Polish cochineal | |
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Life cycle of the Polish cochineal in Breyne's Historia naturalis Cocci Radicum... (1731) | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Class: | Insecta |
Order: | Hemiptera |
Suborder: | Sternorrhyncha |
Family: | Margarodidae |
Genus: | Porphyrophora |
Species: | P. polonica |
Binomial name | |
Porphyrophora polonica | |
Area where the Polish cochineal was found in commercial quantities [1] | |
Synonyms [2] | |
Polish cochineal (Porphyrophora polonica, Margarodes polonicus), also known as Polish carmine scales (Polish : czerwiec polski), is a scale insect formerly used to produce a crimson dye of the same name, colloquially known as "Saint John's blood". The larvae of P. polonica are sessile parasites living on the roots of various herbs – especially those of the perennial knawel (Polish : czerwiec trwały) – growing on the sandy soils of Central Europe and other parts of Eurasia. Before the development of aniline, alizarin, and other synthetic dyes, the insect was of great economic importance, although its use was in decline after the introduction of Mexican cochineal to Europe in the 16th century. Historically earlier was used Armenian cochineal scale insect, Porphyrophora hamelii, which is in a same taxonomic family Porphyrophora of the Polish cochineal and in different taxonomic family from cochineal found in the Americas.
In mid-July, the female Polish cochineal lays approximately 600-700 eggs, encased with a white waxy ootheca, in the ground. When the larvae hatch in late August or early September, they do not leave the egg case but remain inside until the end of winter. In late March or early April, the larvae emerge from the ground to feed for a short time on the low-growing leaves of the host plant before returning underground to feed on the plant's roots. At this point, the larvae undergo ecdysis, shedding their exoskeletons together with their legs and antennae, and they encyst by forming outer protective coatings (cysts) within the root tissues. [4]
The cysts are small dark red or violet bubbles clustered on the host plant's roots. Female cysts are 3–4 millimetres (0.12–0.16 in) in diameter. Males are half the size of their female counterparts and fewer in number, with only one male per 500 females. The cysts undergo ecdysis a number of times. When the male larva reaches the third-instar developmental stage, it forms a delicate white cocoon and transforms into a pupa in early June. In late June or early July, females, which are neotenous and retain their larval form, re-emerge from the ground and slowly climb to the top of the host plant, where they wait until winged adult males, with characteristic plumes at the end of their abdomens, leave the cocoons and join them a few days later. Male imagines (adult insects) do not feed and die shortly after mating, while their female counterparts return underground to lay eggs. After oviposition, the female insects shrink and die. [4]
The Polish cochineal lives on herbaceous plants growing in sandy and arid, infertile soils. Its primary host plant is the perennial knawel (Scleranthus perennis), but it has also been known to feed on plants of 20 other genera, including mouse-ear hawkweed (Hieracium pilosella), bladder campion (Silene inflata), velvet bent (Agrostis canina), Caragana , [4] smooth rupturewort (Herniaria glabra), strawberry (Fragaria), and cinquefoil ( Potentilla ). [3]
The insect was once commonly found throughout the Palearctic [4] and was recognised across Eurasia, from France and England [3] to China, but it was mainly in Central Europe where it was common enough to make its industrial use economically viable. Excessive economic exploitation as well as the shrinking and degradation of its habitat have made the Polish cochineal a rare species. In 1994, it was included in the Ukrainian Red Book of endangered species. [5] In Poland, where it was still common in the 1960s, there is insufficient data to determine its conservation status, and no protective measures are in place. [4]
Ancient Slavs developed a method of obtaining red dye from the larvae of the Polish cochineal. Despite the labor-intensive process of harvesting the cochineal and a relatively modest yield, the dye continued to be a highly sought-after commodity and a popular alternative to kermes throughout the Middle Ages until it was superseded by Mexican cochineal in the 16th century.[ citation needed ]
