Helotidae

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Helotidae
Temporal range: Aptian–Recent
Helotidae Munsiyari.jpg
Neohelota sp.
Scientific classification Red Pencil Icon.png
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Order: Coleoptera
Suborder: Polyphaga
Infraorder: Cucujiformia
Superfamily: Cucujoidea
Family: Helotidae
Chapuis, 1876
Genera

Helotidae is a family of beetles, in the suborder Polyphaga. The family includes about five extant genera, Helota MacLeay, Neohelota Ohta, Afrohelotina Kirejtshuk, Metahelotella Kirejtshuk, and Strophohelota Kirejtshuk. [1] [2] Helotidae are found mainly in the Old World tropics and are absent from Australia and Madagascar. The antennae are clubbed on the final three segments and is retractable within grooves under the head. The wings have reduced venation with just 4 anal veins. [3] Helotids are known to be associated with sap, fruit and flowers, and the larvae of some species are known to bore into wood in order to pupate. [4]

A few fossil genera have been described including Palaeohelota from the Lower Cretaceous (Aptian) Yixian Formation in China, Burmahelota,Metahelotella and Trihelota from the Upper Cretaceous (Cenomanian) Burmese amber of Northern Myanmar. Another fossil genus Laodiscis placed in this family originally is now considered of uncertain taxonomic position. [5]

Related Research Articles

<span class="mw-page-title-main">Cucujidae</span> Family of beetles

The Cucujidae, "flat bark beetles," are a family of distinctively flat beetles found worldwide under the bark of dead trees. The family has received considerable taxonomic attention in recent years and now consists of 70 species distributed in five genera. It was indicated Cucujus species are scavengers, only feeding on pupae and larvae of other insects and on other subcortical beetles such as their own. Since the Cucujidae prey on larvae of potentially tree damaging beetles that spread fungal diseases, they are considered to be beneficial to the health of living trees.

<span class="mw-page-title-main">Myxophaga</span> Suborder of beetles

Myxophaga is the second-smallest suborder of the Coleoptera after Archostemata, consisting of roughly 65 species of small to minute beetles in four families. The members of this suborder are aquatic and semiaquatic, and feed on algae.

<span class="mw-page-title-main">Cucujoidea</span> Superfamily of beetles

Cucujoidea is a superfamily of beetles. This group formerly included all of the families now included in the superfamily Coccinelloidea. They include some fungus beetles and a diversity of lineages of "bark beetles" unrelated to the "true" bark beetles (Scolytinae), which are weevils.

<span class="mw-page-title-main">Cerophytidae</span> Family of beetles

The Cerophytidae are a family of Coleoptera known as the rare click beetles. Adults are infrequently encountered, and larvae are found within wood, where they are believed to feed on fungi. The family contains over 20 species in five genera, primarily distributed in the New World: 17 fossil genera are known extending to the Early Jurassic.

<i>Lepicerus</i> Genus of beetles

Lepicerus is a genus of myxophagan beetles containing three described species in the family Lepiceridae; it is the only extant genus in the family, with another genus, Lepiceratus only known from fossils. Extant species occur in the Neotropics, from Mexico south to Venezuela and Ecuador. Fossils referrable to the genus are known from the early Late Cretaceous of Southeast Asia.

<span class="mw-page-title-main">Endomychidae</span> Family of beetles

Endomychidae, or handsome fungus beetles, is a family of beetles with representatives found in all biogeographic realms. There are around 120 genera and 1300 species. The family was established based on the type genus Endomychus, a genus erected in 1795 by Panzer which was applied to a species that Linnaeus called Chrysomela coccinea. As the common name suggests, Endomychidae feed on fungi. Crowson, in his influential treatment of the beetles, placed the family within the Cucujoidea. They have a tarsal formal of 4-4-4 or 3-3-3 and the wings lack a closed radial cell. The second antennal segment has a sensory appendage that is as long as the third antennal segment. The family has also been grouped with the Coccinellidae in a group called the Trimera for having pseudotrimerous tarsi. A 2015 molecular phylogeny study found that the Cucujoidea were found to be non-monophyletic and the Endomychidae was refined with the removal of the Anamorphinae from within the family and elevated to the status of a full family, Anamorphidae. Mycetaeinae and Eupsilobiinae were also found not to belong within the clades of the core Endomychidae, and likewise reclassified into the families Mycetaeidae and Eupsilobiidae.

