Burmese amber

Last updated
Wing of the dragonfly Burmalindenia in a cabochon of Burmese amber, showing typical red colouration of the amber. Scale bar = 5mm Burmalindenia imperfecta whole amber.png
Wing of the dragonfly Burmalindenia in a cabochon of Burmese amber, showing typical red colouration of the amber. Scale bar = 5mm

Burmese amber, also known as Burmite or Kachin amber, is amber from the Hukawng Valley in northern Myanmar. The amber is dated to around 100 million years ago, during the latest Albian to earliest Cenomanian ages of the mid-Cretaceous period. The amber is of significant palaeontological interest due to the diversity of flora and fauna contained as inclusions, particularly arthropods including insects and arachnids but also birds, lizards, snakes, frogs and fragmentary dinosaur remains. The amber has been known and commercially exploited since the first century AD, and has been known to science since the mid-nineteenth century. Research on the deposit has attracted controversy due to the potential role of the amber trade in funding internal conflict in Myanmar and hazardous working conditions in the mines where it is collected.

Contents

Geological context, depositional environment and age

Geological context of Burmese amber in northern Myanmar Burmese amber geological context PNAS Fig1 A (cropped).jpg
Geological context of Burmese amber in northern Myanmar

The amber is found in the Hukawng Basin, a large Cretaceous-Cenozoic sedimentary basin within northern Myanmar. The strata have undergone folding and faulting. The Hukawng basin is part of the larger Myanmar Central Basin, a N-S orientated synclinal basin extending to the Gulf of Martaban to the south. The basin is considered to be a part of the West Burma block (a.k.a. Burma Terrane), which has a debated tectonic history, it is considered to be associated with the concepts of the Cimmeria and Sibumasu terranes. The block was part of Gondwana during at least the Early Paleozoic, but the timing of rifting is very uncertain, with estimates ranging from the Devonian to Early Cretaceous. It is also disputed whether the block had accreted onto the Asian continental margin by the time of the amber deposition. [1] Some members of the flora and fauna have Gondwanan affinities, [2] while others have Laurasian affinities. [3] A recent paleomagnetic reconstruction finds that the Burma Terrane formed an island land mass in the Tethys Ocean during the Mid Cretaceous at a latitude around 5-10 degrees south of the equator. [4]

At Noije Bum, located on a ridge, amber is found within fine grained clastic rocks, typically medium to greyish green in colour, resulting from the constituent grains being black, yellow, grey and light green. The fine grained rocks are primarily fine to very fine grained sandstone, with beds of silt and shale and laterally persistent thin (1–2 mm thick) coal horizons. Massive micritic limestone interbeds of 6-8 centimetre thickness, often containing coalified plant material also occur. This facies association is typically around 1 metre thick and typically thinly bedded and laminated. Associated with the fine grained facies is a set of medium facies primarily consisting of medium to fine grained sandstones also containing thin beds of siltstone, shale and conglomerate, alongside a persistent conglomerate horizon. A specimen of the ammonite Mortoniceras has been found in a sandstone bed 2 metres above the amber horizon, alongside indeterminate gastropods and bivalves. [5] Lead-uranium dating of zircon crystals of volcanic clasts within the amber bearing horizons has given a maximum age of 98.79 ± 0.62  million years ago (Ma), making the deposit earliest Cenomanian in age. [6] Unpublished data by Wang Bo on other layers suggests an age range of deposition of at least 5 million years. [7] The amber does not appear to have undergone significant transport since hardening or be redeposited. The strata at the site are younging upwards, striking north north-east and dipping 50-70 degrees E and SE north of the ridge and striking between south south-east and south-east and dipping 35-60 degrees south-west south of the ridge, suggesting the site is on the northwest limb of a syncline plunging to the northeast. A minor fault with a conspicuous gouge zone was noted as present, though it appeared to have no significant displacement. [5] Several other localities are known, including the colonial Khanjamaw and the more recent Inzutzut, Angbamo, and Xipiugong sites, within the vicinity of Tanai. The Hkamti site SW of the Hukawng basin has been determined to be significantly older, dating to the early Albian around ca. 110 Ma and is therefore considered distinct. [8]

Paleoenvironment

A Puzosia (Bhimaites) species juvenile shell in Burmese amber Puzosia Bhimaites species Burmese amber PNAS Fig2 A.jpg
A Puzosia ( Bhimaites ) species juvenile shell in Burmese amber

The Burmese amber paleoforest is considered to have been a tropical rainforest, situated near the coast, where resin was subsequently transported into a shallow marine environment. The shell of a dead juvenile Puzosia (Bhimaites) ammonite, four marine gastropod shells (including Mathilda ) and littoral or supralittoral isopods entombed in a piece of amber with shell sand, [9] along with growth of Isocrinid crinoids, corals and oysters on the surface of some amber pieces indicate marine conditions for final deposition. [10] Additionally pholadid (piddock) bivalve borings into amber specimens along with at least one pholadid which became trapped was interpreted to show that the resin was still fresh and unhardened when it was being moved into the tidal areas. [11] However, the phloladids in question, belonging to the extinct genus Palaeolignopholas, were later interpreted as a freshwater species, and the presence of numerous freshwater insects suggests that the initial environment of deposition was a downstream estuarine to freshwater section of a river, with the forests extending across coastal rivers, river deltas, lakes, lagoons, and coastal bays. [12] The forest environment may have been prone to fire, similar to modern tropical peat swamps, based on the presence of fire adapted plants and burned plant remains found in the amber. [13]

The amber itself is primarily disc-shaped and flattened along the bedding plane, and is typically reddish brown, with the colour ranging from shades of yellow to red. The opacity of the amber ranges from clear to opaque. Many amber pieces have thin calcite veins that are typically less than 1 mm (0.04 in), but up to 4–5 mm (0.16–0.20 in) thick. The number and proportion of veins in a piece of amber varies significantly, in some pieces veins are virtually absent, while others are described as being "packed with veinlets". [5] The amber is considered to be of coniferous origin, with a likely araucarian source tree, based on spectroscopic analysis and wood fragment inclusions, [14] though a pine origin has also been suggested. [15]

