The peppered moth is a temperate species of night-flying moth. Peppered moth evolution is an example of population genetics and natural selection.
A natural experiment is an empirical study in which individuals are exposed to the experimental and control conditions that are determined by nature or by other factors outside the control of the investigators. The process governing the exposures arguably resembles random assignment. Thus, natural experiments are observational studies and are not controlled in the traditional sense of a randomized experiment. Natural experiments are most useful when there has been a clearly defined exposure involving a well defined subpopulation such that changes in outcomes may be plausibly attributed to the exposure. In this sense, the difference between a natural experiment and a non-experimental observational study is that the former includes a comparison of conditions that pave the way for causal inference, but the latter does not.
Lamarckism, also known as Lamarckian inheritance or neo-Lamarckism, is the notion that an organism can pass on to its offspring physical characteristics that the parent organism acquired through use or disuse during its lifetime. It is also called the inheritance of acquired characteristics or more recently soft inheritance. The idea is named after the French zoologist Jean-Baptiste Lamarck (1744–1829), who incorporated the classical era theory of soft inheritance into his theory of evolution as a supplement to his concept of orthogenesis, a drive towards complexity.
Edmund Brisco "Henry" Ford was a British ecological geneticist. He was a leader among those British biologists who investigated the role of natural selection in nature. As a schoolboy Ford became interested in lepidoptera, the group of insects which includes butterflies and moths. He went on to study the genetics of natural populations, and invented the field of ecological genetics. Ford was awarded the Royal Society's Darwin Medal in 1954.
Henry Bernard Davis Kettlewell was a British geneticist, lepidopterist and medical doctor, who performed research on the influence of industrial melanism on peppered moth coloration, showing why moths are darker in polluted areas. This experiment is cited as a classic demonstration of natural selection in action. After live video record of the experiment with Niko Tinbergen, Sewall Wright called the study as "the clearest case in which a conspicuous evolutionary process has actually been observed."
Michael Eugene Nicolas Majerus was a British geneticist and professor of evolution at the University of Cambridge. He was also a teaching fellow at Clare College, Cambridge. He was an enthusiast in Darwin's theory of evolution by natural selection and became a world authority in his field of insect evolutionary biology. He was widely noted for his work on moths and ladybirds and as an advocate of the science of evolution. He was also an enthusiastic educator and the author of several books on insects, evolution and sexual reproduction. He is best remembered as an ardent supporter and champion of experiments on peppered moth evolution.
Of Moths and Men is a book by journalist Judith Hooper about the Oxford University ecological genetics school led by E.B. Ford. The book specifically concerns Bernard Kettlewell's experiments on the peppered moth which were intended as experimental validation of evolution. She highlights concerns about the methodology of Kettlewell's experiments and suggests that these issues could invalidate the results obtained, ignoring or disparaging evidence supporting natural selection while repeatedly implying that Kettlewell and his colleagues committed fraud or made careless errors. Subject matter experts have described the book as presenting a "conspiracy theory" with "errors, misrepresentations, misinterpretations and falsehoods".
The evolution of the peppered moth is an evolutionary instance of directional colour change in the moth population as a consequence of air pollution during the Industrial Revolution. The frequency of dark-coloured moths increased at that time, an example of industrial melanism. Later, when pollution was reduced, the light-coloured form again predominated. Industrial melanism in the peppered moth was an early test of Charles Darwin's natural selection in action, and remains as a classic example in the teaching of evolution. In 1978 Sewall Wright described it as "the clearest case in which a conspicuous evolutionary process has actually been observed."
Bruce S. Grant is emeritus professor of biology at the College of William and Mary. He has a particular research interest in the peppered moth, He is a defender of the teaching of evolution and has criticized creationist Jonathan Wells, who has cited his work, as "dishonest."
The Evolution of Melanism: a study of recurring necessity; with special reference to industrial melanism in the Lepidoptera is a 1973 science book by the lepidopterist Bernard Kettlewell.
The term melanism refers to black pigment and is derived from the Greek: μελανός. Melanism is the increased development of the dark-colored pigment melanin in the skin or hair.
James William Tutt was an English schoolteacher and entomologist. He was a founding editor of the journal Entomologists' Record from 1890 and published a landmark series on the British Lepidoptera in which he described numerous species of moths and was among the first to notice industrial melanism in the pepper moth Biston betularia and was among the first to provide a clear explanation of their increasing frequency based on the role of crypsis, natural selection by predators, and the effect of changed environmental conditions brought on by industrialism.
Industrial melanism is an evolutionary effect prominent in several arthropods, where dark pigmentation (melanism) has evolved in an environment affected by industrial pollution, including sulphur dioxide gas and dark soot deposits. Sulphur dioxide kills lichens, leaving tree bark bare where in clean areas it is boldly patterned, while soot darkens bark and other surfaces. Darker pigmented individuals have a higher fitness in those areas as their camouflage matches the polluted background better; they are thus favoured by natural selection. This change, extensively studied by Bernard Kettlewell, is a popular teaching example in Darwinian evolution, providing evidence for natural selection. Kettlewell's results have been challenged by zoologists, creationists and the journalist Judith Hooper, but later researchers have upheld Kettlewell's findings.
Kettlewell's experiment was a biological experiment in the mid-1950s to study the evolutionary mechanism of industrial melanism in the peppered moth. It was executed by Bernard Kettlewell, working as a research fellow in the Department of Zoology, University of Oxford. He was investigating the cause of the appearance of dark-coloured moth since Industrial Revolution in England in the 19th century. He conducted his first experiment in 1953 in the polluted woodland of Birmingham, and his second experiment in 1955 in Birmingham as well as in the clean woods of Dorset.
Judith Hooper is an American journalist.
Helen Spurway Haldane was a British biologist and the second wife of J. B. S. Haldane. She emigrated to India in 1957 along with Haldane and conducted research in field biology along with Krishna Dronamraju, Suresh Jayakar, and others.
Endless Forms Most Beautiful: The New Science of Evo Devo and the Making of the Animal Kingdom is a 2005 book by the molecular biologist Sean B. Carroll. It presents a summary of the emerging field of evolutionary developmental biology and the role of toolkit genes. It has won numerous awards for science communication.
Animal coloration provided important early evidence for evolution by natural selection, at a time when little direct evidence was available. Three major functions of coloration were discovered in the second half of the 19th century, and subsequently used as evidence of selection: camouflage ; mimicry, both Batesian and Müllerian; and aposematism.
Dazzled and Deceived: Mimicry and Camouflage is a 2009 book on camouflage and mimicry, in nature and military usage, by the science writer and journalist Peter Forbes. It covers the history of these topics from the 19th century onwards, describing the discoveries of Henry Walter Bates, Alfred Russel Wallace and Fritz Müller, especially their studies of butterflies in the Amazon. The narrative also covers 20th-century military camouflage, begun by the painter Abbot Thayer who advocated disruptive coloration and countershading and continued in the First World War by the zoologist John Graham Kerr and the marine artist Norman Wilkinson, who developed dazzle camouflage. In the Second World War, the leading expert was Hugh Cott, who advised the British army on camouflage in the Western Desert.
