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Blister beetle Temporal range: | |
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Hycleus lugens | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Class: | Insecta |
Order: | Coleoptera |
Superfamily: | Tenebrionoidea |
Family: | Meloidae Gyllenhaal, 1810 |
Subfamilies | |
Blister beetles are beetles of the family Meloidae, so called for their defensive secretion of a blistering agent, cantharidin. About 7,500 species are known worldwide. Many are conspicuous and some are aposematically colored, announcing their toxicity to would-be predators.
Blister beetles are hypermetamorphic, going through several larval stages, the first of which is typically a mobile triungulin. The larvae are insectivorous, mainly attacking bees, though a few feed on grasshopper eggs. While sometimes considered parasitoids, in general, the meloid larva apparently consumes the immature host along with its provisions, and can often survive on the provisions alone; thus it is not an obligatory parasitoid, but rather a facultative parasitoid, or simply a kleptoparasite. The adults sometimes feed on flowers and leaves of plants of such diverse families as the Amaranthaceae, Asteraceae, Fabaceae, and Solanaceae. [1]
Cantharidin, a poisonous chemical that causes blistering of the skin, is secreted as a defensive agent. It is used medically to remove warts [2] and is collected for this purpose from species of the genera Mylabris and Lytta , especially Lytta vesicatoria , better known as "Spanish fly".
Cantharidin is the principal irritant in "Spanish fly", a folk medicine prepared from dried beetles in the family Meloidae.
The largest genus, Epicauta , contains many species toxic to horses. A few beetles consumed in a single feeding of alfalfa hay may be lethal. [3] In semiarid areas of the western United States, modern harvesting techniques may contribute to cantharidin content in harvested forage. The practice of hay conditioning, crushing the stalks to promote drying, also crushes any beetles present and causes the release of cantharidin into the fodder. Blister beetles are attracted to alfalfa and weeds during bloom. Reducing weeds and timing harvests before and after bloom are sound management practices. Using equipment without hay conditioners may reduce beetle mortality and allow them to escape before baling. [4]
The family is thought to have begun diversifying during the Early Cretaceous. The oldest fossil of the group is a larva (triangulin) found phoretic on a schizopterid bug from the mid Cretaceous Burmese amber, dated to around 99 million years ago. [5]
Tribe Derideini
Tribe Morphozonitini
Tribe Eleticini
Tribe Spasticini
Tribe Cerocomini
Tribe Epicautini
Tribe Eupomphini
Tribe Lyttini
Tribe Meloini
Tribe Mylabrini
Tribe Pyrotini
Genera incertae sedis
Tribe Horiini
Tribe Nemognathini
Tribe Sitarini
Genera incertae sedis
Tribe Tetraonycini
The sap beetles, also known as Nitidulidae, are a family of beetles.
Lytta vesicatoria, also known as the Spanish fly, is an aposematic emerald-green beetle in the blister beetle family (Meloidae). It is distributed across Eurasia.
Cantharidin is an odorless, colorless fatty substance of the terpenoid class, which is secreted by many species of blister beetles. Its main current use in pharmacology is treating molluscum contagiosum and warts topically. It is a burn agent and poisonous in large doses, and has been historically used as aphrodisiacs. In its natural form, cantharidin is secreted by the male blister beetle, and given to the female as a copulatory gift during mating. Afterwards, the female beetle covers her eggs with it as a defense against predators.
The family Oedemeridae is a cosmopolitan group of beetles commonly known as false blister beetles, though some recent authors have coined the name pollen-feeding beetles. There are some 100 genera and 1,500 species in the family, mostly associated with rotting wood as larvae, though adults are quite common on flowers. The family was erected by Pierre André Latreille in 1810.
Ripiphoridae is a cosmopolitan family of some 450 described species of beetles sometimes called "wedge-shaped beetles". Ripiphoridae are unusual among beetle families in that many species are hypermetamorphic parasitoids, an attribute that they share with the Meloidae. Members of the family differ in their choice of hosts, but most attack various species of bees or wasps, while some others attack cockroaches or beetles. Many species of Ripiphoridae have abbreviated elytra, and flabellate or pectinate antennae.