Fauna and flora

The list of taxa is extraordinarily diverse, with 50 classes (or equivalent), 133 orders (or equivalent), 726 families, 1,757 genera and 2,770 species described as of 2023. The vast majority of the species are arthropods, mostly insects. [16]

Arthropods

Fossil of the tailed stem-group spider Chimerarachne in Burmese amber M466395.jpg
Fossil of the tailed stem-group spider Chimerarachne in Burmese amber

Over 2500 species of arthropods (with over 2000 of these species beings insects) are known from the deposit, [16] notably including the oldest members of the arachnid orders Palpigradi ( Electrokoenenia ) [17] and Schizomida [18] and the only known fossil members of Ricinulei since the Paleozoic. [19] Chimerarachne is a unique stem-group spider still possessing a tail, with similar forms only known from the Paleozoic, [20] with over 360 other species of spiders (including the only non Paleozoic fossils of Mesothelae) being known from Burmese amber, along with over 40 species of scorpions. [16] Other significant arachnid fossils include some of the oldest ticks, [21] as well as the oldest mesostigmatan [22] and opilioacarid mites. [23] Beetles (Coleoptera), reflecting their modern diversity, are represented by over 100 famiilies and over 500 species, with Hymenoptera (bees, wasps, ants and sawflies) represented by 70 families and 350 species including some of the oldest known ants. Flies (Diptera) are represented by over 50 families and over 250 species and true bugs (Hemiptera) are represented by over 80 families and over 220 species. Dictyoptera (cockroaches, termites and mantises, among others), are represented by over 100 species in 28 families. Odonata (which contains dragonflies and damselflies, among other extinct groups), is represented by over 19 families and over 40 species, [16] far higher than in other Cretaceous amber deposits. [24] Among the oldest members of the insect order Zoraptera are known from Burmese amber, [25] as well as the oldest parasitic lice [26] and the oldest members of the parasitic insect order Strepsiptera. [27] The winged insect Aristovia is transitional between the extinct "Grylloblattodea" and modern wingless Grylloblattidae (ice crawlers). [28] Over a dozen species each of Myriapoda (millipedes and centipedes, among others) and Entognatha (springtails, among others) have also been reported, along with a number of woodlice (representing some of the oldest records of the group) and various aquatic crustaceans. [16]

Other invertebrates

A wide variety of other invertebrates have been reported. These include gastropods, including freshwater and land snails, [16] the oldest modern onychophoran (also known as velvet worms) Cretoperipatus, [29] as well as nematodes, nematomorphs, annelids and flatworms. [16]

Vertebrates

Fossil of the frog Electrorana in Burmese amber Fossil frog in amber.png
Fossil of the frog Electrorana in Burmese amber

Only a handful of vertebrates have been described from Burmese amber, these include the albanerpetontid (a group of extinct salamander-like amphibians) Yaksha , the frog Electrorana , a number of primitive toothed birds belonging to the extinct clade Enantiornithes, including the named species Elektorornis and Fortipesavis [16] , the lizards Protodraco (suggested to be the oldest agamid [30] ), Cretaceogekko (suggested to be the oldest modern gecko [31] ), Electroscincus (the oldest known skink [32] ) Barlochersaurus , [31] Oculudentavis (formerly erroneously considered to be a bird), [33] and the snake Xiaophis. [34] An indeterminate theropod dinosaur is known from a tail with preserved feathers. [35]

Flora

A wide variety of plants have been reported from the deposit. These include flowering plants, conifers, ferns, lycophytes and bryophytes. [16]

Other

A number of fungi species have been reported, as well as various microorganisms. [16]

History

The amber is recorded as originating from Yunnan Province as early as the first century AD according to the Book of the Later Han. The trade of Burmese amber into China during the Han Dynasty has been confirmed by Fourier-transform infrared spectroscopy (FTIR) analysis of amber artifacts from the Eastern Han Dynasty (25 - 220 CE). [36] It was first mentioned in European sources by the Jesuit Priest Álvaro Semedo who visited China in 1613, it was described as being "digged out of mines, and sometimes in great pieces, it is redder than our amber though not so cleane". [37] The locality itself has been known to European explorers since the 1800s with visitation to the Hukawng Valley by Simon Fraser Hannay in 1836–1837. [38] [39] At that time the principle products of the valley mines were salt, gold, and amber, with the majority of gold and amber being bought by Chinese traders. Hannay visited the amber mines themselves on March 21, 1836, and he noted that the last three miles to the mines were marked with numerous abandoned pits, up to 15 ft (4.6 m) in depth, where amber had been dug in the past. The mining had moved over the hill to a series of 10 pits but no visible amber was seen, suggesting that miners possibly hid the amber found that day before the party arrived. Mining was being performed manually at the time through the use of sharpened bamboo rods and small wooden shovels. Finer pieces of amber were recovered from the deeper pits, with clear yellow being recovered from depths of 40 ft (12 m) The recovered amber was bought with silver or often exchanged for jackets, hats, copper pots, or opium among other goods. mixed and lower quality amber was sold from around ⁠1/⁠ ticals to 4 rupees per seer. Pieces that were considered high quality or fit for use as ornamentation were described as expensive, and price varied depending on the clarity and color of the amber. Women of the valley were noted to wear amber earrings as part of their jewelry. [38] In 1885 the Konbaung dynasty was annexed to the British Raj and a survey of the area was conducted by Dr. Fritz Noetling on behalf of the Geological Survey of India. [37] The final research before Burmese independence in 1947 was conducted by Dr. H.L. Chhibber in 1934, who provided the most detailed description of Burmite occurrences. [37]

