Eucnemidae

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Eucnemidae
Temporal range: Tithonian–Recent
Dirrhagofarsus lewisi (10.3897-zookeys.781.22335) Figure 1 (cropped).jpg
Dirrhagofarsus lewisi
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Order: Coleoptera
Suborder: Polyphaga
Infraorder: Elateriformia
Superfamily: Elateroidea
Family: Eucnemidae
Eschscholtz, 1829
Subfamilies
Synonyms
  • Anischiidae Fleutiaux, 1936
  • Phylloceridae Reitter, 1905
  • Melasidae Leach, 1817
  • Perothopidae Lacordaire, 1857

Eucnemidae, or false click beetles, are a family of elateroid beetles based on the type genus Eucnemis ; they include about 1700 species, distributed worldwide.

Contents

Description

Closely related to the family Elateridae, specimens of Eucnemidae can reach a length of 2–30 mm (0.079–1.181 in). Bodies are slightly flattened and convex. The upper surfaces of the body usually has hairs, setae or scales.

Ecology

The larvae are typically legless, and generally develop feeding on the fluids of rotting wood, likely vomiting digestive enzymes into the wood to break apart the fungal hyphae, moving using their shovel shaped heads to force apart the wood. Adults, which are typically found on broken surfaces of trunks and stumps, have a short lifespan and it is unclear whether they feed, though they are capable fliers, and like some other elateroids are capable of clicking. [1]

Taxonomy

Extant Genera

The Global Biodiversity Information Facility [2] includes:

