and diphyodontous, which means that they have teeth in more than one shape, and have two successive sets of teeth, the deciduous and permanent sets.
Dentition pertains to the development of teeth and their arrangement in the mouth. In particular, it is the characteristic arrangement, kind, and number of teeth in a given species at a given age. That is, the number, type, and morpho-physiology of the teeth of an animal.
The molars or molar teeth are large, flat teeth at the back of the mouth. They are more developed in mammals. They are used primarily to grind food during chewing. The name molar derives from Latin, molaris dens, meaning "millstone tooth", from mola, millstone and dens, tooth. Molars show a great deal of diversity in size and shape across mammal groups. The third molar of humans is sometimes vestigial.
Ferugliotherium is a genus of fossil mammals in the family Ferugliotheriidae from the Campanian and/or Maastrichtian period of Argentina. It contains a single species, Ferugliotherium windhauseni, which was first described in 1986. Although originally interpreted on the basis of a single brachydont (low-crowned) molar as a member of Multituberculata, an extinct group of small, rodent-like mammals, it was recognized as related to the hypsodont (high-crowned) Sudamericidae following the discovery of additional material in the early 1990s. After a jaw of the sudamericid Sudamerica was described in 1999, these animals were no longer considered to be multituberculates and a few fossils that were previously considered to be Ferugliotherium were assigned to unspecified multituberculates instead. Since 2005, a relationship between gondwanatheres and multituberculates has again received support. A closely related animal, Trapalcotherium, was described in 2009 on the basis of a single tooth.
Lavanify is a mammalian genus from the late Cretaceous of Madagascar. The only species, L. miolaka, is known from two isolated teeth, one of which is damaged. The teeth were collected in 1995–1996 and described in 1997. The animal is classified as a member of Gondwanatheria, an enigmatic extinct group with unclear phylogenetic relationships, and within Gondwanatheria as a member of the family Sudamericidae. Lavanify is most closely related to the Indian Bharattherium; the South American Sudamerica and Gondwanatherium are more distantly related. Gondwanatheres probably ate hard plant material.
Elasmotherium is an extinct genus of large rhinoceros endemic to Eurasia during Late Miocene through the Pleistocene, existing at least as recently as 39,000 years ago in the Late Pleistocene. A more recent date of 26,000 BP is considered less reliable. It was the last surviving member of Elasmotheriinae, a distinctive group of rhinoceroses separate from the group that contains living rhinoceros (Rhinocerotinae). The two groups are estimated to have split at least 35 million years ago according to fossils and molecular evidence.
Ferugliotheriidae is one of three known families in the order Gondwanatheria, an enigmatic group of extinct mammals. Gondwanatheres have been classified as a group of uncertain affinities or as members of Multituberculata, a major extinct mammalian order. The best-known representative of Ferugliotheriidae is the genus Ferugliotherium from the Late Cretaceous epoch in Argentina. A second genus, Trapalcotherium, is known from a single tooth, a first lower molariform, from a different Late Cretaceous Argentinean locality. Another genus known from a single tooth, Argentodites, was first described as an unrelated multituberculate, but later identified as possibly related to Ferugliotherium. Finally, a single tooth from the Paleogene of Peru, LACM 149371, perhaps a last upper molariform, and a recent specimen from Mexico, may represent related animals.
Toxodontia is a suborder of the meridiungulate order Notoungulata. Most of the members of the five included families, including the largest notoungulates, share several dental, auditory and tarsal specializations. The group is named after Toxodon, the first example of the group to be discovered by science.
Veterinary dentistry is the field of dentistry applied to the care of animals. It is the art and science of prevention, diagnosis, and treatment of conditions, diseases, and disorders of the oral cavity, the maxillofacial region, and its associated structures as it relates to animals.