History of research

While a 1892 study considered the amber likely to be Miocene in age, a study by Theodore Dru Alison Cockerell in 1916 noted archaic nature of the insects, and concluded that the amber must be older, and possibly as old as the Late Cretaceous. Various later authors during the mid-late 20th century suggested either a Paleocene-Eocene or an Late Cretaceous age. A Cretaceous age was confirmed during the early 2000s. [37] While during the 20th century and the first decade of the 21st century research on the deposit was low intensity, during the mid-2010s there was a great increase number of papers published on the deposit, numbering hundreds every year, with a plurality coming from Chinese researchers, [40] rising from a few dozen species described per year prior to 2014 to over 350 in 2020, though this number had fallen to just over 250 by 2023. [16] Both the great increase in the amber volume coming out of the mines, as well as a 2013 conference and a 2015 Burmese amber special issue in the journal Cretaceous Research have been cited as factors in the boom of publications. [41]

Modern exploitation and controversy

Miners digging and sorting Burmese amber at Noije Bum in the Hukawng Valley, undated photograph Noje Bum amber.jpg
Miners digging and sorting Burmese amber at Noije Bum in the Hukawng Valley, undated photograph

Leeward Capital Corp, a small Canadian mining firm, began exporting amber from Myanmar during the 1990s, resulting in the description of a number of fossil species from the deposit. [42] During the late 2000s, Burmese amber production began to increase as a result of the exhaustion of Chinese amber mines and increased trade between the Myanmar and China generally, with the amber trade being initially unaffected by the outbreak of renewed conflict between the Kachin Independence Army, an armed rebel group seeking to scede from Myanmar, and the Burmese armed forces (Tatmadaw) in 2011. Despite the conflict between them, there was apparently a tacit agreement between the two groups to keep the amber trade flowing, with the KIA controlling the mines, while the Tatmadaw controlled their export to China, with both collecting taxes on the trade. [43] In 2016, the KIA took full control of the amber mines, and controlled amber export via "various licenses, taxes, restrictions on the movement of labor and enforced auctions". [44] In June 2017 the Tatmadaw seized control of the mines from the KIA. [45]

The main amber market in Myanmar is Myitkyina. Most Burmese amber is exported (primarily smuggled) into China, with the primarily Chinese market being in Tengchong, Yunnan, with an estimated 100 tonnes of Burmese amber passing into the city 2015, with a then estimated value between five and seven billion yuan, where it is primarily used for jewelry. Burmese amber was estimated to make up 30% of Tenchong's gemstone market (the rest being Myanmar Jade), and was declared one of the city's eight main industries by the local government. [46] The presence of calcite veins are a major factor in determining the gem quality of pieces, with pieces with a large number of veins having significantly lower value. [5]

The working conditions at the mines have been described as extremely unsafe, down 100 m (330 ft) deep pits barely wide enough to crawl through, with no accident compensation. [7] Sales of amber were alleged to help fund the Kachin conflict by various news organisations in 2019. [47] [48] Interest in this discussion rose in March 2020 after the highly publicised description of Oculudentavis, which made the cover of Nature. [49] On April 21, 2020, the Society of Vertebrate Paleontology (SVP) published a letter of recommendation to journal editors asking for “a moratorium on publication for any fossil specimens purchased from sources in Myanmar after June 2017 when the Myanmar military began its campaign to seize control of the amber mining”. [50] On April 23, 2020 Acta Palaeontologica Polonica stated that it would not accept papers on Burmese amber material collected from 2017 onwards, after the Burmese military took control of the deposit, requiring "certification or other demonstrable evidence, that they were acquired before the date both legally and ethically". [51] On May 13, 2020, the Journal of Systematic Palaeontology published an editorial stating that it would no longer consider papers based whole or in part on Burmese amber material, regardless of whether in historic collections or not. [52] On 30 June 2020, a statement from the International Palaeoentomological Society was published in response to the SVP, criticising the proposal to ban publishing on Burmese amber material. [53] In August 2020, a comment from over 50 authors was published in PalZ responding to the SVP statement. The authors disagreed with the proposal of a moratorium, describing the focus on the Burmese amber as "arbitrary" and that "The SVP’s recommendation for a moratorium on Burmese amber affects fossil non-vertebrate research much more than fossil vertebrate research and clearly does not represent this part of the palaeontological community." [54]

The conclusion that Burmese amber funded the Tatmadaw was disputed by George Poinar and Sieghard Ellenberger, who found that the supply of amber collapsed after the 2017 takeover of the mines by the Tatmadaw, and that most of the current circulation of amber in Chinese markets was extracted prior to 2017. [55] A story in Science in 2019 stated: "Two former mine owners, speaking through an interpreter in phone interviews, say taxes have been even steeper since government troops took control of the area. Both shut their mines when they became unprofitable after the government takeover, and almost all deep mines are now out of business, dealers here corroborate. Only shallow mines and perhaps a few secret operations are still running." [7] There were around 200,000 miners working in the Hukawng valley mines prior to the takeover by the Tatmadaw, which shrunk to 20,000 or less after the military operations. Adolf Peretti, a gemologist who owns a museum with Burmese amber specimens, noted that the 2017 cutoff suggested by the SVP does not take into account that the export of Burmese amber prior to 2017 was also funding internal conflict in Myanmar due to the control by the KIA. Much of the amber cutting since 2017 has been done in internally displaced person camps, under humanitarian and non-conflict conditions. [44]

Other Burmese ambers

Other deposits of amber are known from several regions in Myanmar, with noted deposits in the Shwebo District of the Sagaing Region, from the Pakokku and Thayet districts of Magway Region and the Bago District of the Bago Region. [56] [39]