<span class="mw-page-title-main">Ommatidae</span> Family of beetles

The Ommatidae are a family of beetles in the suborder Archostemata. The Ommatidae are considered the extant beetle family that has most ancestral characteristics. There are only seven extant species, confined to Australia and South America. However, the geographical distribution was much wider during the Mesozoic spanning across Eurasia and Australia, suggesting that they were widespread on Pangea. So far, over 26 extinct genera containing over 170 species of these beetles have been described. Three extant genera have been assigned to this family: Omma,Tetraphalerus and Beutelius. The family is considered to be a subfamily of Cupedidae by some authors, but have been found to be more closely related to Micromalthidae in molecular phylogenies. A close relationship with Micromalthidae is supported by several morphological characters, including those of the mandibles and male genitalia. Due to their rarity, their ecology is obscure, it is likely that their larvae feed on deadwood.

Boganiidae is a family of beetles, in the suborder Polyphaga. Members of the family are found in southern Africa, Australia and New Caledonia. Adults and larvae are pollenivorous, feeding on the pollen of cycads and Myrtaceae, Meliaceae, Cunoniaceae and Elaeocarpaceae angiosperms. Metacucujus and Paracucujus act as pollinators for cycads Encephalartos and Macrozamia respectively. This association with cycads goes back to at least the Mid-Cretaceous, with an extinct form being found with preserved cycad pollen in 99 million year old Burmese amber.

<span class="mw-page-title-main">Silvanidae</span> Family of beetles

Silvanidae, "silvan flat bark beetles", is a family of beetles in the superfamily Cucujoidea, consisting of 68 described genera and about 500 described species. The family is represented on all continents except Antarctica, and is most diverse at both the generic and species levels in the Old World tropics.

<span class="mw-page-title-main">Sphindidae</span> Family of beetles

Sphindidae is a family of beetles, in the suborder Polyphaga. They are also known as slime mold beetles due to their exclusive feeding on slime molds during adult and larval stages, other aspects of their life history are obscure. Palaeontological discoveries since 2015 have added to the geologic history of Sphindidae, including the discovery of Libanopsis, placed in the extinct subfamily Libanopsinae.

<span class="mw-page-title-main">Phloeostichidae</span> Family of beetles

Phloeostichidae is a family of beetles in the superfamily Cucujoidea. There are at least three genera in Phloeostichidae.

The Brachypsectridae are a family of beetles commonly known as the Texas beetles. There are only two extant genera, Brachypsectra and Asiopsectra. The type species, Brachypsectra fulva, occurs in North America. Other species in the genus have a cosmopolitan distribution. Two other extant and fossil species have been described from the Dominican Republic. While formerly considered monogeneric, in 2016 Asiopsectra was described from specimens found in Iran and Tajikistan. Two extinct genera, Vetubrachypsectra and Hongipsectra, known from adults, and a larval genus, Cretopsectra are known from Cenomanian aged Burmese amber.

Brochocoleus is an extinct genus of beetles in the family Ommatidae, known from the Early Jurassic to the Early Late Cretaceous. 9 species are currently recognised, with many species being reassigned to other genera by Kirejtshuk's major systematic revision in 2020.

<span class="mw-page-title-main">Monotomidae</span> Family of beetles

Monotomidae is a family of beetles in the superfamily Cucujoidea. The family is found worldwide, with approximately 240 species in 33 genera. The ecological habits of the family are diverse, with different members of the group being found under tree bark, in decaying vegetation, on flowers and in ant nests. Their ecology is obscure, while at least some species are mycophagous, feeding on the fruiting bodies of ascomycete fungi, Rhyzophagus are predators on bark beetles and possibly Phoridae larvae, with the larvae of some species also being mycophagous.

