List of largest insects

Last updated

Graphic showing the largest living and extinct insects, from top to bottom, left column: actaeon beetle, Coscinocera hercules, Homoioptera gigantea, titan beetle and Gigatitan similis, central column: Mazothairos enormis, Eurycnema versirubra, Meganeuropsis permiana and Phryganistria sp., right column: Clatrotitan scullyi, Kalligramma haeckeli, Bojophlebia prokopi, Arachnacris tenuipes and Thysania agrippina Largest insects size comparison.png
Graphic showing the largest living and extinct insects, from top to bottom, left column: actaeon beetle, Coscinocera hercules , Homoioptera gigantea , titan beetle and Gigatitan similis, central column: Mazothairos enormis , Eurycnema versirubra , Meganeuropsis permiana and Phryganistria sp., right column: Clatrotitan scullyi , Kalligramma haeckeli , Bojophlebia prokopi , Arachnacris tenuipes and Thysania agrippina

Insects, which are a type of arthropod, are the most numerous group of multicellular organisms on the planet, with over a million species identified so far. [1] The title of heaviest insect in the world has many contenders, the most frequently crowned of which is the larval stage of the goliath beetle, Goliathus goliatus , the maximum size of which is at least 115 g (4.1 oz) and 11.5 cm (4.5 in). The highest confirmed weight of an adult insect is 71 g (2.5 oz) for a gravid female giant weta, Deinacrida heteracantha , [2] although it is likely one of the elephant beetles, Megasoma elephas and Megasoma actaeon , or goliath beetles, both of which can commonly exceed 50 g (1.8 oz) and 10 cm (3.9 in), can reach a greater weight. [2]

Contents

The longest insects are the stick insects, see below.

Representatives of the extinct dragonfly-like order Meganisoptera (also known as griffinflies) such as the Carboniferous Meganeura monyi and the Permian Meganeuropsis permiana are the largest insect species ever known. These creatures had a wingspan of some 71 cm (28 in). Their maximum body mass is uncertain, with estimates varying between 34 g [3] and 210 g. [4]

Dragonflies and damselflies (Odonata)

The largest wingspan of any living species of odonate (the order that includes dragonflies and damselflies) belongs to Megaloprepus caerulatus , reaching 19 centimeters (7.5 inches) in wingspan. Petalura ingentissima has the largest wingspan of any dragonfly at 165 mm (6.5 in), although Tetracanthagyna plagiata comes in with a close second of 163 mm (6.4 in).

The largest wing surface area of any living species of odonate belongs to Chlorogomphus papilio , a dragonfly.

The longest body length of any living species of odonate belongs to Mecistogaster lucretia with a body length of 155 mm (6.1 in).

The heaviest living species of odonate is probably either Petalura ingentissima or Tetracanthagyna plagiata . However, other species such as Anax strenuus and Anotogaster sieboldii and other species in Petalura , Tetracanthagyna , Anax and Anotogaster could also be contenders.

See also the extinct genera Meganeuropsis and Meganeura , although they are not certain to be included in the modern dragonfly order.

Mayflies (Ephemeroptera)

The largest mayflies are members of the genus Proboscidoplocia from Madagascar. These insects can reach a length of 5 cm (2.0 in).

Palaeodictyoptera (extinct)

The largest insect of the order Palaeodictyoptera, Mazothairos enormis from the Carboniferous period, reached a wingspan of about 56 centimeters (22 in). [5]

Grasshoppers, crickets, and relatives (Orthoptera)

Giant weta Knights.weta.750pix.jpg
Giant weta

Arachnacris katydids and Tropidacris grasshoppers reach up to 12–15 cm (4.7–5.9 in) in length and 23–27.5 cm (9.1–10.8 in) in wingspan, making them the largest by these measurements. [6] [7] The largest Saga and Pseudophyllus bush crickets are only a few centimeters smaller. [7]

The heaviest of this widespread, varied complex of insects is the Little Barrier Island giant weta, Deinacrida heteracantha , of New Zealand; one specimen weighed 71 g (2.5 oz) and measured nearly 10 cm (3.9 in), [2] giving it one of the largest insect weights ever known. These heavyweight insects can be over 9 cm (3.5 in) long. [8]

Titanopterans (Titanoptera) (extinct)

Related to modern orthopterans, Triassic insects of the extinct suborder of Titanoptera [9] surpassed them in size. The wingspan of Gigatitan vulgaris was as large as of 40 centimetres (16 in). [10] Clatrotitan andersoni also reached a huge size, having a forewing of 13.8 centimetres (5.4 in) long. [11]