Similar to some other red dyes obtained from scale insects, the red coloring is derived from carminic acid with traces of kermesic acid. The Polish cochineal carminic acid content is approximately 0.6% of the insect's dried body weight. [6] The insects were harvested shortly before the female larvae reached maturity, i.e. in late June, usually around Saint John the Baptist's day (June 24), hence the dye's folk name, Saint John's blood. The harvesting process involved uprooting the host plant and picking the female larvae, averaging approximately ten insects from each plant. [7] In Poland, including present-day Ukraine, and elsewhere in Europe, plantations were operated in order to deal with the high toll on the host plants. [1] The larvae were killed with boiling water or vinegar and dried in the sun or in an oven, ground, and dissolved in sourdough or in light rye beer called kvass [8] in order to remove fat. The extract could then be used for dyeing silk, wool, cotton, or linen. [8] The dyeing process requires roughly 3-4 oz of dye per pound (180-250 g per kilogram) of silk [7] and one pound of dye to color almost 20 pounds (50 g per kilogram) of wool. [8]
Polish cochineal was widely traded in Europe during the Middle Ages and the Renaissance. In the 15th and 16th centuries, along with grain, timber, and salt, it was one of Poland's chief exports, mainly to southern Germany and northern Italy as well as to France, England, the Ottoman Empire, and Armenia. [7] In Poland, the cochineal trade was mostly monopolized by Jewish merchants, [7] who bought the dye from peasants in Red Ruthenia and other regions of Poland and Lithuania. The merchants shipped the dye to major Polish cities such as Kraków, Danzig (Gdańsk), and Poznań. From there, the merchandise was exported to wholesalers in Breslau (Wrocław), Nuremberg, Frankfurt, Augsburg, Venice, [7] and other destinations. The Polish cochineal trade was a lucrative business for the intermediaries; according to Marcin of Urzędów (1595), one pound of Polish cochineal cost between four and five Venetian pounds. In terms of quantities, the trade reached its peak in the 1530s. In 1534, 1963 stones (about 30 metric tons) of the dye were sold in Poznań alone. [7]
The advent of cheaper Mexican cochineal led to an abrupt slump in the Polish cochineal trade, and the 1540s saw a steep decline in quantities of the red dye exported from Poland. In 1547, Polish cochineal disappeared from the Poznań customs registry; a Volhynian clerk noted in 1566 that the dye no longer paid in Gdańsk. Perennial knawel plantations were replaced with cereal fields or pastures for raising cattle. Polish cochineal, which until then was mostly used by the rich nobles and the royals and as an export product, continued to be used locally by the peasants who collected it; it was employed not only for dyeing fabric but also as a vodka colorant, an ingredient in folk medicine, crafts or even for decorative coloring of horses' tails. [7]
With the partitions of Poland at the end of the 18th century, vast markets in Russia and Central Asia opened to Polish cochineal, which became an export product again – this time, to the East. In the 19th century, Bukhara, Uzbekistan, became the principal Polish cochineal trading center in Central Asia; from there the dye was shipped to Kashgar in Xinjiang, and Kabul and Herat in Afghanistan. It is possible that the Polish dye was used to manufacture some of the famous oriental rugs. [1]
The earliest known scientific study of the Polish cochineal is found in the Herbarz Polski (Polish Herbal ) by Marcin of Urzędów (1595), where it was described as "small red seeds" that grow under plant roots, becoming "ripe" in April and from which a little "bug" emerges in June. [7] The first scientific comments by non-Polish authors were written by Segerius (1670) and von Bernitz (1672). [1] In 1731, Johann Philipp Breyne, wrote Historia naturalis Cocci Radicum Tinctorii quod polonicum vulgo audit (translated into English during the same century), the first major treatise about the insect, including the results of his research on its physiology and life cycle. [7] In 1934, Polish biologist Antoni Jakubski wrote Czerwiec polski (Polish cochineal), a monograph taking into account both the insect's biology and historical role.[ citation needed ]
The historical importance of the Polish cochineal is still reflected in most modern Slavic languages where the words for the color red and for the month of June both derive from the Proto-Slavic *čьrvь (probably pronounced [t͡ʃĭrwĭ] ), meaning "a worm" or "larva". [9] (See examples in the table below.)
