Dentition pertains to the development of teeth and their arrangement in the mouth. In particular, it is the characteristic arrangement, kind, and number of teeth in a given species at a given age. [1] That is, the number, type, and morpho-physiology (that is, the relationship between the shape and form of the tooth in question and its inferred function) of the teeth of an animal. [2]
Animals whose teeth are all of the same type, such as most non-mammalian vertebrates, are said to have homodont dentition, whereas those whose teeth differ morphologically are said to have heterodont dentition. The dentition of animals with two successions of teeth (deciduous, permanent) is referred to as diphyodont , while the dentition of animals with only one set of teeth throughout life is monophyodont. The dentition of animals in which the teeth are continuously discarded and replaced throughout life is termed polyphyodont . [2] The dentition of animals in which the teeth are set in sockets in the jawbones is termed thecodont.
The evolutionary origin of the vertebrate dentition remains contentious. Current theories suggest either an "outside-in" or "inside-out" evolutionary origin to teeth, with the dentition arising from odontodes on the skin surface moving into the mouth, or vice versa. [3] Despite this debate, it is accepted that vertebrate teeth are homologous to the dermal denticles found on the skin of basal Gnathostomes (i.e. Chondrichtyans). [4] Since the origin of teeth some 450 mya, the vertebrate dentition has diversified within the reptiles, amphibians, and fish: however most of these groups continue to possess a long row of pointed or sharp-sided, undifferentiated teeth (homodont) that are completely replaceable. The mammalian pattern is significantly different. The teeth in the upper and lower jaws in mammals have evolved a close-fitting relationship such that they operate together as a unit. "They 'occlude', that is, the chewing surfaces of the teeth are so constructed that the upper and lower teeth are able to fit precisely together, cutting, crushing, grinding or tearing the food caught between." [5]
Mammals have up to four distinct types of teeth, though not all types are present in all mammals. These are the incisor (cutting), the canine, the premolar, and the molar (grinding). The incisors occupy the front of the tooth row in both upper and lower jaws. They are normally flat, chisel-shaped teeth that meet in an edge-to-edge bite. Their function is cutting, slicing, or gnawing food into manageable pieces that fit into the mouth for further chewing. The canines are immediately behind the incisors. In many mammals, the canines are pointed, tusk-shaped teeth, projecting beyond the level of the other teeth. In carnivores, they are primarily offensive weapons for bringing down prey. In other mammals such as some primates, they are used to split open hard-surfaced food. In humans, the canine teeth are the main components in occlusal function and articulation. The mandibular teeth function against the maxillary teeth in a particular movement that is harmonious to the shape of the occluding surfaces. This creates the incising and grinding functions. The teeth must mesh together the way gears mesh in a transmission. If the interdigitation of the opposing cusps and incisal edges are not directed properly the teeth will wear abnormally (attrition), break away irregular crystalline enamel structures from the surface (abrasion), or fracture larger pieces (abfraction). This is a three-dimensional movement of the mandible in relation to the maxilla. There are three points of guidance: the two posterior points provided by the temporomandibular joints and the anterior component provided by the incisors and canines. The incisors mostly control the vertical opening of the chewing cycle when the muscles of mastication move the jaw forwards and backwards (protrusion/retrusion). The canines come into function guiding the vertical movement when the chewing is side to side (lateral). The canines alone can cause the other teeth to separate at the extreme end of the cycle (cuspid guided function) or all the posterior teeth can continue to stay in contact (group function). The entire range of this movement is the envelope of masticatory function. The initial movement inside this envelope is directed by the shape of the teeth in contact and the Glenoid Fossa/Condyle shape. The outer extremities of this envelope are limited by muscles, ligaments and the articular disc of the TMJ. Without the guidance of anterior incisors and canines, this envelope of function can be destructive to the remaining teeth resulting in periodontal trauma from occlusion seen as wear, fracture or tooth loosening and loss. The premolars and molars are at the back of the mouth. Depending on the particular mammal and its diet, these two kinds of teeth prepare pieces of food to be swallowed by grinding, shearing, or crushing. The specialised teeth—incisors, canines, premolars, and molars—are found in the same order in every mammal. [6] In many mammals, the infants have a set of teeth that fall out and are replaced by adult teeth. These are called deciduous teeth, primary teeth, baby teeth or milk teeth. [7] [8] Animals that have two sets of teeth, one followed by the other, are said to be diphyodont. Normally the dental formula for milk teeth is the same as for adult teeth except that the molars are missing.
