List of Copris species

These species belong to Copris , a genus of dung beetles in the family Scarabaeidae. ^{[1] [2]}

Copris species

• Copris acutidens Motschulsky, 1860
• Copris afgoi Cambefort, 1992
• Copris agnus Sharp, 1875
• Copris algol Nguyen-phung, 1989
• Copris amabilis Kolbe, 1914
• Copris amyntor Klug, 1855
• Copris anceus (Olivier, 1789)
• Copris andrewesi Waterhouse, 1891
• Copris angusticornis Arrow, 1933
• Copris angustus Nguyen-phung & Cambefort, 1986
• Copris anomiopseoides Boucomont, 1924
• Copris antares Ferreira, 1958
• Copris arcturus Gillet, 1907
• Copris ares Zelenka, 1993
• Copris arimotoi Ochi & Araya, 1992
• Copris arizonensis Schaeffer, 1906
• Copris armatus Harold, 1869
• Copris armeniacus Faldermann, 1835
• Copris armiger Gillet, 1910
• Copris arrowi Felsche, 1910
• Copris aspericollis Gillet, 1910
• Copris atropolitus Gillet, 1910
• Copris bacchus Thunberg, 1787
• Copris barclayi Ochi, Kon & Bai, 2009
• Copris basilewskyi Ferreira, 1962
• Copris basipunctatus Balthasar, 1942
• Copris bellator Chevrolat, 1844
• Copris bengalensis Gillet, 1911
• Copris bidens Kolbe, 1893
• Copris binodis
• Copris bootes Klug, 1855
• Copris boucardi Harold, 1869
• Copris bovinus Gillet, 1908
• Copris brachypterus Nomura, 1964
• Copris brittoni Ferreira, 1962
• Copris caelatus (Fabricius, 1794)
• Copris caliginosus Kohlmann, Cano & Delgado, 2003
• Copris cambeforti Nguyen-phung, 1988
• Copris cambodiensis Ochi, Kon & Kawahara, 2008
• Copris camerunus Felsche, 1904
• Copris caobangensis Bui, Dumack & Bonkowski, 2018 ^[3]
• Copris capensis Waterhouse, 1891
• Copris cariniceps Felsche, 1910
• Copris carinicus Gillet, 1910
• Copris carmelita Fabricius, 1801
• Copris cassius Péringuey, 1901
• Copris celebensis Lansberge, 1886
• Copris cheni Ochi, Kon & Bai, 2007
• Copris chimalapensis Mora-Aguilar & Delgado, 2015
• Copris ciliatus
• Copris colmanti Gillet, 1908
• Copris complexus Nguyen-phung, 1989
• Copris compressipennis Gillet, 1910
• Copris confucius Harold, 1877
• Copris coriarius Gillet, 1907
• Copris cornifrons Boheman, 1860
• Copris corniger Sahlberg, 1823
• Copris corpulentus Gillet, 1910
• Copris costaricensis Gahan, 1894
• Copris costariensis Matthews
• Copris crassus Deschodt & Davis, 2015
• Copris cribratus Gillet, 1927
• Copris cribricollis Gillet, 1910
• Copris curlettii Moretto, 2011
• Copris davidis (Deyrolle, 1878)
• Copris davisi Nguyen-phung & Cambefort, 1986
• Copris davisoni Waterhouse, 1891
• Copris decellei Cambefort, 1992
• Copris delicatus Arrow, 1931
• Copris denticulatus Nguyen-phung, 1988
• Copris diversus Waterhouse, 1891
• Copris draco Arrow, 1906
• Copris dracunculus Fereira, 1959
• Copris druidum Heer, 1862
• Copris dudleyi Cambefort, 1992
• Copris eburneus Cambefort, 1992
• Copris elpheneroides Felsche, 1910
• Copris elphenor Klug, 1855
• Copris erratus Lansberge, 1886
• Copris evanidus Klug, 1855
• Copris excisus Waterhouse, 1891
• Copris fallaciosus Gillet, 1907
• Copris fallax Felsche, 1910
• Copris felschei Reitter, 1892
• Copris fidius (Olivier, 1789)
• Copris frankenbergerianus Balthasar, 1958
• Copris freyi Ferreira, 1961
• Copris fricator (Fabricius, 1787)
• Copris fukiensis Balthasar, 1952
• Copris furciceps Felsche, 1910