  1. Acedax de Bonvouloir, 1875
  2. Achaica Muona, 1991
  3. Adelorhagus Horn, 1890
  4. Adelothyreus Chevrolat, 1868
  5. Adelothyreus Horn, 1890
  6. Agalba Broun, 1893
  7. Agastocerus de Bonvouloir, 1871
  8. Anabolodes Strand, 1928
  9. Anelastes Kirby, 1818
  10. Anelastidius Jacquelin du Val, 1860
  11. Anischia Fleutiaux, 1896
  12. Arganus de Bonvouloir, 1871
  13. Arhipis Bonvouloir, 1871
  14. Arisus de Bonvouloir, 1871
  15. Arrhipis Bonvouloir, 1871
  16. Aruanus Fleutiaux, 1902
  17. Asiocnemis Mamaev, 1976
  18. Aubailius Fleutiaux, 1921
  19. Balgus Fleutiaux, 1920
  20. Balistica Motschulsky, 1861
  21. Baryus Fleutiaux, 1921
  22. Basilewskydia Cobos, 1959
  23. Bermillus de Bonvouloir, 1871
  24. Bioxylus Fleutiaux, 1923
  25. Bonvouloiriella Otto & Kovalev, 2013
  26. Bossionus Fleutiaux, 1922
  27. Bothrion Fleutiaux, 1921
  28. Buckia Cobos, 1956
  29. Buckiella Cobos, 1964
  30. Cafolus de Bonvouloir, 1875
  31. Calyptocerus Guérin, 1843
  32. Ceballosmelasis Cobos, 1965
  33. Ceratogonys Perty, 1830
  34. Ceratus Bonvouloir, 1871
  35. Chapianus Fleutiaux, 1921
  36. Chropoecilus Fleutiaux, 1902
  37. Cladidus Fleutiaux, 1918
  38. Cladoscython Fleutiaux, 1921
  39. Cladus Bonvouloir, 1871
  40. Clyeorhagus Olexa, 1975
  41. Clypeorhagus Olexa, 1975
  42. Cobosiantha Alonso-Zarazaga, 1998
  43. Cobosidea Chassain, 2015
  44. Compsocnemis de Bonvouloir, 1875
  45. Coomanius Fleutiaux, 1924
  46. Cretopityobius Otto, 2019
  47. Cryptostoma (beetle) Latreille, 1829
  48. Cupressicharis Muona, 2020
  49. Curtocephalus Fleutiaux, 1912
  50. Cylus Muona, 2020
  51. Cyrtostethus de Bonvouloir, 1872
  52. Deltometopus Bonvouloir, 1871
  53. Dendrocharis Guérin-Méneville, 1843
  54. Diacerus de Bonvouloir, 1871
  55. Diapodius de Bonvouloir, 1875
  56. Dictyeucnemis Lea, 1919
  57. Diphytaxis Horn, 1890
  58. Dirrhagofarsus Fleutiaux, 1935
  59. Discaptothorax Blackburn, 1900
  60. Dorsifornax Fleutiaux, 1926
  61. Dromaeocnemis de Bonvouloir, 1872
  62. Dromaeoloides Cobos, 1964
  63. Dromaeolus Kiesenwetter, 1858
  64. Dyscharachthis Blackburn, 1900
  65. Dyscolotaxia Horn, 1890
  66. Echthrogaster Blackburn, 1900
  67. Entomophthalmus Bonvouloir, 1871
  68. Entomosatopus de Bonvouloir, 1871
  69. Epiphanis Eschscholtz, 1829
  70. Epipleurus Fleutiaux, 1928
  71. Erdaia Muona, 1993
  72. Eucalodemas Bonvouloir, 1871
  73. Eucalosoma Laporte, 1840
  74. Eucnemis Ahrens, 1812
  75. Eudorus (beetle) Laporte, 1835
  76. Eurachis Horn, 1890
  77. Euryaulacus Bonvouloir, 1871
  78. Euryostus de Bonvouloir, 1871
  79. Euryptychoides Muona, 2021
  80. Euryptychus Leconte, 1852
  81. Falsothambus Muona, 2020
  82. Farsus Jacquelin du Val, 1860
  83. Feaia Fleutiaux, 1896
  84. Fiegelia Muona, 2020
  85. Fleutimelasa Otto & Kovalev, 2013
  86. Fornax Laporte, 1835
  87. Fornaxiella Cobos, 1965
  88. Freudenia Cobos, 1965
  89. Freyiola Cobos, 1965
  90. Fryanus Fleutiaux, 1902
  91. Gagatellus Fleutiaux, 1912
  92. Galbimorpha Fleutiaux, 1920
  93. Galbites Fleutiaux, 1918
  94. Galbodema Laporte, 1835
  95. Gastraulacus Guérin, 1843
  96. Golbachia Cobos, 1955
  97. Goudotus Fleutiaux, 1926
  98. Haywardinia Cobos, 1965
  99. Haywardius Cobos, 1958
  100. Hemiopsida MacLeay, 1873
  101. Henecocrus Bonvouloir, 1871
  102. Henecosoma Horn, 1890
  103. Heterotaxis de Bonvouloir, 1875
  104. Hodocerus de Bonvouloir, 1872
  105. Hylis Gozis, 1886
  106. Hylochares Latreille, 1834
  107. Hylotastella Fisher, 1945
  108. Hylotastes de Bonvouloir, 1875
  109. Hyperpalpus Lucht, 1990
  110. Hypocoelus Latreille, 1834
  111. Hyporrhagus Reitter, 1921
  112. Idiotarsus Bonvouloir, 1871
  113. Insulanus Fleutiaux, 1930
  114. Isarthrus LeConte, 1852
