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Photosynthesis is a process used by plants and other organisms to convert light energy into chemical energy that can later be released to fuel the organisms' activities. This chemical energy is stored in carbohydrate molecules, such as sugars, which are synthesized from carbon dioxide and water – hence the name photosynthesis, from the Greek phōs (φῶς), "light", and sunthesis (σύνθεσις), "putting together". In most cases, oxygen is also released as a waste product. Most plants, most algae, and cyanobacteria perform photosynthesis; such organisms are called photoautotrophs. Photosynthesis is largely responsible for producing and maintaining the oxygen content of the Earth's atmosphere, and supplies most of the energy necessary for life on Earth.
In botany, a stoma, also called a stomate is a pore, found in the epidermis of leaves, stems, and other organs, that controls the rate of gas exchange. The pore is bordered by a pair of specialized parenchyma cells known as guard cells that are responsible for regulating the size of the stomatal opening.
Inside a plant, the apoplast is the space outside the plasma membrane within which material can diffuse freely. It is interrupted by the Casparian strip in roots, by air spaces between plant cells and by the plant cuticle.
Thigmotropism is a directional growth movement which occurs as a mechanosensory response to a touch stimulus. Thigmotropism is typically found in twining plants and tendrils, however plant biologists have also found thigmotropic responses in flowering plants and fungi. This behavior occurs due to unilateral growth inhibition. That is, the growth rate on the side of the stem which is being touched is slower than on the side opposite the touch. The resultant growth pattern is to attach and sometimes curl around the object which is touching the plant. However, flowering plants have also been observed to move or grow their sex organs toward a pollinator that lands on the flower, as in Portulaca grandiflora.
Water potential is the potential energy of water per unit volume relative to pure water in reference conditions. Water potential quantifies the tendency of water to move from one area to another due to osmosis, gravity, mechanical pressure and matrix effects such as capillary action. The concept of water potential has proved useful in understanding and computing water movement within plants, animals, and soil. Water potential is typically expressed in potential energy per unit volume and very often is represented by the Greek letter ψ.
Mimosa pudica is a creeping annual or perennial flowering plant of the pea/legume family Fabaceae and Magnoliopsida taxon, often grown for its curiosity value: the compound leaves fold inward and droop when touched or shaken, defending themselves from harm, and re-open a few minutes later. In the UK it has gained the Royal Horticultural Society's Award of Garden Merit.
Thermotropism or thermotropic movement is the movement of a plant or part of a plant in response to a change in temperature. A common example is the curling of Rhododendron leaves in response to cold temperatures. Mimosa pudica also show thermotropism by the collapsing of leaf petioles leading to the folding of leaflets, when temperature drops.
Tonicity is a measure of the effective osmotic pressure gradient; the water potential of two solutions separated by a semipermeable cell membrane. In other words, tonicity is the relative concentration of solutes dissolved in solution which determine the direction and extent of diffusion. It is commonly used when describing the response of cells immersed in an external solution.
Phenotypic plasticity refers to some of the changes in an organism's behavior, morphology and physiology in response to a unique environment. Fundamental to the way in which organisms cope with environmental variation, phenotypic plasticity encompasses all types of environmentally induced changes that may or may not be permanent throughout an individual's lifespan. The term was originally used to describe developmental effects on morphological characters, but is now more broadly used to describe all phenotypic responses to environmental change, such as acclimation (acclimatization), as well as learning. The special case when differences in environment induce discrete phenotypes is termed polyphenism.
Turgor pressure is the force within the cell that pushes the plasma membrane against the cell wall.
Thigmonasty or seismonasty is the nastic response of a plant or fungus to touch or vibration. Conspicuous examples of thigmonasty include many species in the leguminous subfamily Mimosoideae, active carnivorous plants such as Dionaea and a wide range of pollination mechanisms.
Guard cells are specialized plant cells in the epidermis of leaves, stems and other organs that are used to control gas exchange. They are produced in pairs with a gap between them that forms a stomatal pore. The stomatal pores are largest when water is freely available and the guard cells turgid, and closed when water availability is critically low and the guard cells become flaccid. Photosynthesis depends on the diffusion of carbon dioxide (CO2) from the air through the stomata into the mesophyll tissues. Oxygen (O2), produced as a byproduct of photosynthesis, exits the plant via the stomata. When the stomata are open, water is lost by evaporation and must be replaced via the transpiration stream, with water taken up by the roots. Plants must balance the amount of CO2 absorbed from the air with the water loss through the stomatal pores, and this is achieved by both active and passive control of guard cell turgor pressure and stomatal pore size.