Tilin amber

A 2018 study on an amber deposit from Tilin in central Myanmar indicated that deposit to be 27 million years younger than the Hukawng deposit, dating to approximately 72 million years old, placing it in the latest Campanian age. The deposit was associated with an overlying tuffaceous layer, and underlying nodules of brown sandstone yielded remains of the ammonite Sphenodiscus. Within a number of arthropod specimens were described though much more poorly preserved than specimens in the Hukawng amber. These include members of Hymenoptera (Braconidae, Diapriidae, Scelionidae) Diptera (Ceratopogonidae, Chironomidae) Dictyoptera (Blattaria, Mantodea) planthoppers, Berothidae and bark lice (Lepidopsocidae) as well as extant ant subfamilies Dolichoderinae and tentatively Ponerinae, as well as fragments of moss. [57]

Hkamti amber

The Hkamti site is located ca. 90 km southwest of the Angbamo site and predominantly consists of limestone, interbedded with mudstone and tuff, the amber is found in the unconsolidated mudstone/tuff layers. A crinoid was found attached to one amber specimen, alongside marine plant remains in the surrounding sediment, indicating deposition in a shallow marine setting. The amber is generally red-brown, and yellow colouration is rare, the amber is generally found as angular clasts, indicating short transport distance and is more brittle than other northern Myanmar ambers. Zircon dating has constrained the age of the deposit to the early Albian, c. 110 Ma, significantly older than the dates obtained from other deposits. Fauna found in the amber include: Archaeognatha, Diplopoda, Coleoptera, Araneae, Trichoptera, Neuroptera, Psocodea, Isoptera Diptera, Orthoptera, Pseudoscorpionida, Hymenoptera and Thysanoptera. [8] A lizard, Retinosaurus, has also been described from the locality. [58]

See also

Related Research Articles

Michael S. Engel, FLS, FRES is an American paleontologist and entomologist, notable for contributions to insect evolutionary biology and classification. In connection with his studies he has undertaken field expeditions in Central Asia, Asia Minor, the Levant, Arabia, eastern Africa, the high Arctic, and South and North America, and has published more than 925 papers in scientific journals. Some of Engel's research images were included in exhibitions on the aesthetic value of scientific imagery.

<span class="mw-page-title-main">Hemiphlebiidae</span> Family of damselflies

Hemiphlebiidae is a family of damselflies. It contains only one extant species, the ancient greenling, native to Southern Australia and Tasmania. The fossil record of the group extends back to the Late Jurassic, making them the oldest known crown group damselflies.

2018 in paleoentomology is a list of new fossil insect taxa that were described during the year 2018, as well as other significant discoveries and events related to paleoentomology that were scheduled to occur during the year.

2019 in paleoentomology is a list of new fossil insect taxa that were described during the year 2019, as well as other significant discoveries and events related to paleoentomology that were scheduled to occur during the year.

2017 in paleoentomology is a list of new fossil insect taxa that were described during the year 2017, as well as other significant discoveries and events related to paleoentomology that were scheduled to occur during the year.

Burmese amber is fossil resin dating to the early Late Cretaceous Cenomanian age recovered from deposits in the Hukawng Valley of northern Myanmar. It is known for being one of the most diverse Cretaceous age amber paleobiotas, containing rich arthropod fossils, along with uncommon vertebrate fossils and even rare marine inclusions. A mostly complete list of all taxa described up until 2018 can be found in Ross 2018; its supplement Ross 2019b covers most of 2019.

Discoscapa apicula is an extinct species of crabronid wasp, formerly considered one of the two oldest-known species of bees. The species was described from an amber inclusion in Burmese Amber in 2020 by George Poinar Jr., a zoologist at Oregon State University. The fossil was found in a mine in the Hukawng Valley of northern Myanmar and is believed to date from the Cretaceous Period, 100 million years ago, the same age as Melittosphex burmensis, likewise previously considered the oldest known bee species; as it comes from the same amber deposit, these two specimens are considered to be the same approximate age. More recent research has concluded that D. apicula is a wasp belonging to the subfamily Crabroninae, placed in its own tribe, Discoscapini.

2020 in paleoentomology is a list of new fossil insect taxa that were described during the year 2020, as well as other significant discoveries and events related to paleoentomology that were scheduled to occur during the year.

2015 in paleoentomology is a list of new fossil insect taxa that were described during the year 2016, as well as other significant discoveries and events related to paleoentomology that were scheduled to occur during the year.

2015 in paleoentomology is a list of new fossil insect taxa that were described during the year 2015, as well as other significant discoveries and events related to paleoentomology that were scheduled to occur during the year.

Burmese amber is fossil resin dating to the early Late Cretaceous Cenomanian age recovered from deposits in the Hukawng Valley of northern Myanmar. It is known for being one of the most diverse Cretaceous age amber paleobiotas, containing rich arthropod fossils, along with uncommon vertebrate fossils and even rare marine inclusions. A mostly complete list of all taxa described up until 2018 can be found in Ross 2018; its supplement Ross 2019b covers most of 2019.

<span class="mw-page-title-main">Mimarachnidae</span> Extinct family of true bugs

Mimarachnidae is an extinct family of planthoppers known from the Cretaceous period. Their name is derived from spots on the wings of the first described genera, Mimarachne and Saltissus, being suggestive of spider mimicry, but these characters are not distinctive for the family as a whole. The family is characterised by "simplified venation and setigerous metatibial pecten and hind leg armature". as well as "rounded anterior margin of pronotum, double carination of pronotum and mesonotum"

<span class="mw-page-title-main">Yuripopovinidae</span> Extinct family of true bugs

Yuripopovinidae is an extinct family of Coreoidea Hemipteran true bugs. Member species are known from the Early Cretaceous and early Late Cretaceous of Asia and northern Gondwana. Among the distinguishing characters are "the hemelytral costal vein apically much thickened and pterostigma-like, the corium with two large cells separated by one longitudinal straight vein." Dehiscensicoridae, described from the Yixian Formation of China has been deemed a junior synonym of Yuripopovinidae per Du et al. (2019). The family was named after Russian paleoentomologist Yuri Alexandrovich Popov.

