Goliathus | |
---|---|
Goliathus regius | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Class: | Insecta |
Order: | Coleoptera |
Family: | Scarabaeidae |
Subfamily: | Cetoniinae |
Tribe: | Goliathini |
Subtribe: | Goliathina |
Genus: | Goliathus Lamarck, 1801 |
The Goliath beetles (named after the biblical giant Goliath) are any of the six species in the genus Goliathus. Goliath beetles are among the largest insects on Earth, if measured in terms of size, bulk and weight. [1] [2] They are members of subfamily Cetoniinae, within the family Scarabaeidae. [1] Goliath beetles can be found in many of Africa's tropical forests, [1] where they feed primarily on tree sap and fruit. [1] [2] Little appears to be known of the larval cycle in the wild, but in captivity, Goliathus beetles have been successfully reared from egg to adult using protein-rich foods, such as commercial cat and dog food. Goliath beetles measure from 60–110 millimetres (2.4–4.3 in) for males and 50–80 millimetres (2.0–3.1 in) for females, as adults, and can reach weights of up to 80–100 grams (2.8–3.5 oz) in the larval stage, though the adults are only about half this weight. The females range from a dark chestnut brown to silky white, but the males are normally brown/white/black or black/white. [1] Goliath beetles, while not currently evaluated on the IUCN Red List, are facing growing conservation challenges across their African range due to habitat loss, over-collection for the international pet trade, and the potential impacts of climate change. [3]
There are six species of Goliath beetles, [4] with several different subspecies and forms only partially described:
Goliathus larvae are somewhat unusual among cetoniine scarabs in that they have a greater need for high-protein foods than do those of most other genera. [1] [2] Pellets of dry or soft dog or cat food (buried in the rearing substrate on a regular schedule) provide a suitable diet for Goliathus larvae in captivity. [2] However, a substrate of somewhat moistened, decayed leaves and wood should still be provided in order to create a suitable medium for larval growth. [2] The young stage larvae (1st instar) will eat some of this material. Even under optimum conditions, the larvae take a number of months to mature fully because of the great size they attain. They are capable of growing up to 250 millimetres (9.8 in) in length and reaching weights in excess of 100 grams (3.5 oz). [2]
When maximum size is reached, the larva constructs a rather thin-walled, hardened cell of sandy soil in which it will undergo pupation and metamorphose to the adult state. [2] Once building of this cocoon is completed, the larva transforms to the pupal stage, which is an intermediate phase between the larval and adult stages. [2] During the pupal duration, the insect's tissues are broken down and re-organized into the form of the adult beetle. Once metamorphosis is complete, the insect sheds its pupal skin and undergoes a period of hibernation as an adult beetle until the dry season ends. [2] When the rains begin, the beetle breaks open its cocoon, locates a mate, and the entire life cycle starts over again. The adult beetles feed on materials rich in sugar, especially tree sap and fruit. [1] Under captive conditions, adults can sometimes live for about a year after emerging from their pupal cells. [2] Longevity in the wild is likely to be shorter on average due to factors such as predators and weather. The adult phase concentrates solely on reproduction, and once this function is performed, the time of the adult beetle is limited, as is true for the vast majority of other insect species.
The bulky bodies of Goliath beetles are composed of a thick and hardened exoskeleton, which protects their organs and hindwings. Like most beetles, they possess reinforced forewings (called elytra) that act as protective covers for their hindwings and abdomen. Only the hindwings (which are large and membranous) are actually used for flying, while the elytra are kept completely closed; flying with closed elytra is universal among cetoniine scarabs but rare in other beetles. [10] When not in use, the wings are kept completely folded beneath the elytra. Each of the beetle's legs ends in a pair of sharp claws, which provide a strong grip used for climbing on tree trunks and branches.
Males have a Y-shaped horn on the head, which is used as a pry bar in battles with other males over feeding sites or mates. Females lack horns and instead have a wedge-shaped head that assists in burrowing when they lay eggs. In addition to their massive size, Goliath beetles are strikingly patterned; prominent markings common to all of the Goliathus species are the sharply contrasting black vertical stripes on the pronotum (thoracic shield), while the various species may be most reliably distinguished based on their distinctive mix of elytral colors and patterns. [1]
The family Scarabaeidae, as currently defined, consists of over 35,000 species of beetles worldwide; they are often called scarabs or scarab beetles. The classification of this family has undergone significant change. Several groups formerly treated as subfamilies have been elevated to family rank, and some reduced to lower ranks. The subfamilies listed in this article are in accordance with those in Catalog of Life (2023).
