Kukersella | |
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Scientific classification | |
Kingdom: | Animalia |
Phylum: | Bryozoa |
Class: | Stenolaemata |
Order: | Cyclostomatida |
Family: | † Crownoporidae |
Genus: | † Kukersella Toots, 1952 |
Species [1] | |
See text | |
Synonyms [1] | |
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Kukersella is an extinct genus of bryozoan of the family Crownoporidae, known from the Ordovician period. Its colonies consist of cylindrical branches growing from an encrusting base. [2]
The Ordovician is a geologic period and system, the second of six periods of the Paleozoic Era. The Ordovician spans 41.6 million years from the end of the Cambrian Period 485.4 million years ago (Mya) to the start of the Silurian Period 443.8 Mya.
The PaleozoicEra is the earliest of three geologic eras of the Phanerozoic Eon. The name Paleozoic was coined by the British geologist Adam Sedgwick in 1838 by combining the Greek words palaiós and zōḗ, "life", meaning "ancient life").
The Silurian is a geologic period and system spanning 24.6 million years from the end of the Ordovician Period, at 443.8 million years ago (Mya), to the beginning of the Devonian Period, 419.2 Mya. The Silurian is the shortest period of the Paleozoic Era. As with other geologic periods, the rock beds that define the period's start and end are well identified, but the exact dates are uncertain by a few million years. The base of the Silurian is set at a series of major Ordovician–Silurian extinction events when up to 60% of marine genera were wiped out.
Trilobites are extinct marine arthropods that form the class Trilobita. Trilobites form one of the earliest-known groups of arthropods. The first appearance of trilobites in the fossil record defines the base of the Atdabanian stage of the Early Cambrian period and they flourished throughout the lower Paleozoic before slipping into a long decline, when, during the Devonian, all trilobite orders except the Proetida died out. The last extant trilobites finally disappeared in the mass extinction at the end of the Permian about 252 million years ago. Trilobites were among the most successful of all early animals, existing in oceans for almost 270 million years, with over 22,000 species having been described.
Acritarchs are organic microfossils, known from approximately 1800 million years ago to the present. The classification is a catch all term used to refer to any organic microfossils that cannot be assigned to other groups. Their diversity reflects major ecological events such as the appearance of predation and the Cambrian explosion.
Stenolaemata are a class of exclusively marine bryozoans. Stenolaemates originated and diversified in the Ordovician, and more than 600 species are still alive today. All extant (living) species are in the order Cyclostomatida, the third-largest order of living bryozoans.
The Late Ordovician mass extinction (LOME), sometimes known as the end-Ordovician mass extinction or the Ordovician-Silurian extinction, is the first of the "big five" major mass extinction events in Earth's history, occurring roughly 443 Mya. It is often considered to be the second-largest known extinction event, in terms of the percentage of genera that became extinct. Extinction was global during this interval, eliminating 49–60% of marine genera and nearly 85% of marine species. Under most tabulations, only the Permian-Triassic mass extinction exceeds the Late Ordovician mass extinction in biodiversity loss. The extinction event abruptly affected all major taxonomic groups and caused the disappearance of one third of all brachiopod and bryozoan families, as well as numerous groups of conodonts, trilobites, echinoderms, corals, bivalves, and graptolites. Despite its taxonomic severity, the Late Ordovician mass extinction did not produce major changes to ecosystem structures compared to other mass extinctions, nor did it lead to any particular morphological innovations. Diversity gradually recovered to pre-extinction levels over the first 5 million years of the Silurian Period.
Nautiloids are a group of marine cephalopods (Mollusca) which originated in the Late Cambrian and are represented today by the living Nautilus and Allonautilus. Fossil nautiloids are diverse and speciose, with over 2,500 recorded species. They flourished during the early Paleozoic era, when they constituted the main predatory animals. Early in their evolution, nautiloids developed an extraordinary diversity of shell shapes, including coiled morphologies and giant straight-shelled forms (orthocones). Only a handful of rare coiled species, the nautiluses, survive to the present day.
The Hirnantian is the final internationally recognized stage of the Ordovician Period of the Paleozoic Era. It was of short duration, lasting about 1.4 million years, from 445.2 to 443.8 Ma. The early part of the Hirnantian was characterized by cold temperatures, major glaciation, and a severe drop in sea level. In the latter part of the Hirnantian, temperatures rose, the glaciers melted, and sea level returned to the same or to a slightly higher level than it had been prior to the glaciation.
The Tremadocian is the lowest stage of Ordovician. Together with the later Floian Stage it forms the Lower Ordovician Epoch. The Tremadocian lasted from 485.4 to 477.7 million years ago. The base of the Tremadocian is defined as the first appearance of the conodont species Iapetognathus fluctivagus at the Global Boundary Stratotype Section and Point (GSSP) section on Newfoundland.
The Cambrian–Ordovician extinction event occurred approximately 488 million years ago (mya). This early Phanerozoic Eon extinction event eliminated many brachiopods and conodonts, and severely reduced the number of trilobite species. The Period in the Cambrian extinction in which most of the extinction occurred was the Caerfai Period.