• Copris garambae Cambefort, 1992
• Copris gazellarum Gillet, 1918
• Copris gibbulus Lansberge, 1886
• Copris gilleti Kolbe, 1907
• Copris gladiator Arrow, 1933
• Copris gopheri Hubbard, 1894
• Copris gracilis Waterhouse, 1891
• Copris graueri Kolbe, 1914
• Copris halffteri Matthews, 1959
• Copris harrisi Waterhouse, 1891
• Copris hartli Frey, 1975
• Copris hatayamai Ochi & Araya, 1992
• Copris hispanus (Linnaeus, 1764)
• Copris howdeni Matthews & Halffter, 1959
• Copris humilis Gillet, 1908
• Copris hybridus Gillet, 1910
• Copris igualensis Warner, 1990
• Copris illotus Balthasar, 1942
• Copris imitans Felsche, 1910
• Copris inaequabilis Zhang, 1997
• Copris incertus Say, 1835
• Copris inemarginatus Blatchley, 1918
• Copris inhalatus Quedenfeldt, 1884
• Copris insidiosus Péringuey, 1901
• Copris integer Reiche, 1847
• Copris interioris Kolbe, 1897
• Copris iris Sharp, 1875
• Copris jacchoides Nguyen-phung & Cambefort, 1987
• Copris jacchus (Fabricius, 1775)
• Copris jahi Nguyen-phung & Cambefort, 1986
• Copris johkii Ochi & Kon, 2014
• Copris juanmorai Mora-Aguilar & Delgado, 2015
• Copris jucundus Gillet, 1932
• Copris kachinensis Ochi, Kon & Kawahara, 2008
• Copris kartlinus Kabakov, 1988
• Copris kasagii Ochi & Kon, 1996
• Copris kasanka Josso & Prévost, 2016
• Copris katangae Cambefort & Nguyen-phung, 1996
• Copris keralensis Gill, 1986
• Copris kiuchii Masumoto, 1989
• Copris klapperichi Balthasar, 1942
• Copris klugi Harold, 1869
• Copris kusakabei Ochi, Kon & Kawahara, 2005
• Copris laeviceps Harold, 1862
• Copris laevigatus Gillet, 1927
• Copris laiiformis Nguyen-phung & Cambefort, 1987
• Copris laioides Boucomont, 1932
• Copris laius Harold, 1868
• Copris lannathai Hanboonsong & Masumoto & Ochi, 2003
• Copris leakeyorum Paulian, 1976
• Copris lecontei Matthews, 1962
• Copris leruei Josso & Prévost, 2017
• Copris lugubris Boheman, 1858
• Copris lunaris  [ fr ](Linnaeus, 1758)
• Copris macacus Lansberge, 1886
• Copris macclevei Warner, 1990
• Copris macer Péringuey, 1901
• Copris maesi Ratcliffe, 1998
• Copris magicus Harold, 1881
• Copris manni Ochi, Kon & Bai, 2009
• Copris marcus Gillet, 1921
• Copris martinae Nguyen-phung, 1988
• Copris martinezi Matthews and Halffter, 1968
• Copris matthewsi Delgado & Kohlmann, 2001
• Copris medogensis Zhang, 1988
• Copris megaceratoides Waterhouse, 1891
• Copris megasoma Matthews & Halffter, 1959
• Copris mesacanthus Harold, 1878
• Copris mexicanus Matthews & Halffter, 1959
• Copris minutus (Drury, 1773)
• Copris misellus Péringuey, 1901
• Copris moechus LeConte, 1854
• Copris moffartsi Gillet, 1907
• Copris mongkhoni Hanboonsong & Masumoto & Ochi, 2003
• Copris montivagus Gillet, 1908
• Copris morphaeus Gillet, 1932
• Copris mourgliai Zelenka, 1993
• Copris neglectus Moxey, 1963
• Copris neotridens Génier & Prévost, 2017
• Copris nepos Gillet, 1908
• Copris nevinsoni Waterhouse, 1891
• Copris novaki Zelenka, 1992
• Copris nubilosus Kohlmann & Cano & Delgado, 2003
• Copris numa Lansberge, 1886
• Copris obenbergeri Balthasar, 1933
• Copris obesus Boheman, 1857
• Copris ochus (Motschulsky, 1860)
• Copris orion Klug, 1835
• Copris orphanus Guérin-Méneville, 1847
• Copris parapecuarius Ochi, Kon & Kawahara, 2008