A tooth is a hard, calcified structure found in the jaws of many vertebrates and used to break down food. Some animals, particularly carnivores and omnivores, also use teeth to help with capturing or wounding prey, tearing food, for defensive purposes, to intimidate other animals often including their own, or to carry prey or their young. The roots of teeth are covered by gums. Teeth are not made of bone, but rather of multiple tissues of varying density and hardness that originate from the outermost embryonic germ layer, the ectoderm.
Mesowear is a method, used in different branches and fields of biology. This method can apply to both extant and extinct animals, according to the scope of the study. Mesowear is based on studying an animal's tooth wearing fingerprint. In brief, each animal has special feeding habits, which cause unique tooth wearing. Rough feeds cause serious tooth abrasion, while smooth one triggers moderate abrasion, so browsers have teeth with moderate abrasion and grazers have teeth with rough abrasion. Scoring systems can quantify tooth abrasion observations and ease comparisons between individuals.
Many different terms have been proposed for features of the tooth crown in mammals.
Teeth are common to most vertebrates, but mammalian teeth are distinctive in having a variety of shapes and functions. This feature first arose among early therapsids during the Permian, and has continued to the present day. All therapsid groups with the exception of the mammals are now extinct, but each of these groups possessed different tooth patterns, which aids with the classification of fossils.
Bharattherium is a mammal that lived in India during the Maastrichtian and possibly the Paleocene. The genus has a single species, Bharattherium bonapartei. It is part of the gondwanathere family Sudamericidae, which is also found in Madagascar and South America during the latest Cretaceous. The first fossil of Bharattherium was discovered in 1989 and published in 1997, but the animal was not named until 2007, when two teams independently named the animal Bharattherium bonapartei and Dakshina jederi. The latter name is now a synonym. Bharattherium is known from a total of eight isolated fossil teeth, including one incisor and seven molariforms.
The grit, not grass hypothesis is an evolutionary hypothesis that explains the evolution of high-crowned teeth, particularly in New World mammals. The hypothesis is that the ingestion of gritty soil is the primary driver of hypsodont tooth development, not the silica-rich composition of grass, as was previously thought.
Patagonia is an extinct genus of non-placental mammal from the Miocene of Argentina. Traditionally considered a metatherian incertae sedis, one analysis suggested it to be a gondwanathere. However, this has been rejected by other authors.
Dinosaur teeth have been studied since 1822 when Mary Ann Mantell (1795-1869) and her husband Dr Gideon Algernon Mantell (1790-1852) discovered an Iguanodon tooth in Sussex in England. Unlike mammal teeth, individual dinosaur teeth are generally not considered by paleontologists to be diagnostic to the genus or species level for unknown taxa, due morphological convergence and variability between teeth. and many historically named tooth taxa like Paronychodon and Richardoestesia are today considered nomina dubia, and are used as form taxa to refer to isolated teeth from other localities displaced considerably in time and space from the type specimens. However, it is possible to refer isolated teeth to known taxa provided that the tooth morphology is known and the teeth originate from a similar time and place.
Argyrohippus is an extinct genus of notoungulate, belonging to the family Notohippidae. It lived from the Late Oligocene to the Early Miocene, and its fossilized remains were found in South America.
Eomorphippus is an extinct genus of notohippid notoungulate that lived from the Late Eocene to the Early Oligocene in what is today South America.
References
- ↑ Phillip E. Jardine, Christine M. Janis, Sarda Sahney, Michael J. Benton. “Grit not grass: Concordant patterns of early origin of hypsodonty in Great Plains ungulates and Glires.” Palaeogeography, Palaeoclimatology, Palaeoecology. December 2012:365–366, 1–10
- ↑ Flynn, John J., André R. Wyss, and Reynaldo Charrier. “South America’s Missing Mammals.” Scientific American. May 2007: 68-75.
- ↑ Kwan, Paul W.L. (2007). "Digestive system I" (PDF). Tufts University. Archived from the original (PDF) on 20 October 2013. Retrieved 4 July 2019.
- ↑ Gondwanatheria