Earwigs (Dermaptera)

The largest of the earwigs is the Saint Helena earwig (Labidura herculeana), which is up to 8.4 cm (3.3 in) in length. There are no recent records of this species and it is generally considered extinct. [12] The largest certainly living species is the Australian giant earwig ( Titanolabis colossea), which is about 5 cm (2.0 in) long. [7]

Scorpionflies (Mecoptera)

The largest scorpionfly, the common scorpionfly (Panorpa communis), can reach a body length of about 30 mm (1.2 in). [13] [ full citation needed ]

Stick insects (Phasmatodea)

Phobaeticus serratipes, one of the longest stick insects Giant Stick Insect (Phobaeticus serratipes) on Sylvain (8727651923).jpg
Phobaeticus serratipes , one of the longest stick insects
Distributions of stick insect species known to exceed 50 cm (20 in) in total length Distribution insects exceeding 50cm.png
Distributions of stick insect species known to exceed 50 cm (20 in) in total length

The longest known stick insects are also the longest known insects, notably species in the tribe Pharnaciini, but they are generally relatively lightweight because of their slender shape. Female of these stick insects usually reach considerably larger sizes than males of the same species. [14] The longest known is a currently undescribed species of Phryganistria , informally referred to as Phryganistria "chinensis" , that was discovered in China in 2014. [15] A wild collected female kept at the Insect Museum of West China in Chengdu was the record holder with a total length, including extended legs, of 62.4 cm (24.6 in), and a body length of 36.1 cm (14.2 in), [16] [17] but it was surpassed by one of its captive bred young that reached 64 cm (25 in) in total length. [18] [19]

Other very large species, formerly believed to be longest but now considered second longest is Phobaeticus chani ; a specimen held in the Natural History Museum in London is 56.7 cm (22.3 in) in total length, including extended legs, and it has a body length of 35.7 cm (14.1 in). [20] [21] [22] Females of the Australian Ctenomorpha gargantua have been confirmed at up to 56.5 cm (22.2 in) in total length (including extended legs and cerci, the latter of which are unusually long in this species); one captive individual was measured at 61.5 cm (24.2 in), but that size remains unconfirmed. [23] [24] [25] Another very large species is Phobaeticus kirbyi where the total length (including extended legs) is up to 54.6 cm (21.5 in) and the body alone up to 32.8 cm (12.9 in). [26] Another of the longest insect in terms of total length is Phobaeticus serratipes of Malaysia and Singapore, measuring up to 55.5 cm (21.9 in) in total length. [27]

The spiny stick insect ( Heteropteryx dilatata ) of Southeast Asia does not reach the extreme lengths of its cousins, the body reaching up to 16 cm (6.3 in) long, but it is much bulkier. The largest Heteropteryx weighed about 65 g (2.3 oz) and was 3.5 cm (1.4 in) wide across the thickest part of the body. [8]

Cockroaches and termites (Blattodea)

Giant burrowing cockroach MP - Macropanesthia rhinoceros 2.jpg
Giant burrowing cockroach

The largest cockroach in length and wingspan is the South American Megaloblatta longipennis , at up to 9.7 cm (3.8 in) and 18–20 cm (7.1–7.9 in), respectively. [6] [28] Other contenders for longest are Blaberus giganteus , which is found in the same general region and may reach a length of up to 9 or 10 cm (3.5 or 3.9 in), depending on source, [29] [30] and Princisia vanwaerebeki of Madagascar, which grows between 5.6 and 10 cm in length. [31] The heaviest is the Australian giant burrowing cockroach (Macropanesthia rhinoceros), which can attain a length of 8.4 cm (3.3 in) and a weight of 33.5 g (1.18 oz). [6]

Termites

The largest of the termites is the African species Macrotermes bellicosus . The queen of this species can attain a length of 10.6 cm (4.2 in) and breadth of 5.5 cm (2.2 in); other adults, however, are about a third of this size.

Praying mantises (Mantodea)

The giant stick mantises Toxodera maxima and Ischnomantis gigas can reach lengths of over 17 cm, but are more gracile in build than other large mantises. Other species of Toxodera and Ischnomantis , and other genera of “stick mantises” ( Heterochaeta , Solygia and Tauromantis ), can attain lengths almost as large (over 16 cm).