Because every mammal's teeth are specialised for different functions, many mammal groups have lost the teeth that are not needed in their adaptation. Tooth form has also undergone evolutionary modification as a result of natural selection for specialised feeding or other adaptations. Over time, different mammal groups have evolved distinct dental features, both in the number and type of teeth and in the shape and size of the chewing surface. [9]
The number of teeth of each type is written as a dental formula for one side of the mouth, or quadrant, with the upper and lower teeth shown on separate rows. The number of teeth in a mouth is twice that listed, as there are two sides. In each set, incisors (I) are indicated first, canines (C) second, premolars (P) third, and finally molars (M), giving I:C:P:M. [9] [10] So for example, the formula 2.1.2.3 for upper teeth indicates 2 incisors, 1 canine, 2 premolars, and 3 molars on one side of the upper mouth. The deciduous dental formula is notated in lowercase lettering preceded by the letter d: for example: di:dc:dp. [10]
An animal's dentition for either deciduous or permanent teeth can thus be expressed as a dental formula, written in the form of a fraction, which can be written as I.C.P.MI.C.P.M, or I.C.P.M / I.C.P.M. [10] [11] For example, the following formulae show the deciduous and usual permanent dentition of all catarrhine primates, including humans:
The greatest number of teeth in any known placental land mammal[ specify ] was 48, with a formula of 3.1.5.33.1.5.3. [9] However, no living placental mammal has this number. In extant placental mammals, the maximum dental formula is 3.1.4.33.1.4.3 for pigs. Mammalian tooth counts are usually identical in the upper and lower jaws, but not always. For example, the aye-aye has a formula of 1.0.1.31.0.0.3, demonstrating the need for both upper and lower quadrant counts. [10]
Teeth are numbered starting at 1 in each group. Thus the human teeth are I1, I2, C1, P3, P4, M1, M2, and M3. [12] (See next paragraph for premolar naming etymology.) In humans, the third molar is known as the wisdom tooth, whether or not it has erupted. [13]
Regarding premolars, there is disagreement regarding whether the third type of deciduous tooth is a premolar (the general consensus among mammalogists) or a molar (commonly held among human anatomists). [8] There is thus some discrepancy between nomenclature in zoology and in dentistry. This is because the terms of human dentistry, which have generally prevailed over time, have not included mammalian dental evolutionary theory. There were originally four premolars in each quadrant of early mammalian jaws. However, all living primates have lost at least the first premolar. "Hence most of the prosimians and platyrrhines have three premolars. Some genera have also lost more than one. A second premolar has been lost in all catarrhines. The remaining permanent premolars are then properly identified as P2, P3 and P4 or P3 and P4; however, traditional dentistry refers to them as P1 and P2". [7]
The order in which teeth emerge through the gums is known as the dental eruption sequence. Rapidly developing anthropoid primates such as macaques, chimpanzees, and australopithecines have an eruption sequence of M1 I1 I2 M2 P3 P4 C M3, whereas anatomically modern humans have the sequence M1 I1 I2 C P3 P4 M2 M3. The later that tooth emergence begins, the earlier the anterior teeth (I1–P4) appear in the sequence. [12]
Species | Dental formula | Comment |
---|---|---|
Non placental | Non-placental mammals such as marsupials (e.g., opossums) can have more teeth than placentals. | |
Bilby | 5.1.3.43.1.3.4 [15] | |
Kangaroo | 3.1.2.41.0.2.4 [16] | |
Musky rat-kangaroo | 3.1.1.42.0.1.4 [17] | |
Rest of Potoroidae | 3.1.1.41.0.1.4 [17] | The marsupial family Potoroidae includes the bettongs, potoroos, and two of the rat-kangaroos. All are rabbit-sized, brown, jumping marsupials and resemble a large rodent or a very small wallaby. |
Tasmanian devil | 4.1.2.43.1.2.4 [18] | |
Opossum | 5.1.3.44.1.3.4 [19] | |
Placental | Some examples of dental formulae for placental mammals. | |