An elytron is a modified, hardened forewing of beetles (Coleoptera), though a few of the true bugs (Hemiptera) such as the family Schizopteridae are extremely similar; in true bugs, the forewings are called hemelytra, and in most species only the basal half is thickened while the apex is membranous, but when they are entirely thickened the condition is referred to as "coleopteroid". An elytron is sometimes also referred to as a shard.
The huhu beetle is a longhorn beetle endemic to New Zealand. It is the heaviest beetle found in New Zealand.
The Hercules beetle is a species of rhinoceros beetle native to the rainforests of southern Mexico, Central America, South America, and the Lesser Antilles. It is the longest extant species of beetle in the world, and is also one of the largest flying insects in the world.
Dynastes tityus, the eastern Hercules beetle, is a species of rhinoceros beetle native to the Eastern United States. The adult's elytra are green, gray or tan, with black markings, and the whole animal, including the male's horns, may reach 60 mm (2.4 in) in length. The larvae feed on decaying wood from various trees.
The Japanese rhinoceros beetle, also known as the Japanese rhino beetle, the Japanese horned beetle, or by its Japanese name kabutomushi, is a species of rhinoceros beetle. They are commonly found in continental Asia in countries such as China, the Korean peninsula, Japan, and Taiwan. In these areas, this species of beetle is often found in broad-leaved forests with tropical or sub-tropical climates. This beetle is well known for the prominent cephalic horn found on males. Male Japanese rhinoceros beetles will use this horn to fight other males for territory and access to female mating partners. Upon contact, males will attempt to flip each other onto their backs or off of their feeding tree. In response to selective pressures, smaller male A. dichotoma have adapted a "sneak-like behavior". These smaller beetles will attempt to avoid physical confrontation with larger males and try to mate with females.
Flower chafers are a group of scarab beetles comprising the subfamily Cetoniinae. Many species are diurnal and visit flowers for pollen and nectar, or to browse on the petals. Some species also feed on fruit. The group is also called fruit and flower chafers, flower beetles and flower scarabs. There are around 4,000 species, many of them still undescribed.
Cotinis nitida, commonly known as the green June beetle, June bug or June beetle, is a beetle of the family Scarabaeidae. It is found in the eastern United States and Canada, where it is most abundant in the South. It is sometimes confused with the related southwestern species figeater beetle Cotinis mutabilis, which is less destructive.
Goliathus cacicus, the chief goliath, is a species of beetles of the family Scarabaeidae.
Goliathus albosignatus is a species of beetle of the family Scarabaeidae, described by Carl Henrik Boheman in 1857. It is one of several species of Goliath beetles that inhabit Africa, but it is the only one exclusively found in subtropical sections of the continent.
Goliathus goliatus is a very large species of beetle of the family Scarabaeidae, native to tropical Africa.
Goliathus orientalis is a species of beetles belonging to the family Scarabaeidae.
Goliathus regius, the Royal Goliath beetle, is a species of beetles of the family Scarabaeidae.
Melolonthinae is a subfamily of the scarab beetles. It is a very diverse group; distributed over most of the world, it contains over 11,000 species in over 750 genera. Some authors include the scarab subfamilies Euchirinae and Pachypodinae as tribes in the Melolonthinae.
Dynastes satanas, the Satanas beetle, is a species of beetle belonging to the family Scarabaeidae. The name is sometimes misspelled as "satanus".
Stephanorrhina princeps, common name spotted flower beetle, is a beetle from the family Scarabaeidae, subfamily Cetoniinae and tribe Goliathini.
Mecynorhina torquata is a beetle from the subfamily Cetoniinae, tribe Goliathini.
Bolitotherus cornutus is a North American species of darkling beetle known as the horned fungus beetle or forked fungus beetle. All of its life stages are associated with the fruiting bodies of a wood-decaying shelf fungus, commonly Ganoderma applanatum, Ganoderma tsugae, and Ganoderma lucidum.
Chondrorrhina abbreviata is a species of fruit and flower chafers belonging to the family Scarabaeidae, subfamily Cetoniinae.
The white worm beetle is a species of beetle in the family Scarabaeidae. It is the only species in the genus Hylamorpha. This beetle is native to South America, particularly in regions of Argentina, Brazil, and Uruguay.