Mantises of the tribe Hierodulini (of which Rhombodera fratricida is the largest formally described by science) can reach lengths of nearly 13 cm (5.1 in) and are more robust than comparably sized mantises of other genera ( Tenodera , Macromantis , Idolomantis , Sphodromantis , Deroplatys , Plistospilota and Stagmatoptera ). Some larger species have been known to capture and consume frogs, lizards, mice, small birds, small fish and even small snakes.

True bugs (Hemiptera)

Giant water bug walking over land Giantwaterbug.jpg
Giant water bug walking over land

The heaviest species of this order are the giant water bugs Lethocerus grandis and Lethocerus maximus . [32] These can surpass a length of 12 cm (4.7 in), [33] although they are more slender and less heavy than most other insects of this size (principally the huge beetles).

The largest wingspan of any hemipteran belongs to the largest species of cicada: Megapomponia imperatoria , which has a head-body length of about 7 cm (2.8 in) and a wingspan of 18–20 cm (7–8 in). [34] [35] The cicadas of the genus Tacua can also grow to comparably large sizes.

The largest total length of any hemipteran belongs to Gigantometra gigas (the largest species of water strider), which has a leg span of more than 25 cm (9.8 in).

The largest type of aphid is the giant oak aphid ( Stomaphis quercus ), which can reach an overall length of 2 cm (0.79 in). [36] The biggest species of leafhopper is Ledromorpha planirostris , which can reach a length of 2.8 cm (1.1 in). [37]

Dobsonflies and relatives (Megaloptera)

Megaloptera includes dobsonflies, alderflies and relatives. The largest is the dobsonfly Acanthacorydalis fruhstorferi , which can have a wingspan of up to 21.6 cm (8.5 in), making it the largest aquatic insect in the world by this measurement. [38] This species is native to China and Vietnam, and its body can be up to 10.5 cm (4.1 in) long. [39]

Net-winged insects (Neuroptera)

Blue eyes lacewing Nymphes myrmeleonides 4.jpg
Blue eyes lacewing

These flying insects reach their largest size in Palparellus voeltzkowi , which can have a wingspan over 16 cm (6.3 in). [40] The largest lacewing is the Australian "blue eyes lacewing" ( Nymphes myrmeleonides ), which can measure up to 4 cm (1.6 in) in length and span 11 cm (4.3 in) across the wings. [41] Some forms of this ancient order could grow extremely large during the Jurassic period and may have ranked among the largest insects ever. [42] Found in the Early Cretaceous sedimentary rocks, Makarkinia adamsi had wings nearly 140–160 mm (5.5–6.3 in) in length. [43]

Lice (Phthiraptera)

These parasitic insects are typically modest in size. The largest known species is the hog louse, Haematopinus suis , a sucking louse that lives on large livestock like pigs and cattle. It can range up to 6 mm (0.24 in) in length. [44]

Stoneflies (Plecoptera)

Pteronarcys californica Pteronarcys californica 2544.jpg
Pteronarcys californica

The largest species of stonefly is Pteronarcys californica of western North America, a species favored by fishermen as lures. This species can attain a length of 5 cm (2.0 in) and a wingspan of over 8 cm (3.1 in). [45]

Beetles (Coleoptera)

The hercules beetle, the world's longest beetle Dynastes hercules ecuatorianus MHNT.jpg
The hercules beetle, the world's longest beetle

Beetles constitute the most diverse order of organisms on earth, with about 400,000 species identified to this day. The most massive of them belong to the genera Goliathus , Megasoma , Chalcosoma , Titanus , Macrodontia , and Xixuthrus . The longest is the Hercules beetle, Dynastes hercules, with a maximum overall length of at least 18.1 cm (7.1 in) including the very long pronotal horn. The longest overall beetle is a species of longhorn beetle, Batocera wallacei , from New Guinea, which can attain a length of 26.6 cm (10.5 in), about 19 cm (7.5 in) of which is comprised by the long antennae. [8]

Icebugs (Grylloblattodea)

The largest icebug species, Grylloblatta campodeiformis , 3 cm (1.2 in) long, excluding ovipositors and cerci, and are fairly elongate, wingless insects. They are a uniform honey-yellow in colour and covered with very short hair. Unlike some other species of grylloblatid, G. campodeiformis has eyes which have roughly 70 facets. The head is fairly flat and rounded. The thorax is elongate and over a third of the body length. The abdomen is composed of 10 segments and over half the body length. The legs are long and narrow (cursorial) with stout coxae and long femora. [46] Their antennae are long ~9 mm (0.35 in) and thread-like. In adults, the number of antennal subsegments is variable, ranging from 24 to 27. [47]

Ants, bees, and wasps (Hymenoptera)

Megachile pluto, the largest bee. Stavenn Megachile pluto.jpg
Megachile pluto , the largest bee.