Burmese amber is fossil resin dating to the early Late Cretaceous Cenomanian age recovered from deposits in the Hukawng Valley of northern Myanmar. It is known for being one of the most diverse Cretaceous age amber paleobiotas, containing rich arthropod fossils, along with uncommon vertebrate fossils and even rare marine inclusions. A mostly complete list of all taxa described up until 2018 can be found in Ross 2018; its supplement Ross 2019b covers most of 2019.

Jouault, Corentin; Rosse-Guillevic, Simon. "A new genus of praeaulacid wasp from the mid-Cretaceous Kachin amber (Myanmar)". Annales de Paléontologie. 109 (1): 102599. doi:10.1016/j.annpal.2023.102599.

This paleoentomology list records new fossil insect taxa that were to be described during the year 2021, as well as notes other significant paleoentomology discoveries and events which occurred during that year.

This paleoentomology list records new fossil insect taxa that are to be described during the year 2022, as well as notes other significant paleoentomology discoveries and events which occurred during that year.

This list of 2023 in paleoentomology records new fossil insect taxa that are to be described during the year, as well as documents significant paleoentomology discoveries and events which occurred during that year.

<i>Paleotriatoma</i> Extinct species of true bug

Paleotriatoma metaxytaxa is a species of fossil insect belonging to the subfamily Triatominae of the family Reduviidae. Living kissing bugs are blood-sucking insects responsible for the transmission of Chagas disease. Chagas is a parasitic disease affecting millions of people mainly in South America, Central America and Mexico.

This list of 2024 in paleoentomology records new fossil insect taxa that are to be described during the year, as well as documents significant paleoentomology discoveries and events which occurred during that year.