Apes | 2.1.2.32.1.2.3 | All apes (excluding 20–23% of humans) and Old World monkeys share this formula, sometimes known as the cercopithecoid dental formula. [13] |
Armadillo | 0.0.7.10.0.7.1 [20] | |
Aye-aye | 1.0.1.31.0.0.3 [21] | A prosimian. The aye-aye's deciduous dental formula (dI:dC:dM) is 2.1.22.1.2. [10] |
Badger | 3.1.3.13.1.3.2 [22] | |
Big brown bat | 2.1.1.33.1.2.3 [19] | |
Red bat, hoary bat, Seminole bat, Mexican free-tailed bat | 1.1.2.33.1.2.3 [19] | |
Camel | 1.1.3.33.1.2.3 [23] | |
Cat (deciduous) | 3.1.3.03.1.2.0 [24] | |
Cat (permanent) | 3.1.3.13.1.2.1 [11] | The last upper premolar and first lower molar of the cat, since it is a carnivore, are called carnassials and are used to slice meat and skin. |
Cow | 0.0.3.33.1.3.3 [25] | The cow has no upper incisors or canines, the rostral portion of the upper jaw forming a dental pad. The lower canine is incisiform, giving the appearance of a 4th incisor. |
Dog (deciduous) | 3.1.3.03.1.3.0 [24] | |
Dog (permanent) | 3.1.4.23.1.4.3 [22] | |
Eared Seal | 3.1.4.1-32.1.4.1 [26] | |
Eulemur | 3.1.3.33.1.3.3 | Prosimian genus to which the large Malagasy or 'true' lemurs belong. [27] Ruffed lemurs (genus Varecia), [28] dwarf lemurs (genus Mirza), [29] and mouse lemurs (genus Microcebus) also have this dental formula, but the mouse lemurs have a dental comb. [30] |
Euoticus | 2.1.3.32.1.3.3 | Prosimian genus to which the needle-clawed bushbabies (or galagos) belong. Specialised morphology for gummivory includes procumbent dental comb and caniniform upper anterior premolars. [27] |
Fox (red) | 3.1.4.23.1.4.3 [22] | |
Guinea pig | 1.0.1.31.0.1.3 [31] | |
Hedgehog | 3.1.3.32.1.2.3 [22] | |
Horse (deciduous) | 3.0.3.03.0.3.0 [32] [33] | |
Horse (permanent) | 3.0-1.3-4.33.0-1.3.3 | Permanent dentition varies from 36 to 42, depending on the presence or absence of canines and the number of premolars. [34] The first premolar (wolf tooth) may be absent or rudimentary, [32] [33] and is mostly present only in the upper (maxillary) jaw. [33] The canines are small and spade-shaped, and usually present only in males. [34] Canines appear in 20–25% of females and are usually smaller than in males. [33] [35] |
Human (deciduous teeth) | See comment | Either 2.1.2.02.1.2.0 or 2.1.0.22.1.0.2. Human anatomists and mammal anatomists differ on whether the two anterior deciduous teeth are premolars (mammalogists) or molars (human anatomists) |
Human (permanent teeth) | 2.1.2.2-32.1.2.2-3 | Wisdom teeth are congenitally absent in 20–23% of the human population; the proportion of agenesis of wisdom teeth varies considerably among human populations, ranging from a near 0% incidence rate among Aboriginal Tasmanians to near 100% among Indigenous Mexicans. [36] |
Indri | See comment | A prosimian. Dental formula disputed. Either 2.1.2.32.0.2.3 or 2.1.2.31.1.2.3. Proponents of both formulae agree there are 30 teeth and that there are only four teeth in the dental comb. [37] |
Lepilemur | 0.1.3.32.1.3.3 | A prosimian. The upper incisors are lost in the adult, but are present in the deciduous dentition. [38] |
Lion | 3.1.3.13.1.2.1 [39] | |
Mole | 3.1.4.33.1.4.3 [22] | |
Mouse | 1.0.0.31.0.0.3 [22] | Plains pocket mouse (Perognathus flavescens) have dental formula of 1.0.1.31.0.1.3. [40] |
New World monkeys | See comment | All New World monkeys have a dentition formula of 2.1.3.32.1.3.3 or 2.1.3.22.1.3.2. [13] |
Pantodonta | 3.1.4.33.1.4.3 [41] | Extinct suborder of early eutherians. |
Pig (deciduous) | 3.1.4.03.1.4.0 [24] | |
Pig (permanent) | 3.1.4.33.1.4.3 [22] | |
Rabbit | 2.0.3.31.0.2.3 [11] | |
Raccoon | 3.1.4.23.1.4.2 | |
Rat | 1.0.0.31.0.0.3 [22] | |
Sheep (deciduous) | 0.0.3.04.0.3.0 [24] | |
Sheep (permanent) | 0.0.3.33.1.3.3 [19] | |
Shrew | 3.1.3.33.1.3.3 [22] | |
Sifakas | See comment | Prosimians. Dental formula disputed. Either 2.1.2.32.0.2.3 or 2.1.2.31.1.2.3. Possess dental comb comprising four teeth. [42] |
Slender loris Slow loris | 2.1.3.32.1.3.3 | Prosimians. Lower incisors and canines form a dental comb; upper anterior dentition is peg-like and short. [43] [44] |
Squirrel | 1.0.2.31.0.1.3 [22] | |