The largest of ants, and the heaviest species of the family, are the queens of Dorylus helvolus , reaching a length of 5 cm (2.0 in). The ant that averages the largest for the mean size of the whole colony is Dinoponera gigantea , averaging up to 3.3 cm (1+14 in). Another ant that is native to Australia, Myrmecia brevinoda , workers are reported to be 3.7 cm (1.5 in) on average and queens are more than 4 cm (1.6 in) in length. [8]

The largest of the bees is Megachile pluto , the females of which can be 3.8 cm (1+12 in) long, with a 6.3-cm (2.5-in) wingspan.

The largest wingspan of any wasp (and of any hymenopteran) probably belongs to the so-called tarantula hawk species Pepsis heros , measuring up to 12.2 cm (4.8 in) in wingspan, although many other Pepsis species approach a similar size and other species of wasp can probably attain greater weights.

The heaviest wasp (and heaviest hymenopteran) is probably the giant scoliid wasp Megascolia procer , it can reach a body length of over 7.7 cm (3.0 in) and have a wingspan of 11.6 cm (4.6 in).

The largest known social wasps are the queens of the Asian giant hornet (Vespa mandarinia). They can have a body length exceeding 6 cm (2.4 in) and a 9.3 cm (3.7 in) wingspan. [48]

Butterflies and moths (Lepidoptera)

The Queen Alexandra's birdwing is the largest species of butterfly. MP - Ornithoptera alexandrae 3.jpg
The Queen Alexandra's birdwing is the largest species of butterfly.

The largest lepidopteran species overall is often claimed to be either the Queen Alexandra's birdwing (Ornithoptera alexandrae), a butterfly from Papua New Guinea, or the Atlas moth ( Attacus atlas ), a moth from Southeast Asia. Both of these species can reach a body length of 8 cm (3.1 in), a wingspan of 28 cm (11 in) and a weight of 12 g (0.42 oz). One Atlas moth allegedly had a wingspan of 30 cm (12 in) but this measurement was not verified. [8] The larvae in the previous species can weigh up to 58 and 54 g (2.0 and 1.9 oz), respectively.

However, the white witch of Central and South America, Thysania agrippina , has the widest recorded wingspan of the order, and indeed of any living insect, at up to 30 cm (12 in). The verified record-sized Thysania spanned 30.8 cm (12.1 in) across the wings, although specimens have been reported to 36 cm (14 in). [8]

The largest lepidopteran species in terms of adult body mass is the giant wood moth Endoxyla cinereus from Australia and New Zealand. Despite having a smaller wingspan than the other species, it has a mass of about 30 g (1.1 oz) and outweighs them all.

The Hercules moth (Coscinocera hercules), in the family Saturniidae, is endemic to New Guinea and northern Australia, and its wings have the largest documented surface area (300 square centimeters) of any living insect, [49] [50] and a maximum wingspan which is confirmed to 28 cm (11 in) while unconfirmed specimens have spanned up to 35.5 cm (14.0 in).

True flies (Diptera)

Gauromydas heros, one of the world's largest true flies. Mydas sp..JPG
Gauromydas heros , one of the world's largest true flies.

The largest total length of any fly belongs to the largest species of crane fly: Holorusia clavipes . If the legs are extended in front of and behind the body, then an overall length of 25.8 cm (10.2 in) makes it the longest true fly. It also probably has the largest wingspan of any Diptera at over 11 cm (4.3 in).

Gauromydas heros (which can reach a wingspan of 10 cm (3.9 in) and a body length of 7 cm (2.8 in)) has the largest wing surface area of any Diptera. It might also have the longest body length of any fly at 7 cm (2.8 in) (although it is rivaled by Microstylum magnum and Phellus piliferus for this title).

The heaviest flies are the timber flies from the genus Pantophthalmus (with Pantophthalmus bellardi being the largest species of this genus known). The largest members of Asilidae ( Microstylum , Phellus and Blepharotes ) are almost as bulky.