  115. Isoriphis Boisduval & Lacordaire, 1835
  116. Jenibuntor Muona, 1993
  117. Lacus (beetle) Bonvouloir, 1871
  118. Lamprotrichus Bonvouloir, 1871
  119. Langurioscython Heller, 1917
  120. Macraulacus de Bonvouloir, 1872
  121. Macroscython Fleutiaux, 1902
  122. Maelodrus Fleutiaux, 1928
  123. Mahatsinjus Fleutiaux, 1926
  124. Melanocoleus Bonvouloir, 1872
  125. Melanoscython Fleutiaux, 1926
  126. Melasis Olivier, 1790
  127. Melocarvalhosia Cobos, 1957
  128. Mesogenus Bonvouloir, 1871
  129. Microrhagus Dejean, 1833
  130. Microtrigonus Bonvouloir, 1871
  131. Modius (beetle) Fleutiaux, 1918
  132. Monrosina Cobos, 1958
  133. Muonaja Özdikmen, 2008
  134. Myall Muona, 1993
  135. Namolius Bonvouloir, 1875
  136. Nebulatorpidus Lucht, 1998
  137. Nematodes (beetle) Berthold, 1827
  138. Neocharis Sharp, 1877
  139. Neocharoides Muona, 2021
  140. Neodema Fleutiaux, 1921
  141. Neodiaeretus Cobos, 1956
  142. Neodiapodius Fisher, 1945
  143. Neofornax Cobos, 1958
  144. Neomathion Fleutiaux, 1930
  145. Neusiokia Muona, 2022
  146. Oisocerus Bonvouloir, 1868
  147. Olbrechtsia Cobos, 1959
  148. Onichodon Newman, 1838
  149. Onya Muona, 1993
  150. Orodotes Bonvouloir, 1875
  151. Otho (beetle) Lacordaire, 1857
  152. Pachyfornax Fleutiaux, 1902
  153. Palaeoxenus Horn, 1891
  154. Paleoeucnemis Muona, 2020
  155. Paradema Cobos, 1986
  156. Paraxylophilus Cobos, 1965
  157. Pereirella Cobos, 1956
  158. Perothops Eschscholtz, 1836
  159. Perrotius Fleutiaux, 1938
  160. Phaenobolus Horn, 1890
  161. Phaenocerus Bonvouloir, 1871
  162. Phanerochroeus Bonvouloir, 1871
  163. Phlegon (beetle) Laporte de Castelnau, 1840
  164. Phyllocerus Lepeletier & Serville, 1828
  165. Piestocera Perty, 1830
  166. Pinaroo Muona, 1993
  167. Plesiofornax Bonvouloir, 1866
  168. Plesiofornax Coquerel, 1866
  169. Pocoelus Fleutiaux, 1926
  170. Poecilochrus Bonvouloir, 1871
  171. Porraulacus Fleutiaux, 1896
  172. Potergites Britton, 1960
  173. Procladidus Fleutiaux, 1921
  174. Prodirhagus Fleutiaux, 1925
  175. Profornax Fleutiaux, 1896
  176. Prolycaon Fleutiaux, 1923
  177. Prosassanus Fleutiaux, 1945
  178. Protofarsus Muona, 2000
  179. Protomicrorhagus Muona, 2020
  180. Protovitellius Muona, 2020
  181. Proutianus Muona, 1993
  182. Proxylobius Fleutiaux, 1900
  183. Pseudochapianus Lucht, 1987
  184. Pseudodiaeretus Fleutiaux, 1902
  185. Pseudomenes Fleutiaux, 1902
  186. Pseudomyall Muona, 2020
  187. Pseudorhacopus Olexa, 1975
  188. Pseudoscython Fleutiaux, 1902
  189. Pseudotemnillus Muona, 2021
  190. Quirsfeldia Cobos, 1965
  191. Raapia Fleutiaux, 1899
  192. Rhacopus Hampe, 1855
  193. Rhagomicrus Fleutiaux, 1902
  194. Rheanischia Li, Chang & Muona, 2021
  195. Sakalavus Fleutiaux, 1926
  196. Saproxylobius Leiler, 1990
  197. Sarfus Fleutiaux, 1925
  198. Sarpedon (beetle) Bonvouloir, 1871
  199. Sassanus Fleutiaux, 1925
  200. Schizophilus Bonvouloir, 1871
  201. Scopulifer Fleutiaux, 1896
  202. Scython Laporte, 1835
  203. Semnodema Bonvouloir, 1875
  204. Serrifornax Fleutiaux, 1926
  205. Sieglindea Muona, 1993
  206. Silveriola Cobos, 1956
  207. Similifornax Fleutiaux, 1926
  208. Somahenecus Cobos, 1965
  209. Spinifornax Fleutiaux, 1926
  210. Stethon LeConte, 1866
  211. Submesogenus Fleutiaux, 1921
  212. Tacerus Fleutiaux, 1928
  213. Tachycnemis Bonvouloir, 1872
  214. Talerax Sharp, 1877
  215. Temnillus Bonvouloir, 1871
  216. Temnus Fleutiaux, 1920
  217. Thambus Bonvouloir, 1871
  218. Torigaia Hisamatsu & Sato, 1959
  219. Trigonopleurus Bonvouloir, 1875
  220. Vitellius (beetle) Bonvouloir, 1871
  221. Weyrauchiella Cobos, 1972
  222. Xylophilus (beetle) Mannerheim, 1823
  223. Xylotho Lucht, 1990

