The family Scarabaeidae, as currently defined, consists of over 35,000 species of beetles worldwide; they are often called scarabs or scarab beetles. The classification of this family has undergone significant change in recent years. Several subfamilies have been elevated to family rank, and some reduced to lower ranks. The subfamilies listed in this article are in accordance with those in Catalog of Life (2023).
Scarabaeoidea is a superfamily of beetles, the only subgroup of the infraorder Scarabaeiformia. Around 35,000 species are placed in this superfamily and some 200 new species are described each year. Its constituent families are also undergoing revision presently, and the family list below is only preliminary. This superfamily includes some of the largest beetles extant today, including rhinoceros beetles, (Dynastinae), the Hercules beetle and Goliath beetles.
The Hercules beetle is a species of rhinoceros beetle native to the rainforests of southern Mexico, Central America, South America, and the Lesser Antilles. It is the longest extant species of beetle in the world, and is also one of the largest flying insects in the world.
Dynastinae or rhinoceros beetles are a subfamily of the scarab beetle family (Scarabaeidae). Other common names – some for particular groups of rhinoceros beetles – include Hercules beetles, unicorn beetles or horn beetles. Over 1,500 species and 225 genera of rhinoceros beetles are known.
Ludwigia is a genus of about 82 species of aquatic plants with a cosmopolitan but mainly tropical distribution.
Dynastes is a genus of large beetles belonging to the family Scarabaeidae. They occur in the Nearctic realm and in the Neotropical realm, from the United States to Brazil; four North American species, three with distributions extending from Central America either north or south, and two species endemic to South America.
Tyrannasorus rex is an extinct species of hybosorid beetle known to exist in the Miocene epoch and the sole member of the monotypic genus Tyrannasorus. A fossilized example scarabaeoid was found embedded in the amber resin of Hymenaea protera in the Dominican Republic. The species was described by Brett C. Ratcliffe and Federico Carlos Ocampo in 2001.
Coprophanaeus is a genus in the family Scarabaeidae. The genus is almost entirely Neotropical, with a single species, C. pluto, ranging into southernmost Texas in the United States. They are medium-sized to large beetles, with the South American C. ensifer and C. lancifer sometimes exceeding 5 cm (2 in) in length, making these two some of the largest dung beetles in the world and the largest in the Americas. They often have a horn on the head, and are typically a bright metallic color, most often blue or green, or black. These diurnal or crepuscular beetles are excellent diggers and good fliers.
Aphodiites is a genus of fossil beetles from the Lias of Schambelen, and the oldest fossil in the superfamily Scarabaeoidea. Its affinities are not apparent; it was initially placed in the Aphodiinae (Scarabaeidae), but its diagnostic characters are shared by beetles such as Glaresis (Glaresidae).
Sequoia is a genus of redwood coniferous trees in the subfamily Sequoioideae of the family Cupressaceae. The only extant species of the genus is Sequoia sempervirens in the Northern California coastal forests ecoregion of Northern California and Southwestern Oregon in the United States. The two other genera in the subfamily Sequoioideae, Sequoiadendron and Metasequoia, are closely related to Sequoia. It includes the tallest trees, as well as the heaviest, in the world.
Titanomyrma is a genus of extinct giant ants which lived during the Eocene. The type species Titanomyrma gigantea and the smaller Titanomyrma simillima are known from the Eocene of Germany, while the third species Titanomyrma lubei, is known from Wyoming, United States. The presence of Titanomyrma in North America was considered to indicate "the first reported cross-Arctic dispersal by a thermophilic insect group".. However a queen reported from Upland temperate shales in British Columbia raised questions on the exact thermophilic nature of the genus. The type species of this genus, T. gigantea, is the largest-known fossil or extant species of ant in the world.
The western Hercules beetle is a species of rhinoceros beetle that lives in Arizona, New Mexico and Utah in the United States and in parts of northern Mexico. This species is known for its grayish-white elytra, large size, and characteristic horn of the adult males.
Eosacantha is an extinct genus of tortoise beetle in the family Chrysomelidae and containing a single species Eosacantha delocranioides. The species is known only from the Middle Eocene Parachute Member, part of the Green River Formation, in the Piceance Creek Basin, Garfield County, northwestern Colorado, USA.
Afairiguana avius is an extinct iguanid lizard known from a nearly complete and articulated skeleton discovered in rocks of the Early Eocene-aged Green River Formation of Wyoming, United States. As of the initial description, the skeleton represents the oldest complete iguanian from the Western Hemisphere, and is the oldest representative of the extant iguanid family of anoles, Polychrotidae.
Paleopanax is an extinct genus of flowering plant in the Ginseng and Ivy family, Araliaceae, containing the single species Paleopanax oregonensis. The species is solely known from the middle Eocene sediments exposed in north central Oregon and was first described from a series of isolated fossil fruits in siltstones.
Phanaeus, the rainbow scarabs, is a genus of true dung beetles in the family Scarabaeidae, ranging from the United States to northern Argentina, with the highest species richness in Mexico. Depending on species, they can inhabit a wide range of habitats, from tropical to temperate climates and deserts to rainforests. In those living in relatively arid places adults are primarily active during the wet season and those living in relatively cold places are primarily active during the summer. They are excellent diggers and good fliers.
Protognathinus is an extinct genus of stag beetle from the Eocene of Europe, known from the Messel Pit in Germany. This genus is known from the single species P. spielbergi.
Comptonia columbiana is an extinct species of sweet fern in the flowering plant family Myricaceae. The species is known from fossil leaves found in the early Eocene deposits of central to southern British Columbia, Canada, plus northern Washington state, United States, and, tentatively, the late Eocene of Southern Idaho and Earliest Oligocene of Oregon, United States.
Amia? hesperia is an extinct species of bony fish in the bowfin family, Amiidae. The species is known from fossils found in the early Eocene deposits of northern Washington state in the United States and southeastern British Columbia. The species is one of eight fish species identified in the Eocene Okanagan Highlands paleofauna.
Notonuphar is an extinct genus of water lily in the family Nymphaeaceae. It contains a single species, Notonuphar antarctica. It is only known only from the Eocene-aged La Meseta Formation of Seymour Island, Antarctica.