Booklice (Psocoptera)

The largest of this order of very small insects is the barklouse of the genus Psocus , the maximum size of which is about 1 cm (0.39 in).

Fleas (Siphonaptera)

The largest species of flea is Hystrichopsylla schefferi . This parasite, known exclusively from the fur of the mountain beaver, can reach a length of 1.2 cm (0.47 in). [51]

Thrips (Thysanoptera)

Members of the genus Phasmothrips are the largest thrips. The maximum size of these species is about 1.3 cm (0.51 in).

Caddisflies (Trichoptera)

The largest of the small, moth-like caddisflies is Eubasilissa maclachlani . This species can range up to 7 cm (2.8 in) across the wings. [52]

Silverfishes and allies (Zygentoma)

These insects, known to feed on human household objects, are up to 1.5 cm (0.59 in) in length. [53] The largest is the extinct Ramsdelepidion schusteri , which reached 6 cm (2.4 in) long. [4]

Angel insects (Zoraptera)

The largest angel insect species, Hubbard's angel insect (Zorotypus hubbardi), grows up to 3 mm (0.12 in) in length. [54] [55]

See also

Related Research Articles

<span class="mw-page-title-main">Blackpoll warbler</span> Species of bird

The blackpoll warbler is a New World warbler. Breeding males are mostly black and white. They have a prominent black cap, white cheeks, and white wing bars. The blackpoll breeds in forests of northern North America, from Alaska throughout most of Canada, to the Adirondack Mountains of New York as well as New England in the Northeastern United States. They are a common migrant throughout much of North America. In fall, they fly south to the Greater Antilles and the northeastern coasts of South America in a non-stop long-distance migration over open water, averaging 2,500 km (1,600 mi), one of the longest-distance non-stop overwater flights ever recorded for a migratory songbird. Rare vagrants to western Europe, they are one of the more frequent transatlantic passerine wanderers.

<span class="mw-page-title-main">Goliath frog</span> Species of amphibian

The goliath frog, otherwise known commonly as the giant slippery frog and the goliath bullfrog, is a species of frog in the family Conrauidae. The Goliath frog is the largest living frog. Specimens can reach up to about 35 centimetres (14 in) in snout–vent length and 3.3 kilograms (7.3 lb) in weight. This species has a relatively small habitat range in Cameroon and Equatorial Guinea. Its numbers are dwindling due to habitat destruction, collection for food, and the pet trade.

<span class="mw-page-title-main">Green anaconda</span> Species of snake

The green anaconda, also known as the giant anaconda, emerald anaconda, common anaconda, common water boa, or southern green anaconda, is a semi-aquatic boa species found in South America and the Caribbean island of Trinidad. It is the largest, heaviest, and second longest snake in the world, after the reticulated python. No subspecies are currently recognized. Like all boas, it is a non-venomous constrictor. The term "anaconda" often refers to this species, though the term could also apply to other members of the genus Eunectes. Fossils of the snake date back to the Late Pleistocene in the Gruta do Urso locality.

<span class="mw-page-title-main">Dinosaur size</span> Dinosaur mass and length estimates

Size is an important aspect of dinosaur paleontology, of interest to both the general public and professional scientists. Dinosaurs show some of the most extreme variations in size of any land animal group, ranging from tiny hummingbirds, which can weigh as little as two grams, to the extinct titanosaurs, such as Argentinosaurus and Bruhathkayosaurus which could weigh as much as 50–130 t.

<span class="mw-page-title-main">Largest body part</span>

The largest body part is either the largest given body part across all living and extinct organisms or the largest example of a body part within an existing species. The largest animals on the planet are not the only ones to have large body parts, with some smaller animals actually having one particularly enlarged area of the body.

The smallest organisms found on Earth can be determined according to various aspects of organism size, including volume, mass, height, length, or genome size.

The purple needletail, or hagibas in Tagalog, is the largest swift found in the Old World. It is native to the Philippine archipelago and the Minahasa Peninsula (Sulawesi).

<span class="mw-page-title-main">Cephalopod size</span> Body variation

Cephalopods, which include squids and octopuses, vary enormously in size. The smallest are only about 1 centimetre (0.39 in) long and weigh less than 1 gram (0.035 oz) at maturity, while the giant squid can exceed 10 metres (33 ft) in length and the colossal squid weighs close to half a tonne (1,100 lb), making them the largest living invertebrates. Living species range in mass more than three-billion-fold, or across nine orders of magnitude, from the lightest hatchlings to the heaviest adults. Certain cephalopod species are also noted for having individual body parts of exceptional size.