In Czech, as well as old Bulgarian, this is true for both June and July, the two months when harvest of the insect's larvae was possible. In modern Polish, czerwiec is a word for June, as well as for the Polish cochineal (czerwiec polski) and its host plant, the perennial knawel (czerwiec trwały).[ citation needed ]
English | Belarusian | Ukrainian | Polish | Czech | Bulgarian |
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worm, larva | чарвяк charvyak | черв'як cherv'yak | czerw | červ | червей chervey |
red (adj.) | чырвоны chyrvony | червоний chervonyy | czerwony czerwień | červený | червен cherven |
June | чэрвень chervyen' | червень cherven' | czerwiec | červen | червеник chervenik |
July | červenec | чръвенъ chraven | |||
Polish cochineal | чэрвек chervyeк | червець chervets' | czerwiec polski |
Lac is the resinous secretion of a number of species of lac insects, of which the most commonly cultivated is Kerria lacca.
Galls or cecidia are a kind of swelling growth on the external tissues of plants. Plant galls are abnormal outgrowths of plant tissues, similar to benign tumors or warts in animals. They can be caused by various parasites, from viruses, fungi and bacteria, to other plants, insects and mites. Plant galls are often highly organized structures so that the cause of the gall can often be determined without the actual agent being identified. This applies particularly to insect and mite plant galls. The study of plant galls is known as cecidology.
Crimson is a rich, deep red color, inclining to purple. It originally meant the color of the kermes dye produced from a scale insect, Kermes vermilio, but the name is now sometimes also used as a generic term for slightly bluish-red colors that are between red and rose. It is the national color of Nepal.
Carmine – also called cochineal, cochineal extract, crimson lake, or carmine lake – is a pigment of a bright-red color obtained from the aluminium complex derived from carminic acid. Specific code names for the pigment include natural red 4, C.I. 75470, or E120. Carmine is also a general term for a particularly deep-red color.
A lake pigment is a pigment made by precipitating a dye with an inert binder, or mordant, usually a metallic salt. Unlike vermilion, ultramarine, and other pigments made from ground minerals, lake pigments are organic. Manufacturers and suppliers to artists and industry frequently omit the lake designation in the name. Many lake pigments are fugitive because the dyes involved are not lightfast. Red lakes were particularly important in Renaissance and Baroque paintings; they were often used as translucent glazes to portray the colors of rich fabrics and draperies.
Scale insects are small insects of the order Hemiptera, suborder Sternorrhyncha. Of dramatically variable appearance and extreme sexual dimorphism, they comprise the infraorder Coccomorpha which is considered a more convenient grouping than the superfamily Coccoidea due to taxonomic uncertainties. Adult females typically have soft bodies and no limbs, and are concealed underneath domed scales, extruding quantities of wax for protection. Some species are hermaphroditic, with a combined ovotestis instead of separate ovaries and testes. Males, in the species where they occur, have legs and sometimes wings, and resemble small flies. Scale insects are herbivores, piercing plant tissues with their mouthparts and remaining in one place, feeding on sap. The excess fluid they imbibe is secreted as honeydew on which sooty mold tends to grow. The insects often have a mutualistic relationship with ants, which feed on the honeydew and protect them from predators. There are about 8,000 described species.
Carminic acid (C22H20O13) is a red glucosidal hydroxyanthrapurin that occurs naturally in some scale insects, such as the cochineal, Armenian cochineal, and Polish cochineal. The insects produce the acid as a deterrent to predators. An aluminum salt of carminic acid is the coloring agent in carmine, a pigment. Natives of Peru had been producing cochineal dyes for textiles since at least 700 CE. Synonyms are C.I. 75470 and C.I. Natural Red 4.
