Scarab beetle | |
---|---|
![]() | |
Central European scarab beetles with some anatomical details. Edmund Reitter's Fauna Germanica, 1908 | |
Scientific classification ![]() | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Class: | Insecta |
Order: | Coleoptera |
Suborder: | Polyphaga |
Infraorder: | Scarabaeiformia |
Superfamily: | Scarabaeoidea |
Family: | Scarabaeidae Latreille, 1802 |
Subfamilies [1] [2] [3] | |
|
The family Scarabaeidae, as currently defined, consists of over 35,000 species of beetles worldwide; they are often called scarabs or scarab beetles. The classification of this family has undergone significant change. Several groups formerly treated as subfamilies have been elevated to family rank (e.g., Bolboceratidae, Geotrupidae, Glaresidae, Glaphyridae, Hybosoridae, Ochodaeidae, and Pleocomidae), and some reduced to lower ranks. The subfamilies listed in this article are in accordance with those in Catalog of Life (2023). [3]
Scarabs are stout-bodied beetles, many with bright metallic colours, measuring between 1.5 and 160 millimetres (0.059 and 6.3 in). They have distinctive, clubbed antennae composed of plates called lamellae that can be compressed into a ball or fanned out like leaves to sense odours. Many species are fossorial, with legs adapted for digging. In some groups males (and sometimes females) have prominent horns on the head and/or pronotum to fight over mates or resources. [4] The largest fossil scarabaeid was Oryctoantiquus borealis with a length of 50 millimetres (2.0 in). [5]
The C-shaped larvae, called grubs, are pale yellow or white. Most adult beetles are nocturnal, although the flower chafers (Cetoniinae) and many leaf chafers (Rutelinae) are active during the day. The grubs mostly live underground or under debris, so are not exposed to sunlight. Many scarabs are scavengers that recycle dung, carrion, or decaying plant material. [6] Others, such as the Japanese beetle, are plant-eaters, wreaking havoc on various crops and vegetation.
Some of the well-known beetles from the Scarabaeidae are Japanese beetles, dung beetles, June beetles, rose chafers (Australian, European, and North American), rhinoceros beetles, Hercules beetles and Goliath beetles.
Several members of this family have structurally coloured shells which act as left-handed circular polarisers; this was the first-discovered example of circular polarization in nature. [7]
In Ancient Egypt, the dung beetle now known as Scarabaeus sacer (formerly Ateuchus sacer) was revered as sacred. [8] Egyptian amulets representing the sacred scarab beetles were traded throughout the Mediterranean world. [4]
Kheper is a subgenus of Scarabaeus: the typical genus of scarab beetles in the tribe Scarabaeini. The genus name honors the god Khepri in the ancient Egyptian religion, who is depicted as having a scarab for a head. Kheper can be found on the border between Botswana, Namibia, and South Africa in the arid sand dunes. This genus of scarab beetles share a family with the oldest and most revered scarab beetles, the Scarabaeus sacer.
Scarab may refer to:
Dung beetles are beetles that feed on feces. Some species of dung beetles can bury dung 250 times their own mass in one night.
Dynastinae or rhinoceros beetles are a subfamily of the scarab beetle family (Scarabaeidae). Other common names – some for particular groups of rhinoceros beetles – include Hercules beetles, unicorn beetles or horn beetles. Over 1,500 species and 225 genera of rhinoceros beetles are known.
The scarab beetle subfamily Scarabaeinae consists of species collectively called true dung beetles. Most of the beetles of this subfamily feed exclusively on dung. However, some may feed on decomposing matter including carrion, decaying fruits and fungi. Dung beetles can be placed into three structural guilds based on their method of dung processing namely rollers (telecoprids), dwellers (endocoprids) and tunnelers (paracoprids). Dung removal and burial by dung beetles result in ecological benefits such as soil aeration and fertilization; improved nutrient cycling and uptake by plants, increase in Pasture quality, biological control of pest flies and intestinal parasites and secondary seed dispersal. Well-known members include the genera Scarabaeus and Sisyphus, and Phanaeus vindex.