References

  1. ↑ Metcalfe, Ian (June 2017). "Tectonic evolution of Sundaland". Bulletin of the Geological Society of Malaysia. 63: 27–60. doi: 10.7186/bgsm63201702 .
  2. ↑ Poinar, George (2018-03-09). "Burmese amber: evidence of Gondwanan origin and Cretaceous dispersion". Historical Biology: 1–6. doi:10.1080/08912963.2018.1446531. ISSN   0891-2963. S2CID   90373037.
  3. ↑ Hinkelman, Jan (2022-01-10). "Mongolblatta sendii sp. n. (Mesoblattinidae) from North Myanmar amber links record to Laurasian sediments". Palaeontographica Abteilung A. 321 (1–6): 81–96. Bibcode:2022PalAA.321...81H. doi:10.1127/pala/2021/0105. ISSN   0375-0442. S2CID   240162578.
  4. ↑ Westerweel, Jan; Roperch, Pierrick; Licht, Alexis; Dupont-Nivet, G.; Win, Zaw; Poblete, Fernando; Ruffet, Gilles; Swe, Hnin; Thi, Myat; Aung, Day (2019-10-01). "Burma Terrane part of the Trans-Tethyan Arc during collision with India according to palaeomagnetic data". Nature Geoscience. 12 (10): 863–868. Bibcode:2019NatGe..12..863W. doi:10.1038/s41561-019-0443-2. PMC   6774779 . PMID   31579400.
  5. 1 2 3 4 Cruickshank, R.D; Ko, Ko (February 2003). "Geology of an amber locality in the Hukawng Valley, Northern Myanmar". Journal of Asian Earth Sciences. 21 (5): 441–455. Bibcode:2003JAESc..21..441C. doi:10.1016/S1367-9120(02)00044-5.
  6. ↑ Shi, Guanghai; Grimaldi, David A.; Harlow, George E.; Wang, Jing; Wang, Jun; Yang, Mengchu; Lei, Weiyan; Li, Qiuli; Li, Xianhua (October 2012). "Age constraint on Burmese amber based on U–Pb dating of zircons". Cretaceous Research. 37: 155–163. Bibcode:2012CrRes..37..155S. doi:10.1016/j.cretres.2012.03.014.
  7. 1 2 3 Sokol, Joshua (2019-05-23). "Fossils in Burmese amber offer an exquisite view of dinosaur times—and an ethical minefield". Science. doi:10.1126/science.aay1187. ISSN   0036-8075. S2CID   194317225.
  8. 1 2 Xing, Lida; Qiu, Liang (November 2020). "Zircon U Pb age constraints on the mid-Cretaceous Hkamti amber biota in northern Myanmar". Palaeogeography, Palaeoclimatology, Palaeoecology. 558: 109960. Bibcode:2020PPP...55809960X. doi:10.1016/j.palaeo.2020.109960. S2CID   224899464.
  9. ↑ Yu, T.; Kelly, R.; Mu, L; Ross, A.; Kennedy, J.; Broly, P.; Xia, F.; Zhang, H.; Wang, B.; Dilcher, D. (2019-06-04). "An ammonite trapped in Burmese amber". Proceedings of the National Academy of Sciences. 116 (23): 11345–11350. Bibcode:2019PNAS..11611345Y. doi: 10.1073/pnas.1821292116 . ISSN   0027-8424. PMC   6561253 . PMID   31085633.
  10. ↑ Mao, Y.; Liang, K.; Su, Y.; Li, J.; Rao, X.; Zhang, H.; Xia, F.; Fu, Y.; Cai, C.; Huang, D. (2018-12-28). "Various amberground marine animals on Burmese amber with discussions on its age". Palaeoentomology. 1 (1): 91. doi:10.11646/palaeoentomology.1.1.11. ISSN   2624-2834. S2CID   68048811.
  11. ↑ Smith, Ru D. A.; Ross, Andrew J. (January 2018). "Amberground pholadid bivalve borings and inclusions in Burmese amber: implications for proximity of resin-producing forests to brackish waters, and the age of the amber". Earth and Environmental Science Transactions of the Royal Society of Edinburgh. 107 (2–3): 239–247. doi:10.1017/S1755691017000287. ISSN   1755-6910. S2CID   204250232.
  12. ↑ Bolotov, Ivan N.; Aksenova, Olga V.; Vikhrev, Ilya V.; Konopleva, Ekaterina S.; Chapurina, Yulia E.; Kondakov, Alexander V. (2021-03-23). "A new fossil piddock (Bivalvia: Pholadidae) may indicate estuarine to freshwater environments near Cretaceous amber-producing forests in Myanmar". Scientific Reports. 11 (1): 6646. Bibcode:2021NatSR..11.6646B. doi: 10.1038/s41598-021-86241-y . ISSN   2045-2322. PMC   7988128 . PMID   33758318.
  13. ↑ Shi, Chao; Wang, Shuo; Cai, Hao-hong; Zhang, Hong-rui; Long, Xiao-xuan; Tihelka, Erik; Song, Wei-cai; Feng, Qi; Jiang, Ri-xin; Cai, Chen-yang; Lombard, Natasha; Li, Xiong; Yuan, Ji; Zhu, Jian-ping; Yang, Hui-yu (February 2022). "Fire-prone Rhamnaceae with South African affinities in Cretaceous Myanmar amber". Nature Plants. 8 (2): 125–135. Bibcode:2022NatPl...8..125S. doi:10.1038/s41477-021-01091-w. ISSN   2055-0278. PMID   35102275. S2CID   246443363.
  14. ↑ Poinar, G.; Lambert, J.B.; Wu, Y. (2007-08-10). "Araucarian source of fossiliferous Burmese amber: Spectroscopic and anatomical evidence". Journal of the Botanical Research Institute of Texas. 1: 449–455.
  15. ↑ Dutta, S.; Mallick, M.; Kumar, K.; Mann, U.; Greenwood, P. F. (January 2011). "Terpenoid composition and botanical affinity of Cretaceous resins from India and Myanmar". International Journal of Coal Geology. 85 (1): 49–55. Bibcode:2011IJCG...85...49D. doi:10.1016/j.coal.2010.09.006.
  16. 1 2 3 4 5 6 7 8 9 10 11 Ross, Andrew J. (2024-03-28). "Complete checklist of Burmese (Myanmar) amber taxa 2023". Mesozoic. 1 (1): 21–57. doi: 10.11646/mesozoic.1.1.4 . ISSN   3021-1867.
  17. ↑ Engel, Michael S.; Breitkreuz, Laura C. V.; Cai, Chenyang; Alvarado, Mabel; Azar, Dany; Huang, Diying (15 February 2016). "The first Mesozoic microwhip scorpion (Palpigradi): a new genus and species in mid-Cretaceous amber from Myanmar". The Science of Nature. 103 (3–4): 19. Bibcode:2016SciNa.103...19E. doi:10.1007/s00114-016-1345-4. PMID   26879963. S2CID   14816297.
  18. ↑ Müller, Sandro P.; Dunlop, Jason A.; Kotthoff, Ulrich; Hammel, Jörg U.; Harms, Danilo (February 2020). "The oldest short-tailed whipscorpion (Schizomida): A new genus and species from the Upper Cretaceous amber of northern Myanmar". Cretaceous Research. 106: 104227. Bibcode:2020CrRes.10604227M. doi:10.1016/j.cretres.2019.104227. S2CID   202899476.