<span class="mw-page-title-main">Western mastiff bat</span> Species of bat

The western mastiff bat, also known as the western bonneted bat, the greater mastiff bat, or the greater bonneted bat, is a member of the free-tailed bat family, Molossidae. It is found in the Western United States, Mexico and South America. This species is the largest bat native to North America, and some of its distinguishing characteristics are its large ears, wings, and forearms. The subspecies Eumops perotis californicus is a species of concern as identified by the U.S. Fish and Wildlife Service. The range of this subspecies is principally southwest desert regions of the United States, along the border with Mexico; however, the range extends as far north on the Pacific coast as Alameda County, California.

<span class="mw-page-title-main">Pel's fishing owl</span> Species of owl

Pel's fishing owl is a large species of owl in the family Strigidae, found in Africa. It lives near rivers and lakes, and feeds nocturnally on fish and frogs snatched from the surface of lakes and rivers. The species prefers slow-moving rivers with large, overhanging trees to roost in and forage from. It nests in hollows and the forks of large trees. Though as many as two eggs are laid, often only one chick is raised.

<span class="mw-page-title-main">Largest organisms</span> List of largest organisms on Earth

This article lists the largest organisms for various types of life and mostly considers extant species, which found on Earth can be determined according to various aspects of an organism's size, such as: mass, volume, area, length, height, or even genome size. Some organisms group together to form a superorganism, but such are not classed as single large organisms. The Great Barrier Reef is the world's largest structure composed of living entities, stretching 2,000 km (1,200 mi) but contains many organisms of many types of species.

<span class="mw-page-title-main">Largest prehistoric animals</span>

The largest prehistoric animals include both vertebrate and invertebrate species. Many of them are described below, along with their typical range of size. Many species mentioned might not actually be the largest representative of their clade due to the incompleteness of the fossil record and many of the sizes given are merely estimates since no complete specimen have been found. Their body mass, especially, is largely conjecture because soft tissue was rarely fossilized. Generally the size of extinct species was subject to energetic and biomechanical constraints.

<span class="mw-page-title-main">Largest and heaviest animals</span>

The largest animal currently alive is the blue whale. The maximum recorded weight was 190 tonnes for a specimen measuring 27.6 metres (91 ft), whereas longer ones, up to 33 metres (108 ft), have been recorded but not weighed. It is estimated that this individual could have a mass of 250 tonnes or more. The longest non-colonial animal is the lion's mane jellyfish.

<i>Phryganistria</i> Genus of stick insects

Phryganistria is a genus of stick insects belonging to the subfamily Clitumninae, native to northeastern South Asia, northern Mainland Southeast Asia, and southern China. The species in this genus range in size from fairly large to very large.

<i>Tetracanthagyna plagiata</i> Species of dragonfly

The giant hawker or the gigantic riverhawker, is a species of dragonfly in the family Aeshnidae. It is found throughout Sundaland, having been recorded on Thailand, Peninsular Malaysia, Singapore, Sumatra, and Borneo. It is the type species for the genus Tetracanthagyna.

