Endomychidae Temporal range: | |
---|---|
Brachytrycherus humeralis | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Class: | Insecta |
Order: | Coleoptera |
Suborder: | Polyphaga |
Infraorder: | Cucujiformia |
Superfamily: | Coccinelloidea |
Family: | Endomychidae Leach, 1815 |
Type genus | |
Endomychus | |
Synonyms | |
Merophysiidae Seidlitz, 1872 |
Endomychidae, or handsome fungus beetles, is a family of beetles with representatives found in all biogeographic realms. There are around 120 genera and 1300 species. The family was established based on the type genus Endomychus , a genus erected in 1795 by Panzer which was applied to a species that Linnaeus called Chrysomela coccinea. As the common name suggests, Endomychidae feed on fungi. Crowson, in his influential treatment of the beetles, placed the family within the Cucujoidea. They have a tarsal formal of 4-4-4 or 3-3-3 and the wings lack a closed radial cell. The second antennal segment has a sensory appendage that is as long as the third antennal segment. The family has also been grouped with the Coccinellidae in a group called the Trimera for having pseudotrimerous tarsi. [1] A 2015 molecular phylogeny study found that the Cucujoidea were found to be non-monophyletic and the Endomychidae was refined with the removal of the Anamorphinae from within the family and elevated to the status of a full family, Anamorphidae. Mycetaeinae and Eupsilobiinae were also found not to belong within the clades of the core Endomychidae, and likewise reclassified into the families Mycetaeidae and Eupsilobiidae. [2]
The subfamilies that are included:
These 70 genera belong to the family Endomychidae as defined in the past (due to revisions, this may not be entirely accurate):
Data sources: i = ITIS, [3] c = Catalogue of Life, [4] g = GBIF, [5] b = Bugguide.net [6]
Cucujoidea is a superfamily of beetles. This group formerly included all of the families now included in the superfamily Coccinelloidea. They include some fungus beetles and a diversity of lineages of "bark beetles" unrelated to the "true" bark beetles (Scolytinae), which are weevils.
Ripiphoridae is a cosmopolitan family of some 450 described species of beetles sometimes called "wedge-shaped beetles". Ripiphoridae are unusual among beetle families in that many species are hypermetamorphic parasitoids, an attribute that they share with the Meloidae. Members of the family differ in their choice of hosts, but most attack various species of bees or wasps, while some others attack cockroaches or beetles. Many species of Ripiphoridae have abbreviated elytra, and flabellate or pectinate antennae.
Corylophidae is a family of minute hooded beetles, sometimes called minute fungus beetles, in the superfamily Coccinelloidea. There are about 18 genera and at least 120 described species in Corylophidae. They feed on microfungi such as molds, and are often found associated with bark, as well as in leaf litter and other decaying vegetation. In older literature, the family name was often given as Orthoperidae.
Erotylidae, or the pleasing fungus beetles, is a family of beetles belonging to Cucujoidea containing over 100 genera. In the present circumscription, it contains 6 tribes and 10 subfamilies. In other words, the narrowly circumscribed Erotylidae correspond to the subfamily Erotylinae in the definition sensu lato. There are doubts on the monophyly of lower ranked taxa within Erotylidae, with further phylogenetic studies requiring better sampling and studies of unexplored character sets, for example the metendosternite and penile flagellum, which are generally lacking detailed morphological studies within the Coleoptera literature. The Eroytlina taxonomy is based on traits such as their different colors and not off morphological differences like mouthparts, thorax, and abdominal terminalia (Pecci-Maddalena).
Bothrideridae is a family of beetles in the superfamily Coccinelloidea. They are known commonly as the cocoon-forming beetles or dry bark beetles. They occur worldwide with most native to the Old World tropics. In older literature, the family was often included in the family Colydiidae, but is now considered unrelated.
Cerylonidae are small to tiny, smooth, shiny, hairless beetles, only lightly punctured. There are about 450 species worldwide in 50 or so genera, mostly tropical and subtropical. They are most common under the bark of dead trees, but can also occur in compost and other decaying plant material. Little is known specifically about their biology but they are thought to be either predators that feed on other small animals, or fungus-feeders.
Eumorphus is a genus of beetles belonging to the family Endomychidae.
Coccinelloidea is a superfamily of beetles in the order Coleoptera, formerly included in the superfamily Cucujoidea. There are more than 10,000 species in Coccinelloidea, including more than 6000 in the lady beetle family Coccinellidae.
Danae is a genus of handsome fungus beetles in the family Endomychidae. There are more than 90 species in Danae, 1 found in North America, about 10 in Asia, and the remainder in Africa.
Teredidae is a family of beetles in the superfamily Coccinelloidea, formerly included within the family Bothrideridae. There are around 160 species in 10 genera, found worldwide except South America. Teredids are generally found under bark, in the galleries of wood-boring beetles, or in leaf litter. They are thought to be fungivores. The oldest records of the family are Delteredolaemus from mid-Cretaceous aged Burmese amber from Myanmar and a species of Teredolaemus from Eocene aged Baltic amber.
Anamorphidae is a family of beetles in the superfamily Coccinelloidea, formerly included within the family Endomychidae. They are found worldwide. Like enchomyids, they are fungivores, with adult and larval stages thought to exclusively consume fungal spores.
Eupsilobiidae is a family of beetles in the superfamily Coccinelloidea, formerly included within the family Endomychidae. Most genera are restricted to the Neotropics, while the genus Eidoreus is found worldwide. They are fungivores, and have been observed living commensally in bee and ant nests.
Epipocus is a genus of handsome fungus beetles in the family Endomychidae. There are at least 20 described species in Epipocus.
Trochoideus is a genus of handsome fungus beetles in the family Endomychidae. There are about 10 described species in Trochoideus.
Bystus is a genus of beetles in the family Anamorphidae. There are about six described species in Bystus.
Epipocinae is a subfamily of handsome fungus beetles in the family Endomychidae. There are at least 4 genera and more than 40 described species in Epipocinae.
Stenotarsus is a genus of handsome fungus beetles in the family Endomychidae. There are more than 50 described species in Stenotarsus.
Leiestinae is a subfamily of handsome fungus beetles in the family Endomychidae. There are at least three genera and about five described species in Leiestinae.
Merophysiinae is a subfamily of handsome fungus beetles in the family Endomychidae.
Xenomycetes is a genus of handsome fungus beetles in the family Endomychidae. It is the only genus in the subfamily Xenomycetinae. There are at least two described species in Xenomycetes, both endemic to northwestern North America.