• Copris pauliani Nguyen-phung & Cambefort, 1986
• Copris pecuarius Lewis, 1884
• Copris pedarioides Lansberge, 1886
• Copris phungae Cambefort, 1992
• Copris phylax Gillet, 1908
• Copris poggii Ochi & Kon, 2005
• Copris potanini Semenov, 1891
• Copris prevosti Josso, 2011
• Copris pristinus Pierce, 1946
• Copris pseudobootes Ferreira, 1962
• Copris pseudochus Ochi & Kon, 2004
• Copris pseudomoffartsi Moretto, 2011
• Copris pseudosinicus Balthasar, 1958
• Copris pueli Mollandin De Boissy, 1905
• Copris punctatus Gillet, 1910
• Copris puncticollis Boheman, 1857
• Copris punctipennis Boucomont, 1914
• Copris punctulatus Wiedemann, 1823
• Copris punjabensis Gillet, 1921
• Copris quasilaevigatus Ochi, Kon & Bai, 2007
• Copris ramosiceps Gillet, 1921
• Copris rebouchei Harold, 1869
• Copris remotus Leconte, 1866
• Copris renwarti Nguyen-phung & Cambefort, 1986
• Copris repertus Walker, 1858
• Copris ribbei Lansberge, 1886
• Copris riekoae Ochi & Kon, 2003
• Copris ritsemae Harold, 1875
• Copris rugosus Gillet, 1908
• Copris ruricola Balthasar, 1933
• Copris sabinus Gillet, 1910
• Copris sacontala Redtenbacher, 1848
• Copris sallei Harold, 1869
• Copris sarpedon Harold, 1868
• Copris saundersi Harold, 1869
• Copris schoolmeestersi Ochi & Kon, 2010
• Copris scorpio Zelenka, 1993
• Copris serius Nguyen-phung, 1987
• Copris sexdentatus Thunberg, 1818
• Copris siamensis Gillet, 1910
• Copris siangensis Biswas, 1980
• Copris sierrensis Matthews, 1961
• Copris signatus Walker, 1858
• Copris simonettai Cambefort, 1992
• Copris simulator Nguyen-phung & Cambefort, 1986
• Copris singularis Nguyen-phung & Cambefort, 1986
• Copris sinicus Hope, 1842
• Copris sinon (Olivier, 1789)
• Copris sodalis Walker, 1858
• Copris sonensis Bui, Dumack & Bonkowski, 2018 ^[3]
• Copris sorex Balthasar, 1942
• Copris sphaeropterus Harold, 1877
• Copris spinator Harold, 1881
• Copris subdolus Balthasar, 1958
• Copris subpunctatus Gillet, 1910
• Copris subsidens Péringuey, 1901
• Copris subterraneus Heer, 1862
• Copris sumatrensis Gillet, 1921
• Copris surdus Arrow, 1931
• Copris szechouanicus Balthasar, 1958
• Copris tetraodon Gillet, 1910
• Copris tridentatus Solis & Kohlmann, 2003
• Copris tripartitus Waterhouse, 1875
• Copris truncatus Felsche, 1901
• Copris tsukamotoi Ochi, Kon & Kawahara, 2008
• Copris typhoeus Gerstaecker, 1884
• Copris uenoi Hanboonsong & Masumoto & Ochi, 2003
• Copris umbilicatus Abeille De Perrin, 1901
• Copris urus Boheman, 1857
• Copris usambaricus Gillet, 1908
• Copris vankhaii Nguyen-phung, 1988
• Copris victorini Boheman, 1857
• Copris vietnamicus Kabakov, 1994
• Copris vilhenai Ferreira, 1962
• Copris vrydaghi Ferreira, 1962
• Copris warneri Mccleve & Kohlmann, 2005
• Copris wiesei Kolbe, 1914
• Copris yangi Ochi, Kon & Bai, 2007
• Copris youngai Balthasar, 1967
• Copris zhangi Ochi, Kon & Bai, 2009

References

1. "Copris". GBIF. Retrieved 2019-01-05.
2. "Copris Genus Information". BugGuide.net. Retrieved 2019-01-05.
3. Bui, Van Bac; Dumack, Kenneth; Bonkowski, Michael (2018). "Two new species and one new record for the genus Copris (Coleoptera: Scarabaeidae: Scarabaeinae) from Vietnam with a key to Vietnamese species". European Journal of Entomology. 15. doi: 10.14411/eje.2018.016 .