Note misplaced names: AulacosternusMotschoulsky, 1861

Fossil genera

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<span class="mw-page-title-main">Whirligig beetle</span> Family of beetles

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<span class="mw-page-title-main">Elateroidea</span> Superfamily of beetles

The Elateroidea are a large superfamily of beetles. It contains the familiar click beetles, fireflies, and soldier beetles and their relatives. It consists of about 25,000 species.

<span class="mw-page-title-main">Buprestinae</span> Subfamily of beetles

Buprestinae is a subfamily of beetles in the family Buprestidae, containing the following genera in the tribes Anthaxiini, Buprestini, Chrysobothrini, Melanophilini, and Xenorhipidini:

<span class="mw-page-title-main">Chrysochroinae</span> Subfamily of beetles

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Physodeutera is a genus of Tiger Beetles endemic to Madagascar. There are more than 60 described species in Physodeutera.

<i>Pogonostoma</i> Genus of beetles

Pogonostoma is a genus of Tiger Beetles endemic to Madagascar.

<span class="mw-page-title-main">Aleocharinae</span> Subfamily of beetles

The Aleocharinae are one of the largest subfamilies of rove beetles, containing over 12,000 species. Previously subject to large-scale debate whether the subfamily deserved the familial status, it is now considered one of the largest subfamilies of rove beetles.

<span class="mw-page-title-main">Agrilini</span> Tribe of beetles

Agrilini is a tribe of metallic wood-boring beetles in the family Buprestidae. There are at least 40 described genera in Agrilini.

<span class="mw-page-title-main">Ptilodactylidae</span> Family of beetles

Ptilodactylidae is a family of beetles belonging to the Elateriformia. There around 500 extant species in 35 genera. They are generally associated with riparian and aquatic habitats. The larvae generally live associated with rotting wood or vegetation, or within gravel and detritus on the edge of water bodies. The larvae of some species feed on submerged rotting wood or on plant roots, while the adults of some species are known to feed on fungus with modified brush-like maxillae.

<span class="mw-page-title-main">Throscidae</span> Family of beetles

Throscidae is a family of elateroid beetles found worldwide with around 150 species in 5 extant genera. The larvae are soil-dwelling, siphoning fluid from mycorrhizae attached to trees. The adults are short-lived, with the adult males being noted for a complex mating dance. Like some other elateroids, they are capable of clicking.

<span class="mw-page-title-main">Melasinae</span> Subfamily of beetles

Melasinae is a subfamily of false click beetles in the family Eucnemidae.

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Agrypninae is a subfamily of click beetles in the family Elateridae. There are at least 130 genera and more than 430 described species in Agrypninae.

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<i>Melasis</i> Genus of beetles

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<span class="mw-page-title-main">Eucneminae</span> Subfamily of beetles

The Eucneminae are a subfamily of "false click beetle" genera, erected by Johann von Eschscholtz in 1889.

Mesogenus is a genus of mostly Asian click beetle allies in the subfamily Eucneminae and typical of the tribe Mesogenini, erected by Viscount Henri de Bonvouloir in 1871.

References

  1. Muona, Jyrki. "4.5. Eucnemidae Eschscholtz, 1829". Volume 2 Morphology and Systematics (Elateroidea, Bostrichiformia, Cucujiformia partim), edited by Willy Kükenthal, Richard A.B. Leschen, Rolf G. Beutel and John F. Lawrence, Berlin, New York: De Gruyter, 2011, pp. 61–69
  2. Global Biodiversity Information Facility: Eucnemidae (retrieved 2 January 2025)
  3. Li, Yan-Da; Tihelka, Erik; Liu, Zhen-Hua; Huang, Diying; Cai, Chen-Yang (2020-08-31). "Muonabuntor gen. nov., a new genus of false click beetles from mid-Cretaceous Burmese amber (Coleoptera: Elateroidea: Eucnemidae)". Palaeoentomology. 3 (4): 399–406. doi:10.11646/palaeoentomology.3.4.12. ISSN   2624-2834.