References

  1. Zhang Z-Q, ed. (2011) Animal Biodiversity: An Outline of Higher-Level Classification and Survey of Taxonomic Richness. Auckland, N. Z.: Magnolia Press
  2. 1 2 3 Williams, David M (2001-04-21). "Chapter 30 — Largest Insect". Book of Insect Records. University of Florida. Archived from the original on 2014-08-20.
  3. Dorrington, Graham E. (2016-04-01). "Heavily loaded flight and limits to the maximum size of dragonflies (Anisoptera) and griffenflies (Meganisoptera)". Lethaia. 49 (2): 261–274. doi:10.1111/let.12144. ISSN   1502-3931.
  4. 1 2 Polet, Delyle (2011-05-06). "The Biggest Bugs: An investigation into the factors controlling the maximum size of insects". Eureka. 2 (1): 43–46. doi: 10.29173/eureka10299 . ISSN   1923-1520.
  5. Kukalová-Peck, Jarmila; Richardson, Eugene S. (1983). "New Homoiopteridae (Insecta: Paleodictyoptera) with wing articulation from Upper Carboniferous strata of Mazon Creek, Illinois". Canadian Journal of Zoology. 61 (7): 1670–1687. doi:10.1139/z83-218.
  6. 1 2 3 Carwardine, M. (2007). Animal Records. Natural History Museum, London. pp. 229–230. ISBN   978-1-4027-5623-8.
  7. 1 2 3 Flindt, R. (2006). Amazing Numbers in Biology . Springer. p.  10. ISBN   978-3-540-30146-2.
  8. 1 2 3 4 5 6 Wood, Gerald The Guinness Book of Animal Facts and Feats (1983) ISBN   978-0-85112-235-9
  9. Order Orthoptera, Orthoptera Species File (Version 5.0/5.0); retrieved 20 August 2022
  10. Park, Tae-Yoon S.; Kim, Do-Yoon; Nam, Gi-Soo; Lee, Mirinae (2022-05-06). "A new titanopteran Magnatitan jongheoni n. gen. n. sp. from southwestern Korean Peninsula". Journal of Paleontology. 96 (5): 1111–1118. doi: 10.1017/jpa.2022.30 . ISSN   0022-3360. S2CID   248592776.
  11. McKeown, Keith C. (1937-05-15). "New fossil insect wings (Protohemiptera, family Mesotitanidae)". Records of the Australian Museum. 20 (1): 31–37. doi: 10.3853/j.0067-1975.20.1937.565 . ISSN   0067-1975.
  12. "Labidura". Archived from the original on 2011-07-17. Retrieved 2011-02-05.
  13. UK Safari
  14. Bradler, S.; Buckley, T.R. (2018). "Biodiversity of Phasmatodea". In Foottit, R.G.; Adler, P.H. (eds.). Insect Biodiversity: Science and Society. Vol. II. John Wiley & Sons Ltd. pp. 281–313. doi:10.1002/9781118945582.ch11. ISBN   978-1-118-94557-5.
  15. Baggaley, K. (6 May 2016). "World's Longest Insect Is Two Feet Long". PopularScience. Retrieved 25 June 2023.
  16. "Longest Insect discovered in China". Archived from the original on May 8, 2016.
  17. "The discovery of the world's longest insect. Crawling "tree branches"". Google Arts and Culture. Retrieved 10 March 2023.
  18. Xuequan, M. (8 August 2017). "China breeds world's largest insect". xinhuanet. Retrieved 25 June 2023.
  19. "Bugs beyond belief! Shining the spotlight on celebrity creepy-crawlies". Guinness World Records. 29 November 2018. Retrieved 25 June 2023.
  20. "World's longest insect revealed". London: Natural History Museum. 2008-10-16. Archived from the original on 2008-10-19. Retrieved 2008-10-16.
  21. Seow-Choen, F. (1995). "The longest insect in the world". Malayan Naturalist48: 12.
  22. Hennemann, F. H. & Conle, O.V. (October 2008). "Revision of Oriental Phasmatodea: The tribe Pharnaciini Günther, 1953, including the description of the world's longest insect, and a survey of the family Phasmatidae Gray, 1835 with keys to the subfamilies and tribes (Phasmatodea: "Anareolatae": Phasmatidae)" (PDF). Zootaxa. 1906. Auckland, New Zealand: Magnolia Press: 1–316. doi:10.11646/zootaxa.1906.1.1. ISSN   1175-5326 . Retrieved 2008-10-16.
  23. Brock, Paul D.; Hasenpusch, Jack W. (2009). The Complete Field Guide to Stick and Leaf Insects of Australia. CSIRO Publishing. p. 106. ISBN   9780643094185.
  24. Dean, S. (15 February 2016). "Gargantuan Aussie stick insect has babies in a world first". Special Broadcasting Service . Retrieved 25 June 2023.
  25. Cane, S. (10 March 2014). "Australia's biggest stick insect found". Australian National Geographic. Retrieved 25 June 2023.