  19. ↑ Botero-Trujillo, Ricardo; Davis, Steven R.; Michalik, Peter; Prendini, Lorenzo (2022-09-22). "Hirsutisoma grimaldii sp. nov., a ca. 99-million-year-old ricinuleid (Primoricinulei, Hirsutisomidae) from Cretaceous Burmese amber with a corticolous, scansorial lifestyle". Palaeoentomology. 5 (5). doi:10.11646/palaeoentomology.5.5.11. ISSN   2624-2834.
  20. ↑ Dunlop, Jason A. (2022-06-27). "Spider Origins: a Palaeontological Perspective". Arachnology. 19 (sp1). doi:10.13156/arac.2022.19.sp1.182. ISSN   2050-9928. S2CID   250148030.
  21. ↑ Chitimia-Dobler, Lidia; Mans, Ben J.; Handschuh, Stephan; Dunlop, Jason A. (2022). "A remarkable assemblage of ticks from mid-Cretaceous Burmese amber". Parasitology. 149 (6): 820–830. doi: 10.1017/S0031182022000269 . ISSN   0031-1820. PMC   10090602 . PMID   35241194. S2CID   247227499.
  22. ↑ Joharchi, Omid; Vorontsov, Dmitry D.; Walter, David Evans (2021-09-14). "Oldest determined record of a mesostigmatic mite (Acari: Mesostigmata: Sejidae) in Cretaceous Burmese amber". Acarologia. 61 (3): 641–649. doi: 10.24349/goj5-bzms . ISSN   0044-586X. S2CID   239420481.
  23. ↑ Jason A. Dunlop & Leopoldo Ferreira de Oliveira Bernardi (2014). "An opilioacarid mite in Cretaceous Burmese amber". Naturwissenschaften. 101 (9): 759–763. Bibcode:2014NW....101..759D. doi:10.1007/s00114-014-1212-0. PMID   25027588. S2CID   253637881.
  24. ↑ Zheng, Daran; Jarzembowski, Edmund A. (2020-01-02). "A brief review of Odonata in mid-Cretaceous Burmese amber". International Journal of Odonatology. 23 (1): 13–21. Bibcode:2020IJOdo..23...13Z. doi:10.1080/13887890.2019.1688499. ISSN   1388-7890.
  25. ↑ Yin, Ziwei; Cai, Chenyang; Huang, Diying (April 2018). "New zorapterans (Zoraptera) from Burmese amber suggest higher paleodiversity of the order in tropical forests". Cretaceous Research. 84: 168–172. Bibcode:2018CrRes..84..168Y. doi: 10.1016/j.cretres.2017.11.028 .
  26. ↑ Zhang, Yanjie; Rasnitsyn, Alexandr P.; Zhang, Weiwei; Song, Fan; Shih, Chungkun; Ren, Dong; Wang, Yongjie; Li, Hu; Gao, Taiping (February 2024). "Stem chewing lice on Cretaceous feathers preserved in amber". Current Biology. 34 (4): 916–922.e1. Bibcode:2024CBio...34E.916Z. doi:10.1016/j.cub.2024.01.027. PMID   38320551.
  27. ↑ Pohl, Hans; Wipfler, Benjamin; Boudinot, Brendon; Beutel, Rolf Georg (2020). "On the value of Burmese amber for understanding insect evolution: Insights from †Heterobathmilla – an exceptional stem group genus of Strepsiptera (Insecta)". Cladistics. 37 (2): 211–229. doi: 10.1111/cla.12433 . ISSN   1096-0031. PMID   34478185.
  28. ↑ Cui, Yingying; Bardin, Jérémie; Wipfler, Benjamin; Demers-Potvin, Alexandre; Bai, Ming; Tong, Yi-Jie; Chen, Grace Nuoxi; Chen, Huarong; Zhao, Zhen-Ya; Ren, Dong; Béthoux, Olivier (2024-03-07). "A winged relative of ice-crawlers in amber bridges the cryptic extant Xenonomia and a rich fossil record". Insect Science. 31 (5): 1645–1656. doi:10.1111/1744-7917.13338. ISSN   1672-9609. PMID   38454304.
  29. ↑ Oliveira, I.; Bai, M.; Jahn, H.; Gross, V.; Martin, C.; Hammel, J. U.; Zhang, W.; Mayer, G. (2016). "Earliest Onychophoran in Amber Reveals Gondwanan Migration Patterns". Current Biology. 26 (19): 2594–2601. Bibcode:2016CBio...26.2594O. doi: 10.1016/j.cub.2016.07.023 . PMID   27693140.
  30. ↑ Wagner, Philipp; Stanley, Edward L.; Daza, Juan D.; Bauer, Aaron M. (August 2021). "A new agamid lizard in mid-Cretaceous amber from northern Myanmar". Cretaceous Research. 124: 104813. Bibcode:2021CrRes.12404813W. doi:10.1016/j.cretres.2021.104813.
  31. 1 2 Wang, Miaoyan; Xing, Lida (2020-03-04). "A Brief Review of Lizard Inclusions in Amber". Biosis:Biological Systems. 1 (1): 39–53. doi: 10.37819/biosis.001.01.0050 . ISSN   2707-9783.
  32. ↑ Daza, J. D.; Stanley, E. L.; Heinicke, M. P.; Leah, C.; Doucet, D. S.; Fenner, K. L.; Arias, J. S.; Smith, R. D. A.; Peretti, A. M.; Aung, N. N.; Bauer, A. M. (2024). "Compound osteoderms preserved in amber reveal the oldest known skink". Scientific Reports. 14 (1). 15662. Bibcode:2024NatSR..1415662D. doi: 10.1038/s41598-024-66451-w . PMC   11231356 . PMID   38977836.
  33. ↑ Bolet, Arnau; Stanley, Edward L.; Daza, Juan D.; Arias, J. Salvador; Čerňanský, Andrej; Vidal-García, Marta; Bauer, Aaron M.; Bevitt, Joseph J.; Peretti, Adolf; Evans, Susan E. (August 2021). "Unusual morphology in the mid-Cretaceous lizard Oculudentavis". Current Biology. 31 (15): 3303–3314.e3. Bibcode:2021CBio...31E3303B. doi:10.1016/j.cub.2021.05.040. hdl: 11336/148383 . PMID   34129826.
  34. ↑ Xing, Lida; Caldwell, Michael W.; Chen, Rui; Nydam, Randall L.; Palci, Alessandro; Simões, Tiago R.; McKellar, Ryan C.; Lee, Michael S. Y.; Liu, Ye; Shi, Hongliang; Wang, Kuan; Bai, Ming (2018-07-06). "A mid-Cretaceous embryonic-to-neonate snake in amber from Myanmar". Science Advances. 4 (7): eaat5042. Bibcode:2018SciA....4.5042X. doi:10.1126/sciadv.aat5042. ISSN   2375-2548. PMC   6051735 . PMID   30035227.
  35. ↑ Xing, Lida; McKellar, Ryan C.; Xu, Xing; Li, Gang; Bai, Ming; Persons, W. Scott; Miyashita, Tetsuto; Benton, Michael J.; Zhang, Jianping; Wolfe, Alexander P.; Yi, Qiru; Tseng, Kuowei; Ran, Hao; Currie, Philip J. (December 2016). "A Feathered Dinosaur Tail with Primitive Plumage Trapped in Mid-Cretaceous Amber". Current Biology. 26 (24): 3352–3360. Bibcode:2016CBio...26.3352X. doi:10.1016/j.cub.2016.10.008. hdl: 1983/d3a169c7-b776-4be5-96af-6053c23fa52b . PMID   27939315.