Evaniidae is a family of parasitoid wasps also known as ensign wasps, nightshade wasps, hatchet wasps, or cockroach egg parasitoid wasps. They number around 20 extant genera containing over 400 described species, and are found all over the world except in the polar regions. The larvae of these solitary wasps are parasitoids that feed on cockroaches and develop inside the egg-cases, or oothecae, of their hosts.
Meloe is a genus of blister beetles commonly referred to as oil beetles. The name derives from their defensive strategy: when threatened by collectors or predators they release oily droplets of hemolymph from their joints. This fluid is bright orange and contains cantharidin, a poisonous chemical compound. Wiping the chemical on skin can cause blistering and painful swelling of the skin. This defensive strategy is not exclusive to this genus; all meloids possess and exude cantharidin upon threat.
Epicauta is a genus of beetles in the blister beetle family, Meloidae. The genus was first scientifically described in 1834 by Pierre François Marie Auguste Dejean. Epicauta is distributed nearly worldwide, with species native to all continents except Australia and Antarctica. Surveys have found the genus to be particularly diverse in northern Arizona in the United States. Few species occur in the Arctic, with none farther north than the southern edge of the Northwest Territories, Canada.
Hycleus phaleratus, is a species of blister beetle found in China, Taiwan, Thailand, Indonesia, Nepal, India, Sri Lanka, and Pakistan. It is sometimes considered a problem in agricultural cultivation but has been used in traditional Chinese medicine. The species was formerly placed in the genus Mylabris.
Lytta nuttalli, or Nuttall's blister beetle, is a species of North American beetle first described in 1824 by Thomas_Say. The genus Lytta is from a Latin word suggesting madness The specific nuttallii recognizes the contributions of Thomas Nuttall, a contemporary of Say.
Hycleus is a genus of blister beetle belonging to the Meloidae family found in Africa and Asia. The genus contains over 400 species, which historically have been confused with the genus Mylabris.
Gratiana boliviana is a species of beetle in the leaf beetle family, Chrysomelidae. Its common name is tropical soda apple leaf beetle. It is native to South America, where its distribution includes Argentina, Brazil, and Paraguay. It specializes on tropical soda apple, an invasive plant species. It has been released as an agent of biological pest control against the weedy plant in Florida and other parts of the United States.
Epicauta vittata is a species of beetle in the family Meloidae, the blister beetles. It is native to eastern North America, including eastern Canada and the eastern United States. It is known commonly as the striped blister beetle and the old-fashioned potato beetle. It is known as an agricultural pest.
Ceroctis capensis, or spotted blister beetle, is diurnal and endemic to Southern Africa occurring in diverse habitats, and belonging to the Meloidae or Blister beetle family. It secretes a toxic liquid from its leg joints when roughly handled, blistering human skin. This species somewhat resembles Mylabris oculata, a member of the same family.
Hycleus pustulatus is a species of blister beetle found in India, Sri Lanka, China and Java.
Meloinae is a subfamily of beetles in the family Meloidae. There are at least 330 described species in Meloinae.
Meloe americanus is a type of blister beetle (Meloidae) found in North America. It is most relevant to the fields of agriculture and veterinary medicine. Adult beetles feed on different types of plants, which cause crop damage. They also release a fluid containing a chemical that is toxic, and at high concentrations lethal, to mammals. The first instar larvae are uniquely active and mobile, utilizing phoresy and parasitism to feed and mature through their developmental stages.
Tegrodera aloga, the iron cross blister beetle, is a species of blister beetle in the family Meloidae. It is found in Arizona, California, and Sonora.
Megetra vittata is a species of blister beetle in the family Meloidae. It is found in North America. Like other blister beetles, megetra vittata excrete cantharidin, a toxic chemical, to defend itself from predators. Animals such as horses can fall ill and die from this toxin, as a result of eating many of these beetles, as they have been reported to get mixed in with their hay and other feed.
Meloetyphlus fuscatus, the blind blister beetle, is a species of blister beetle in the family Meloidae found in Central and South America. They are kleptoparasites of orchid bees and are entirely blind as adults. Unique among meloids, females do not lay their eggs near flowers, but rather within their hosts' nests.