<span class="mw-page-title-main">Passandridae</span> Family of beetles

Passandridae, the "parasitic flat bark beetles," are a family of beetles notable for being one of the very few beetle families with larvae that are, as far as known, exclusively ectoparasitic on the immature stages of other beetles and Hymenoptera.

<span class="mw-page-title-main">Artematopodidae</span> Family of beetles

Artematopodidae is a family of soft-bodied plant beetles in the superfamily Elateroidea. They are mostly found in understory forest foliage. The life history of the group is obscure, larvae of the genera Eurypogon and Macropogon likely feed on moss, while the larvae of Artematopus have been fed insect remains.The oldest fossils of the family date to the Middle Jurassic.

2017 in paleoentomology is a list of new fossil insect taxa that were described during the year 2017, as well as other significant discoveries and events related to paleoentomology that were scheduled to occur during the year.

2015 in paleoentomology is a list of new fossil insect taxa that were described during the year 2015, as well as other significant discoveries and events related to paleoentomology that were scheduled to occur during the year.

<i>Burmocoleus</i> Extinct genus of beetles

Burmocoelus is an extinct genus of Archostematan beetle belonging to the family Ommatidae. It is known from two species, Burmocoleus zhiyuani, which was initially assigned to the genus Brochocoleus this was placed into the new genus Burmocoleus by Kirejtshuk, 2020, who also described a new species, Burmocoleus prisnyi. Both species are known from the Cenomanian aged Burmese amber from Myanmar. Similar to Stegocoleus, Brochocoleus and Jarzembowskiops it has flat, wide elytral edges, but can be distinguished from these genera in several characters, and is more similar to other ommatids in the characters of the prothorax.

Burmese amber is fossil resin dating to the early Late Cretaceous Cenomanian age recovered from deposits in the Hukawng Valley of northern Myanmar. It is known for being one of the most diverse Cretaceous age amber paleobiotas, containing rich arthropod fossils, along with uncommon vertebrate fossils and even rare marine inclusions. A mostly complete list of all taxa described up until 2018 can be found in Ross 2018; its supplement Ross 2019b covers most of 2019.

References

  1. Lee, Chi-Feng; Votruba, Petr (2013). "A revision of the family Helotidae (Coleoptera: Cucujoidea): VIII. The guerinii species-group of the genus Neohelota". Entomologica Basiliensia et Collectionis Frey. 34: 269–308.
  2. Lee, Chi-Feng (2007). "Revision of Family Helotidae (Coleoptera: Cucujoidea):I. Gemmata Group of Genus Helota". Ann. Entomol. Soc. Am. 100 (5): 623–639.
  3. Crowson, R.A. (1967). The natural classification of the families of Coleoptera. Middlesex: E.W. Classey Ltd. p. 104.
  4. Lawrence, John F.; Ślipinśki, Adam; Lee, Chi-Feng (2010-12-31), Kükenthal, Willy; Leschen, Richard A.B.; Beutel, Rolf G.; Lawrence, John F. (eds.), "10.3. Helotidae Reitter, 1876/Chapuis, 1876", Coleoptera, Beetles, Volume 2, Morphology and Systematics (Elateroidea, Bostrichiformia, Cucujiformia partim), DE GRUYTER, pp. 292–295, doi:10.1515/9783110911213.292, ISBN   978-3-11-019075-5 , retrieved 2022-10-02
  5. Liu, Zhenhua; Ślipiński, Adam; Ren, Dong; Pang, Hong (2019). "The first Mesozoic Helotidae (Coleoptera: Cucujoidea)". Cretaceous Research. 96: 113–119. doi:10.1016/j.cretres.2018.12.005.

Additional sources