The Andean-Saharan glaciation, also known as the Early Palaeozoic Icehouse, the Early Palaeozoic Ice Age, the Late Ordovician glaciation, the end-Ordovician glaciation, or the Hirnantian glaciation, occurred during the Paleozoic from approximately 460 Ma to around 420 Ma, during the Late Ordovician and the Silurian period. The major glaciation during this period, which was formerly thought only to consist of the Hirnantian glaciation itself, but has now been recognized as a longer, more gradual event that began as early as the Darriwilian, is widely considered to be the leading cause of the Ordovician-Silurian extinction event. Evidence of this glaciation can be seen in places such as Arabia, North Africa, South Africa, Brazil, Peru, Bolivia, Chile, Argentina, and Wyoming. More evidence derived from isotopic data is that during the Late Ordovician, tropical ocean temperatures were about 5 °C cooler than present day; this would have been a major factor that aided in the glaciation process.
Tabulata, commonly known as tabulate corals, are an order of extinct forms of coral. They are almost always colonial, forming colonies of individual hexagonal cells known as corallites defined by a skeleton of calcite, similar in appearance to a honeycomb. Adjacent cells are joined by small pores. Their distinguishing feature is their well-developed horizontal internal partitions (tabulae) within each cell, but reduced or absent vertical internal partitions. They are usually smaller than rugose corals, but vary considerably in shape, from flat to conical to spherical.
The Darriwilian is the upper stage of the Middle Ordovician. It is preceded by the Dapingian and succeeded by the Upper Ordovician Sandbian Stage. The lower boundary of the Darriwilian is defined as the first appearance of the graptolite species Undulograptus austrodentatus around 467.3 million years ago. It lasted for about 8.9 million years until the beginning of the Sandbian around 458.4 million years ago. This stage of the Ordovician was marked by the beginning of the Andean-Saharan glaciation.
Kazakhstania, the Kazakh terranes, or the Kazakhstan Block, is a geological region in Central Asia which consists of the area roughly centered on Lake Balkhash, north and east of the Aral Sea, south of the Siberian craton and west of the Altai Mountains. The Junggar basin in Xinjiang, China, is also part of Kazakhstania, though sometimes referred to as the Junggar Block. Because the Kazakh terranes merged during the Late Ordovician as part of the Central Asian Orogenic Belt they are also referred to as the Kazakh Orogen. These terranes are located in what is today Kazakhstan, north-eastern Uzbekistan, northern Kyrgyzstan and south-western China. Today Kazakhstania is surrounded by three large, former continents: to the north-east the Gornostaev Shear Zone separates it from Siberia with which it collided during the Carboniferous; to the north-west is Baltica which lay adjacent to the Kazakh Tourgai terrane but far away from Kazakhstania; to the south and east was Gondwana stretching from the South Pole to the Equator. Not far away from the dispersed Kazakh terranes were South China, North China, and Tarim, but how these continental blocks were positioned relative to Gondwana is not known.
The Sandbian is the first stage of the Upper Ordovician. It follows the Darriwilian and is succeeded by the Katian. Its lower boundary is defined as the first appearance datum of the graptolite species Nemagraptus gracilis around 458.4 million years ago. The Sandbian lasted for about 5.4 million years until the beginning of the Katian around 453 million years ago.
Archeognathus is a fossilized jaw apparatus of a large predatory conodont from the Ordovician period. Its large size has made classification difficult, and it has historically been compared to conodonts and gnathostomes since its remains were first discovered in Missouri. Complete articulated jaw apparatus of Archeognathus primus are common in the Winneshiek Shale lagerstätte of Iowa, allowing its identity as a conodont to be secured.
The Soom Shale is a member of the Late Ordovician (Hirnantian) Cederberg Formation in South Africa, renowned for its remarkable preservation of soft-tissue in fossil material. Deposited in still waters, the unit lacks bioturbation, perhaps indicating anoxic conditions.
Paleontology in Tennessee refers to paleontological research occurring within or conducted by people from the U.S. state of Tennessee. During the early part of the Paleozoic era, Tennessee was covered by a warm, shallow sea. This sea was home to brachiopods, bryozoans, cephalopods, corals, and trilobites. Tennessee is one of the best sources of Early Devonian fossils in North America. During the mid-to-late Carboniferous, the state became a swampy environment, home to a rich variety of plants including ferns and scale trees. A gap in the local rock record spans from the Permian through the Jurassic. During the Cretaceous, the western part of the state was submerged by seawater. The local waters were home to more fossil gastropods than are known from anywhere else in the world. Mosasaurs and sea turtles also inhabited these waters. On land the state was home to dinosaurs. Western Tennessee was still under the sea during the early part of the Cenozoic. Terrestrial portions of the state were swampy. Climate cooled until the Ice Age, when the state was home to Camelops, horses, mammoths, mastodons, and giant ground sloths. The local Yuchi people told myths of giant lizard monsters that may have been inspired by fossils either local or encountered elsewhere. In 1920, after local fossils became a subject of formal scientific study, a significant discovery of a variety of Pleistocene creatures was made near Nashville. The Cretaceous bivalve Pterotrigonia thoracica is the Tennessee state fossil.
The Ordovician meteor event was a dramatic increase in the rate at which L chondrite meteorites fell to Earth during the Middle Ordovician period, about 467.5±0.28 million years ago. This is indicated by abundant fossil L chondrite meteorites in a quarry in Sweden and enhanced concentrations of ordinary chondritic chromite grains in sedimentary rocks from this time. This temporary increase in the impact rate was most likely caused by the destruction of the L chondrite parent body 468 ± 0.3 million years ago having scattered fragments into Earth-crossing orbits, a chronology which is also supported by shock ages in numerous L chondrite meteorites that fall to Earth today. It has been speculated that this influx contributed to, or possibly even instigated, the Great Ordovician Biodiversification Event, although this has been questioned.