Tarsiers | 2.1.3.31.1.3.3 | Prosimians. [45] |
Tiger | 3.1.3.13.1.2.1 [46] | |
Vole (field) | 1.0.0.31.0.0.3 [22] | |
Weasel | 3.1.3.13.1.3.2 [22] |
Dentition, or the study of teeth, is an important area of study for archaeologists, especially those specializing in the study of older remains. [47] [48] [49] Dentition affords many advantages over studying the rest of the skeleton itself (osteometry). The structure and arrangement of teeth is constant and, although it is inherited, does not undergo extensive change during environmental change, dietary specializations, or alterations in use patterns. The rest of the skeleton is much more likely to exhibit change because of adaptation. Teeth also preserve better than bone, and so the sample of teeth available to archaeologists is much more extensive and therefore more representative.
Dentition is particularly useful in tracking ancient populations' movements, because there are differences in the shapes of incisors, the number of grooves on molars, presence/absence of wisdom teeth, and extra cusps on particular teeth. These differences can not only be associated with different populations across space, but also change over time so that the study of the characteristics of teeth could say which population one is dealing with, and at what point in that population's history they are.
A dinosaur's dentition included all the teeth in its jawbones, which consist of the dentary, maxillary, and in some cases the premaxillary bones. The maxilla is the main bone of the upper jaw. The premaxilla is a smaller bone forming the anterior of the animal's upper jaw. The dentary is the main bone that forms the lower jaw (mandible). The predentary is a smaller bone that forms the anterior end of the lower jaw in ornithischian dinosaurs; it is always edentulous and supported a horny beak.
Unlike modern lizards, dinosaur teeth grew individually in the sockets of the jawbones, which are known as the dental alveoli. This thecodont dentition is also present in crocodilians and mammals, but is not found among the non-archosaur reptiles, which instead have acrodont or pleurodont dentition. [50] Teeth that were lost were replaced by teeth below the roots in each tooth socket. Occlusion refers to the closing of the dinosaur's mouth, where the teeth from the upper and lower parts of the jaw meet. If the occlusion causes teeth from the maxillary or premaxillary bones to cover the teeth of the dentary and predentary, the dinosaur is said to have an overbite, the most common condition in this group. The opposite condition is considered to be an underbite, which is rare in theropod dinosaurs.
The majority of dinosaurs had teeth that were similarly shaped throughout their jaws but varied in size. Dinosaur tooth shapes included cylindrical, peg-like, teardrop-shaped, leaf-like, diamond-shaped and blade-like. A dinosaur that has a variety of tooth shapes is said to have heterodont dentition. An example of this are dinosaurs of the group Heterodontosauridae and the enigmatic early dinosaur, Eoraptor . While most dinosaurs had a single row of teeth on each side of their jaws, others had dental batteries where teeth in the cheek region were fused together to form compound teeth. Individually these teeth were not suitable for grinding food, but when joined together with other teeth they would form a large surface area for the mechanical digestion of tough plant materials. This type of dental strategy is observed in ornithopod and ceratopsian dinosaurs as well as the duck-billed hadrosaurs, which had more than one hundred teeth in each dental battery. The teeth of carnivorous dinosaurs, called ziphodont, were typically blade-like or cone-shaped, curved, with serrated edges. This dentition was adapted for grasping and cutting through flesh. In some cases, as observed in the railroad-spike-sized teeth of Tyrannosaurus rex , the teeth were designed to puncture and crush bone. Some dinosaurs had procumbent teeth, which projected forward in the mouth. [51]
Some articles have helpful discussions on dentition, which will be listed as identified.