Kermes is a red dye derived from the dried bodies of the females of a scale insect in the genus Kermes, primarily Kermes vermilio. The Kermes insects are native in the Mediterranean region and are parasites living on the sap of the host plant, the Kermes oak and the Palestine oak.
An animal product is any material derived from the body of a non-human animal. Examples are fat, flesh, blood, milk, eggs, and lesser known products, such as isinglass and rennet.
Delia flies are members of the Anthomyiidae family within the superfamily Muscoidae. The identification of different species of Delia can be very difficult for non-specialists as the diagnostic characteristics used for immature and/or female specimens may be inconsistent between species. Past taxonomic keys were not as comprehensive in their identification of Delia specimens; they were either too reliant on genetic characteristics, focused solely on a specific life stage, or were focused only on certain species. However current taxonomic keys aim to be more thorough by not only including morphological diagnostics for males, females, and immature specimens of various species, but also their genetic make-up or molecular barcode.
Johann Philipp Breyne FRS, son of Jacob Breyne (1637–97), was a German-Polish botanist, palaeontologist, zoologist and entomologist. He is best known for his work on the Polish cochineal, an insect formerly used in production of red dye. Proposed by Hans Sloane, he was elected, on 21 April 1703, a Fellow of the Royal Society. He was also a member of the German Academy of Sciences Leopoldina and the Societas Litteraria
The cochineal is a scale insect in the suborder Sternorrhyncha, from which the natural dye carmine is derived. A primarily sessile parasite native to tropical and subtropical South America through North America, this insect lives on cacti in the genus Opuntia, feeding on plant moisture and nutrients. The insects are found on the pads of prickly pear cacti, collected by brushing them off the plants, and dried.
Insect farming is the practice of raising and breeding insects as livestock, also referred to as minilivestock or micro stock. Insects may be farmed for the commodities they produce, or for them themselves; to be used as food, as feed, as a dye, and otherwise.
Natural dyes are dyes or colorants derived from plants, invertebrates, or minerals. The majority of natural dyes are vegetable dyes from plant sources—roots, berries, bark, leaves, and wood—and other biological sources such as fungi.
Dyeing is the craft of imparting colors to textiles in loose fiber, yarn, cloth or garment form by treatment with a dye. Archaeologists have found evidence of textile dyeing with natural dyes dating back to the Neolithic period. In China, dyeing with plants, barks and insects has been traced back more than 5,000 years. Natural insect dyes such as Tyrian purple and kermes and plant-based dyes such as woad, indigo and madder were important elements of the economies of Asia and Europe until the discovery of man-made synthetic dyes in the mid-19th century. Synthetic dyes quickly superseded natural dyes for the large-scale commercial textile production enabled by the industrial revolution, but natural dyes remained in use by traditional cultures around the world.
Dactylopius is a genus of insect in the superfamily Coccoidea, the scale insects. It is the only genus in the family Dactylopiidae. These insects are known commonly as cochineals, a name that also specifically refers to the best-known species, the cochineal. The cochineal is an insect of economic and historical importance as a main source of the red dye carmine. It has reportedly been used for this purpose in the Americas since the 10th century. Genus Dactylopius is also important because several species have been used as agents of biological pest control, and because several are known as invasive species.
The Armenian cochineal, also known as the Ararat cochineal or Ararat scale, is a scale insect indigenous to the Ararat plain and Aras (Araks) River valley in the Armenian Highlands and in Turkey. It was formerly used to produce an eponymous crimson carmine dyestuff known in Armenia as vordan karmir and historically in Persia as kirmiz. The species is critically endangered within Armenia.
Hemipenthes maura is a species of bee fly belonging to the family Bombyliidae.
Red pigments are materials, usually made from minerals, used to create the red colors in painting and other arts. The color of red and other pigments is determined by the way it absorbs certain parts of the spectrum of visible light and reflects the others. The brilliant opaque red of vermillion, for example, results because vermillion reflects the major part of red light, but absorbs the blue, green and yellow parts of white light.