Ecophysiology, environmental physiology or physiological ecology is a biological discipline that studies the response of an organism's physiology to environmental conditions. It is closely related to comparative physiology and evolutionary physiology. Ernst Haeckel's coinage bionomy is sometimes employed as a synonym.
A pulvinus is a joint-like thickening at the base of a plant leaf or leaflet that facilitates growth-independent movement. Pulvini are common, for example, in members of the bean family Fabaceae (Leguminosae) and the prayer plant family Marantaceae.
Phototropism is the growth of an organism in response to a light stimulus. Phototropism is most often observed in plants, but can also occur in other organisms such as fungi. The cells on the plant that are farthest from the light have a chemical called auxin that reacts when phototropism occurs. This causes the plant to have elongated cells on the furthest side from the light. Phototropism is one of the many plant tropisms or movements which respond to external stimuli. Growth towards a light source is called positive phototropism, while growth away from light is called negative phototropism (skototropism). Most plant shoots exhibit positive phototropism, and rearrange their chloroplasts in the leaves to maximize photosynthetic energy and promote growth. Some vine shoot tips exhibit negative phototropism, which allows them to grow towards dark, solid objects and climb them. The combination of phototropism and gravitropism allow plants to grow in the correct direction.
By definition, stomatal conductance, usually measured in mmol m⁻² s⁻¹, is the measure of the rate of passage of carbon dioxide (CO2) entering, or water vapor exiting through the stomata of a leaf. Stomata are small pores on the top and or bottom of a leaf that are responsible for taking in CO2 and expelling water vapour.
Breeding for drought resistance is the process of breeding plants with the goal of reducing the impact of dehydration on plant growth.
Ruth Lyttle Satter was an American botanist best known for her work on circadian leaf movement.
Paraheliotropism refers to the phenomenon in which plants orient their leaves parallel to incoming rays of light, usually as a means of minimizing excess light absorption. Excess light absorption can cause a variety of physiological problems for plants, including overheating, dehydration, loss of turgor, photoinhibition, photo-oxidation, and photorespiration, so paraheliotropism can be viewed as an advantageous behavior in high light environments. Not all plants exhibit this behavior, but it has developed in multiple lineages.
A mechanoreceptor is a sensory organ or cell that responds to mechanical stimulation such as touch, pressure, vibration, and sound from both the internal and external environment. Mechanoreceptors are well-documented in animals and are integrated into the nervous system as sensory neurons. While plants do not have nerves or a nervous system like animals, they also contain mechanoreceptors that perform a similar function. Mechanoreceptors detect mechanical stimulus originating from within the plant (intrinsic) and from the surrounding environment (extrinsic). The ability to sense vibrations, touch, or other disturbance is an adaptive response to herbivory and attack so that the plant can appropriately defend itself against harm. Mechanoreceptors can be organized into three levels: molecular, cellular, and organ-level.
References
- ↑ Pantin F.; Simmonneau T.; Rolland G.; Dauzat M.; Muller B. (2011). "Control of Leaf Expansion: A Developmental Switch from Metabolics to Hydraulics". Plant Physiology. 156 (2): 803–815. doi:10.1104/pp.111.176289. PMC 3177277 . PMID 21474437.
- ↑ Hsiao T.C. (1973). "Plant Responses to Water Stress". Annual Review of Plant Physiology. 156: 519–570. doi:10.1146/annurev.pp.24.060173.002511.
- 1 2 3 Volkenburgh, E.V. (1999). "Leaf Expansion - an integrating plant behaviour". Plant, Cell & Environment. 22 (12): 1463–1473. doi: 10.1046/j.1365-3040.1999.00514.x .
- ↑ Poorter H., van der Werf A., eds. Lambers H., Poorter H., Van Vuuren M.M.I. (1998). "Is inherent variation in RGR determined by LAR at low irradiance and by NAR at high irradiance? A review of herbaceous species". Physiological Mechanisms and Ecological Consequences: 309–336.CS1 maint: multiple names: authors list (link)
- ↑ Lambers H.; Poorter H.; Van Vuuren M.M.I. (1998). "Inherent Variation in Plant Growth". Netherlands.