Fauna of Australia Volume 1b - Mammalia
The dental formula for both Bactrian and dromedary camels is incisors (I) 1/3, canines (C) 1/1, premolars (P) 3/2, molars (M) 3/3.
The dental formula is 3/2, 1/1, 4/4, 1-3/1 = 34-38.
Horse teeth refers to the dentition of equine species, including horses and donkeys. Equines are both heterodontous and diphyodontous, which means that they have teeth in more than one shape, and have two successive sets of teeth, the deciduous and permanent sets.
In mammalian oral anatomy, the canine teeth, also called cuspids, dogteeth, eye teeth, vampire teeth, or fangs, are the relatively long, pointed teeth. In the context of the upper jaw, they are also known as fangs. They can appear more flattened, however, causing them to resemble incisors and leading them to be called incisiform. They developed and are used primarily for firmly holding food in order to tear it apart, and occasionally as weapons. They are often the largest teeth in a mammal's mouth. Individuals of most species that develop them normally have four, two in the upper jaw and two in the lower, separated within each jaw by incisors; humans and dogs are examples. In most species, canines are the anterior-most teeth in the maxillary bone. The four canines in humans are the two upper maxillary canines and the two lower mandibular canines. They are specially prominent in dogs (Canidae), hence the name.
Ferugliotherium is a genus of fossil mammals in the family Ferugliotheriidae from the Campanian and/or Maastrichtian period of Argentina. It contains a single species, Ferugliotherium windhauseni, which was first described in 1986. Although originally interpreted on the basis of a single brachydont (low-crowned) molar as a member of Multituberculata, an extinct group of small, rodent-like mammals, it was recognized as related to the hypsodont (high-crowned) Sudamericidae following the discovery of additional material in the early 1990s. After a jaw of the sudamericid Sudamerica was described in 1999, these animals were no longer considered to be multituberculates and a few fossils that were previously considered to be Ferugliotherium were assigned to unspecified multituberculates instead. Since 2005, a relationship between gondwanatheres and multituberculates has again received support. A closely related animal, Trapalcotherium, was described in 2009 on the basis of a single tooth.
Incisors are the front teeth present in most mammals. They are located in the premaxilla above and on the mandible below. Humans have a total of eight. Opossums have 18, whereas armadillos have none.
Dorudon ("spear-tooth") is a genus of extinct basilosaurid ancient whales that lived alongside Basilosaurus 40.4 to 33.9 million years ago in the Eocene. It was a small whale, with D. atrox measuring 5 metres (16 ft) long and weighing 1–2.2 metric tons. Dorudon lived in warm seas around the world and fed on small fish and mollusks. Fossils have been found along the former shorelines of the Tethys Sea in present-day Egypt and Pakistan, as well as in the United States, New Zealand and Western Sahara.
Plesiadapis is one of the oldest known primate-like mammal genera which existed about 58–55 million years ago in North America and Europe. Plesiadapis means "near-Adapis", which is a reference to the adapiform primate of the Eocene period, Adapis. Plesiadapis tricuspidens, the type specimen, is named after the three cusps present on its upper incisors.
Permanent teeth or adult teeth are the second set of teeth formed in diphyodont mammals. In humans and old world simians, there are thirty-two permanent teeth, consisting of six maxillary and six mandibular molars, four maxillary and four mandibular premolars, two maxillary and two mandibular canines, four maxillary and four mandibular incisors.
Dental professionals, in writing or speech, use several different dental notation systems for associating information with a specific tooth. The three most common systems are the FDI World Dental Federation notation, the Universal Numbering System, and the Palmer notation. The FDI notation is used worldwide, and the Universal is used widely in the United States. The FDI notation can be easily adapted to computerized charting.
FDI World Dental Federation notation is the world's most commonly used dental notation. It is designated by the International Organization for Standardization as standard ISO 3950 "Dentistry — Designation system for teeth and areas of the oral cavity".