Scarabaeus sacer, common name sacred scarab, is the type species of the genus Scarabaeus and the family Scarabaeidae. This dung beetle is native of southern Europe, northern Africa and western Asia, and it was venerated in ancient Egypt.
The genus Scarabaeus consists of a number of Afro-Eurasian dung beetle species, including the "sacred scarab beetle", Scarabaeus sacer and is the namesake of the tribe Scarabaeini, the family Scarabaeidae, the superfamily Scarabaeoidea and the infraorder Scarabaeiformia. These beetles feed exclusively on dung, which they accomplish by rolling a piece of dung some distance from where it was deposited, and burying it in order to feed on it underground. They also prepare food for their larvae by excavating an underground chamber, and filling it with balls that have eggs laid in them. The growing larva feeds upon the dung ball, pupates, and eventually emerges as an adult.
Anomala binotata is a species of shining leaf chafer in the family of scarab beetles, Scarabaeidae. The common name "Shining leaf chafer" is sometimes applied to this species specifically, but is more often used to describe all members of the subfamily Rutelinae.
Euchirinae is a subfamily of Scarabaeidae or scarab beetles in the superfamily Scarabaeoidea. They are sometimes referred to as "long-armed scarabs" due to the elongated forelegs of the males. These long legs often have median and apical spines that are fixed in the male while females have a movable terminal spine.
The European chafer is a beetle of the family Scarabaeidae. Formerly found only in continental Europe, this invasive species is now found at temperate latitudes in North America. The large, white grubs of A. majale feed on the roots of most cool-latitude grasses, both wild and cultivated. This has made the European chafer an enemy of lawns.
Rutelinae or shining leaf chafers is a subfamily of the scarab beetles. It is a very diverse group; distributed over most of the world, it contains some 200 genera with over 4,000 described species in 7 tribes. Several taxa have yet to be described. A few recent classifications include the tribe Hopliini, but this is not generally accepted.
Sceliages, Westwood,, is a sub-genus of the Scarabaeus dung beetles, and are obligate predators of spirostreptid, spirobolid and julid millipedes, having renounced the coprophagy for which they were named. The genus is near-endemic to Southern Africa, Sceliages augias exceptionally ranging as far north as the Democratic Republic of Congo.
Phanaeus, the rainbow scarabs, is a genus of true dung beetles in the family Scarabaeidae, ranging from the United States to northern Argentina, with the highest species richness in Mexico. Depending on species, they can inhabit a wide range of habitats, from tropical to temperate climates and deserts to rainforests. In those living in relatively arid places adults are primarily active during the wet season and those living in relatively cold places are primarily active during the summer. They are excellent diggers and good fliers.
Pyronota festiva, commonly known as mānuka beetle or mānuka chafer, is a member of the genus Pyronota of the beetle family Scarabaeidae. It is a scarab beetle endemic to New Zealand, and is commonly found in mānuka trees, hence the beetle's name. In some areas it is considered a pasture pest.
Protognathinus is an extinct genus of stag beetle from the Eocene of Europe, known from the Messel Pit in Germany. This genus is known from the single species P. spielbergi.
Gymnetina is a genus of fruit and flower chafers in the family of beetles known as Scarabaeidae. There are about six described species in Gymnetina.
Onthophagus unifasciatus, is a species of true dung beetle native to India and Sri Lanka.
Apogonia ferruginea, is a species of dung beetle found in India and Sri Lanka.
Scarabaeus sanctus, is a species of dung beetle found in India, and Sri Lanka.
Oryctoantiquus is a genus of middle Eocene beetles in the Dynastinae clade. Oryctoantiquus is known for being the oldest known genus of Dynastinae and is also known for being the largest discovered Scarabaeidae so far. The genus only has one known species, O. borealis. The species and genus was found in Oregon, United States and dates from around 44.6 million years to 46.8 million years ago in the Clarno formation, middle Eocene epoch.
The scarab (Scarabaeus sacer) is a species of dung beetle that was considered sacred by the ancient Egyptians. In particular, the insect symbolized rebirth and was commonly associated with solar gods of creation, such as Khepri and Re.