  36. ↑ Chen, Dian; Zeng, Qingshuo; Yuan, Ye; Cui, Benxin; Luo, Wugan (November 2019). "Baltic amber or Burmese amber: FTIR studies on amber artifacts of Eastern Han Dynasty unearthed from Nanyang". Spectrochimica Acta Part A: Molecular and Biomolecular Spectroscopy. 222: 117270. Bibcode:2019AcSpA.22217270C. doi:10.1016/j.saa.2019.117270. PMID   31226615. S2CID   195261188.
  37. 1 2 3 4 Zherikhin, V.V., Ross, A.J., 2000. A review of the history, geology and age of Burmese amber (Burmite). Bulletin of the Natural History Museum, London (Geology) 56 (1), 3–10.
  38. 1 2 Pemberton, R. B. (1837). "Abstract of the Journal of a route travelled by Cap. Hannay from the Capital of Ava to the Amber Mines of the Hukong valley in the South east frontier of Assam". Journal of the Asiatic Society of Bengal. 64: 248–278.
  39. 1 2 Ross, A.; Mellish, C.; York, P.; Craighton, B. (2010). "Chapter 12: Burmese amber". In Penney, D. (ed.). Biodiversity of Fossils in Amber from the Major World Deposits. Siri Scientific Press. pp. 116–136. ISBN   978-0-9558636-4-6.
  40. ↑ "Research on amber from a war-torn part of Myanmar is surging". The Economist. ISSN   0013-0613 . Retrieved 2023-07-04.
  41. ↑ Ortega, Rodrigo Pérez (2022-09-29). Violent conflict in Myanmar linked to boom in fossil amber research, study claims (Report). Science. doi:10.1126/science.adf0973.
  42. ↑ "The return of Burmite amber". Deposits. 2022-04-17. Retrieved 2024-10-24.
  43. ↑ Dunne, Emma M.; Raja, Nussaïbah B.; Stewens, Paul P.; Zin-Maung-Maung-Thein; Zaw, Khin (2022-09-29). "Ethics, law, and politics in palaeontological research: The case of Myanmar amber". Communications Biology. 5 (1): 1023. doi:10.1038/s42003-022-03847-2. ISSN   2399-3642. PMC   9522859 . PMID   36175597.
  44. 1 2 Peretti, Adolf (December 2021). "An alternative perspective for acquisitions of amber from Myanmar including recommendations of the United Nations Human Rights Council". Journal of International Humanitarian Action. 6 (1): 12. doi: 10.1186/s41018-021-00101-y . ISSN   2364-3412. S2CID   235174183.
  45. ↑ Lawton, Graham. "Blood amber: The exquisite trove of fossils fuelling war in Myanmar". New Scientist. Retrieved 2020-02-04.
  46. ↑ Rippa, Alessandro; Yang, Yi (July 2017). "The Amber Road: Cross-Border Trade and the Regulation of the Burmite Market in Tengchong, Yunnan". TRaNS: Trans -Regional and -National Studies of Southeast Asia. 5 (2): 243–267. doi: 10.1017/trn.2017.7 . ISSN   2051-364X.
  47. ↑ Gammon, Katharine. "The Human Cost of Amber". The Atlantic. ISSN   1072-7825 . Retrieved 2020-02-04.
  48. ↑ Lawton, Graham. "Military now controls Myanmar's scientifically important amber mines". New Scientist. Retrieved 2020-02-04.
  49. ↑ Joel, Lucas (2020-03-11). "Some Paleontologists Seek Halt to Myanmar Amber Fossil Research". The New York Times. ISSN   0362-4331 . Retrieved 2020-03-13.
  50. ↑ Rayfield, Emily J.; Theodor, Jessica M.; Polly, P. David (2020). Fossils from conflict zones and reproducibility of fossil-based scientific data. Society for Vertebrate Paleontology. pp. 1–2. Archived from the original on 2021-01-26. Retrieved 2020-04-28.
  51. ↑ "News - Acta Palaeontologica Polonica". www.app.pan.pl. Retrieved 2020-05-19.
  52. ↑ Barrett, Paul M.; Johanson, Zerina (2020-05-13). "Editorial: Myanmar (Burmese) Amber Statement". Journal of Systematic Palaeontology: 1. doi: 10.1080/14772019.2020.1764313 . ISSN   1477-2019.
  53. ↑ Szwedo, Jacek; Wang, Bo; Soszyńska-Maj, Agnieszka; Azar, Dany; Ross, Andrew J. (2020-06-30). "International Palaeoentomological Society Statement". Palaeoentomology. 3 (3): 221–222. doi: 10.11646/palaeoentomology.3.3.1 . ISSN   2624-2834.
  54. ↑ Haug, Joachim T.; Azar, Dany; et al. (September 2020). "Comment on the letter of the Society of Vertebrate Paleontology (SVP) dated April 21, 2020 regarding "Fossils from conflict zones and reproducibility of fossil-based scientific data": Myanmar amber". PalZ. 94 (3): 431–437. Bibcode:2020PalZ...94..431H. doi: 10.1007/s12542-020-00524-9 . hdl: 2445/175747 . ISSN   0031-0220.
  55. ↑ Poinar, George; Ellenberger, Sieghard (April 2020). "Burmese Amber Fossils, Mining, Sales and Profits". Geoconservation Research. 3 (1). doi:10.30486/gcr.2020.1900981.1018.
  56. ↑ Zherikhin, V. V.; Ross, A. J. (2000). "A review of the history, geology and age of Burmese amber burmite". Bulletin of the Natural History Museum, Geology Series.
  57. ↑ Zheng, D.; Chang, S.-C.; Perrichot, V.; Dutta, S.; Rudra, A.; Mu, L.; Kelly, R. S.; Li, S.; Zhang, Q.; Zhang, Q.; Wong, J. (December 2018). "A Late Cretaceous amber biota from central Myanmar". Nature Communications. 9 (1): 3170. Bibcode:2018NatCo...9.3170Z. doi:10.1038/s41467-018-05650-2. ISSN   2041-1723. PMC   6085374 . PMID   30093646.
  58. ↑ Čerňanský, Andrej; Stanley, Edward L.; Daza, Juan D.; Bolet, Arnau; Arias, J. Salvador; Bauer, Aaron M.; Vidal-García, Marta; Bevitt, Joseph J.; Peretti, Adolf M.; Aung, Nyi Nyi; Evans, Susan E. (2022-01-31). "A new Early Cretaceous lizard in Myanmar amber with exceptionally preserved integument". Scientific Reports. 12 (1): 1660. Bibcode:2022NatSR..12.1660C. doi:10.1038/s41598-022-05735-5. ISSN   2045-2322. PMC   8803969 . PMID   35102237.
This page is based on this Wikipedia article
Text is available under the CC BY-SA 4.0 license; additional terms may apply.
Images, videos and audio are available under their respective licenses.