Tooth eruption is a process in tooth development in which the teeth enter the mouth and become visible. It is currently believed that the periodontal ligament plays an important role in tooth eruption. The first human teeth to appear, the deciduous (primary) teeth, erupt into the mouth from around 6 months until 2 years of age, in a process known as "teething". These teeth are the only ones in the mouth until a person is about 6 years old creating the primary dentition stage. At that time, the first permanent tooth erupts and begins a time in which there is a combination of primary and permanent teeth, known as the mixed dentition stage, which lasts until the last primary tooth is lost. Then, the remaining permanent teeth erupt into the mouth during the permanent dentition stage.
Dental anatomy is a field of anatomy dedicated to the study of human tooth structures. The development, appearance, and classification of teeth fall within its purview. Tooth formation begins before birth, and the teeth's eventual morphology is dictated during this time. Dental anatomy is also a taxonomical science: it is concerned with the naming of teeth and the structures of which they are made, this information serving a practical purpose in dental treatment.
A toothcomb is a dental structure found in some mammals, comprising a group of front teeth arranged in a manner that facilitates grooming, similar to a hair comb. The toothcomb occurs in lemuriform primates, treeshrews, colugos, hyraxes, and some African antelopes. The structures evolved independently in different types of mammals through convergent evolution and vary both in dental composition and structure. In most mammals the comb is formed by a group of teeth with fine spaces between them. The toothcombs in most mammals include incisors only, while in lemuriform primates they include incisors and canine teeth that tilt forward at the front of the lower jaw, followed by a canine-shaped first premolar. The toothcombs of colugos and hyraxes take a different form with the individual incisors being serrated, providing multiple tines per tooth.
A tooth is a hard, calcified structure found in the jaws of many vertebrates and used to break down food. Some animals, particularly carnivores and omnivores, also use teeth to help with capturing or wounding prey, tearing food, for defensive purposes, to intimidate other animals often including their own, or to carry prey or their young. The roots of teeth are covered by gums. Teeth are not made of bone, but rather of multiple tissues of varying density and hardness that originate from the outermost embryonic germ layer, the ectoderm.
Hipposideros besaoka is an extinct bat from Madagascar in the genus Hipposideros. It is known from numerous jaws and teeth, which were collected in a cave at Anjohibe in 1996 and described as a new species in 2007. The site where H. besaoka was found is at most 10,000 years old; other parts of the cave have yielded H. commersoni, a living species of Hipposideros from Madagascar, and some material that is distinct from both species. H. besaoka was larger than H. commersoni, making it the largest insectivorous bat of Madagascar, and had broader molars and a more robust lower jaw. As usual in Hipposideros, the second upper premolar is small and displaced from the toothrow, and the second lower premolar is large.
Teeth are common to most vertebrates, but mammalian teeth are distinctive in having a variety of shapes and functions. This feature first arose among early therapsids during the Permian, and has continued to the present day. All therapsid groups with the exception of the mammals are now extinct, but each of these groups possessed different tooth patterns, which aids with the classification of fossils.
Dermotherium is a genus of fossil mammals closely related to the living colugos, a small group of gliding mammals from Southeast Asia. Two species are recognized: D. major from the Late Eocene of Thailand, based on a single fragment of the lower jaw, and D. chimaera from the Late Oligocene of Thailand, known from three fragments of the lower jaw and two isolated upper molars. In addition, a single isolated upper molar from the Early Oligocene of Pakistan has been tentatively assigned to D. chimaera. All sites where fossils of Dermotherium have been found were probably forested environments and the fossil species were probably forest dwellers like living colugos, but whether they had the gliding adaptations of the living species is unknown.
Serial extraction is the planned extraction of certain deciduous teeth and specific permanent teeth in an orderly sequence and predetermined pattern to guide the erupting permanent teeth into a more favorable position.
Plesiopithecus is an extinct genus of early strepsirrhine primate from the late Eocene.
Changes to the dental morphology and jaw are major elements of hominid evolution. These changes were driven by the types and processing of food eaten. The evolution of the jaw is thought to have facilitated encephalization, speech, and the formation of the uniquely human chin.
Saxonella is a genus of extinct primate from the Paleocene Epoch, 66–56 Ma. The genus is present in the fossil record from around ~62–57 Ma. Saxonella has been found in fissure fillings in Walbeck, Germany as well as in the Paskapoo Formation in Alberta, Canada. Saxonella is one of five families within the superfamily Plesiadapoidae, which appears in the fossil record from the mid Paleocene to the early Eocene. Analyses of molars by paleontologists suggest that Saxonella most likely had a folivorous diet.