  26. Brock, P. D. (1999). The amazing world of stick and leaf-insects. Essex, England: Cravitz.
  27. Seow-Choen, F. (1995). "The longest insect in the world". Malayan Nat. 48: 12.
  28. "Largest cockroach". Guinness World Records. Retrieved 30 March 2018.
  29. "Allpet Roaches;Blattodea Rearing". Allpet. Retrieved 2019-07-15.
  30. Davis, Shkora (2015). "Blaberus giganteus (Giant Cave Cockroach)" (PDF). The Online Guide to the Animals of Trinidad and Tobago.
  31. Delfosse, E. (2004). "Les blattes souffleuses de Madagascar". Insectes. 153 (4): 19–22.
  32. P.J., Perez-Goodwyn (2006). "Taxonomic revision of the subfamily Lethocerinae Lauck & Menke (Heteroptera: Belostomatidae)". Stuttgarter Beiträge zur Naturkunde, Serie A (Biologie). 695: 1–71.
  33. Haddad, V.; Schwartz; Schwartz; Carvalho (2010). "Bites Caused by Giant Water Bugs Belonging to Belostomatidae Family (Hemiptera, Heteroptera) in Humans: A Report of Seven Cases". Wilderness & Environmental Medicine. 21 (2): 130–133. doi:10.1016/j.wem.2010.01.002. PMID   20591375.
  34. Burton, Maurice; Burton, Robert (2002). International Wildlife Encyclopedia. 4, Chickaree – crabs (3rd ed.). Tarrytown, NY: Marshall Cavendish Corporation. p. 455. ISBN   0-7614-7270-3
  35. Flindt, R. (2006). Amazing Numbers in Biology, p. 10. ISBN   978-3540301462
  36. Giant Oak Aphid hunt is on. The Telegraph (2007-08-08)
  37. Ledromorpha planirostris. Bugs.bio.usyd.edu.au
  38. "Largest aquatic insect (by wingspan)". Guinness World Records. Retrieved 4 April 2018.
  39. Xing Yue Liu; Ding Yang; Si Qin Ge; Xing Ke Yang (2005). "Phylogenetic review of the Chinese species of Acanthacorydalis (Megaloptera, Corydalidae)". Zoologica Scripta. 34 (4): 373–387. doi:10.1111/j.1463-6409.2005.00197.x. S2CID   84988447.
  40. Palparellus voeltzkowi (Kolbe, 1906). Researcharchive.calacademy.org
  41. Bio-Ditrl, Department of Biological Sciences, University of Alberta Archived July 6, 2011, at the Wayback Machine
  42. Michael S. Engel (2005). "A remarkable kalligrammatid lacewing from the Upper Jurassic of Kazakhstan (Neuroptera: Kalligrammatidae)". Transactions of the Kansas Academy of Science. 108 (1): 59–62. doi:10.1660/0022-8443(2005)108[0059:arklft]2.0.co;2. JSTOR   3628206. S2CID   86240200.
  43. Bechly, G.; Makarkin, V. N. (2016). "A new gigantic lacewing species (Insecta: Neuroptera) from the Lower Cretaceous of Brazil confirms the occurrence of Kalligrammatidae in the Americas". Cretaceous Research. 58: 135–140. doi:10.1016/j.cretres.2015.10.014.
  44. ADW: Haematopinus suis: Information. Animaldiversity.ummz.umich.edu
  45. Pteronarcys californica – aka Giant Stonefly or Giant Salmonfly. Riverwood Blog – Fly Fishing Gear & Guided Fishing Trips in Oregon (2009-04-20) Archived April 2, 2012, at the Wayback Machine
  46. Walker, Edmund (1914). "A new species of Orthoptera, forming a new genus and family". The Canadian Entomologist. 46 (3): 93–99. doi:10.4039/ent4693-3. S2CID   85329926 . Retrieved 2014-12-19.
  47. Slifer, Eleanor H. (1976). "Sense organs on antennal flagellum of Grylloblata campodeiformis E.M. Walker (Orthoptera: Grylloblattodea)". Transactions of the American Entomological Society. 87 (9 & 10): 275–276. Retrieved 2014-12-25.
  48. Vespa mandarinia factsheet
  49. Robert G. Foottit & Peter H. Adler. 2009. Insect Biodiversity: Science and Society. Blackwell Publishing Ltd. ISBN   978-1-405-15142-9
  50. Rainier Flindt. 2006. Amazing Numbers in Biology. Springer-Verlag, Berlin. ISBN   3-540-30146-1
  51. Yoon, Carol Kaesuk (28 July 2014). "The Great Giant Flea Hunt". The New York Times . Retrieved 26 October 2021.
  52. Diptera.info – Discussion Forum: The LARGEST caddisfly of the world.
  53. "Order Zygentoma - Silverfish". bugguide.net. Retrieved 2024-02-12.
  54. Foottit, Robert G.; Adler, Peter H. (2018). Insect Biodiversity: Science and Society. Wiley. pp. 199–207. ISBN   978-1-118-94560-5.
  55. Engel, Michael S. (2005). "Zoraptera". Tree of Life Web Project. Retrieved 9 March 2019.