The obturator externus groove is the groove on the posterior neck of the femur for the insertion of the obturator externus muscle, a muscle that is important during bipedal locomotion. It is located within the depression of the trochanteric fossa.
This landmark is used as evidence of bipedal locomotion in the hominins. [1] The fossil Orrorin tugenensis (6-7 mya) possesses the obturator externus groove, which suggests that it moved bipedally and could represent one of the earliest fossils with evidence of bipedal locomotion. [2]
Bipedalism is a form of terrestrial locomotion where a tetrapod moves by means of its two rear limbs or legs. An animal or machine that usually moves in a bipedal manner is known as a biped, meaning 'two feet'. Types of bipedal movement include walking or running and hopping.
Homininae, also called "African hominids" or "African apes", is a subfamily of Hominidae. It includes two tribes, with their extant as well as extinct species: 1) the tribe Hominini ―and 2) the tribe Gorillini (gorillas). Alternatively, the genus Pan is sometimes considered to belong to its own third tribe, Panini. Homininae comprises all hominids that arose after orangutans split from the line of great apes. The Homininae cladogram has three main branches, which lead to gorillas, and to humans and chimpanzees via the tribe Hominini and subtribes Hominina and Panina. There are two living species of Panina and two living species of gorillas, but only one extant human species. Traces of extinct Homo species, including Homo floresiensis have been found with dates as recent as 40,000 years ago. Organisms in this subfamily are described as hominine or hominines.
Orrorin is an extinct genus of primate within Homininae from the Miocene Lukeino Formation and Pliocene Mabaget Formation, both of Kenya.
Australopithecus is a genus of early hominins that existed in Africa during the Pliocene and Early Pleistocene. The genera Homo, Paranthropus, and Kenyanthropus evolved from some Australopithecus species. Australopithecus is a member of the subtribe Australopithecina, which sometimes also includes Ardipithecus, though the term "australopithecine" is sometimes used to refer only to members of Australopithecus. Species include A. garhi, A. africanus, A. sediba, A. afarensis, A. anamensis, A. bahrelghazali and A. deyiremeda. Debate exists as to whether some Australopithecus species should be reclassified into new genera, or if Paranthropus and Kenyanthropus are synonymous with Australopithecus, in part because of the taxonomic inconsistency.
Sahelanthropus tchadensis is an extinct species of the hominid dated to about 7 million years ago, during the Miocene epoch. The species, and its genus Sahelanthropus, was announced in 2002, based mainly on a partial cranium, nicknamed Toumaï, discovered in northern Chad.
In anatomy, the thigh is the area between the hip (pelvis) and the knee. Anatomically, it is part of the lower limb.
The piriformis muscle is a flat, pyramidally-shaped muscle in the gluteal region of the lower limbs. It is one of the six muscles in the lateral rotator group.
The internal obturator muscle or obturator internus muscle originates on the medial surface of the obturator membrane, the ischium near the membrane, and the rim of the pubis.
The Hominini form a taxonomic tribe of the subfamily Homininae ("hominines"). Hominini includes the extant genera Homo (humans) and Pan and in standard usage excludes the genus Gorilla (gorillas).
Orthograde is a term derived from Greek ὀρθός, orthos + Latin gradi that describes a manner of walking which is upright, with the independent motion of limbs. Both New and Old World monkeys are primarily arboreal, and they have a tendency to walk with their limbs swinging in parallel to one another. This differs from the manner of walking demonstrated by the apes.
The external obturator muscle, obturator externus muscle is a flat, triangular muscle, which covers the outer surface of the anterior wall of the pelvis.
The adductor brevis is a muscle in the thigh situated immediately deep to the pectineus and adductor longus. It belongs to the adductor muscle group. The main function of the adductor brevis is to pull the thigh medially. The adductor brevis and the rest of the adductor muscle group is also used to stabilize left to right movements of the trunk, when standing on both feet, or to balance when standing on a moving surface. The adductor muscle group is used pressing the thighs together to ride a horse, and kicking with the inside of the foot in soccer or swimming. Last, they contribute to flexion of the thigh when running or against resistance.
In vertebrates, the pubis or pubic bone forms the lower and anterior part of each side of the hip bone. The pubis is the most forward-facing of the three bones that make up the hip bone. The left and right pubic bones are each made up of three sections, a superior ramus, inferior ramus, and a body.
The obturator artery is a branch of the internal iliac artery that passes antero-inferiorly on the lateral wall of the pelvis, to the upper part of the obturator foramen, and, escaping from the pelvic cavity through the obturator canal, it divides into an anterior branch and a posterior branch.
In mammals including humans, the medial surface of the greater trochanter has at its base a deep depression bounded posteriorly by the intertrochanteric crest, called the trochanteric fossa. This fossa is the point of insertion of four muscles. Moving from the inferior-most to the superior-most, they are: the tendon of the obturator externus muscle, the obturator internus, the superior gemellus and inferior gemellus. The width and depth of the trochanteric fossa varies taxonomically.
Poposaurus is an extinct genus of pseudosuchian archosaur from the Late Triassic of the southwestern United States. It belongs to the clade Poposauroidea, an unusual group of Triassic pseudosuchians that includes sail-backed, beaked, and aquatic forms. Fossils have been found in Wyoming, Utah, Arizona, and Texas. Except for the skull, most parts of the skeleton are known. The type species, P. gracilis, was described and named by Maurice Goldsmith Mehl in 1915. A second species, P. langstoni, was originally the type species of the genus Lythrosuchus. Since it was first described, Poposaurus has been variously classified as a dinosaur, a phytosaur, and a "rauisuchian".
The evolution of human bipedalism, which began in primates approximately four million years ago, or as early as seven million years ago with Sahelanthropus, or approximately twelve million years ago with Danuvius guggenmosi, has led to morphological alterations to the human skeleton including changes to the arrangement, shape, and size of the bones of the foot, hip, knee, leg, and the vertebral column. These changes allowed for the upright gait to be overall more energy efficient in comparison to quadrupeds. The evolutionary factors that produced these changes have been the subject of several theories that correspond with environmental changes on a global scale.
William L. Jungers was an American anthropologist, Distinguished Teaching Professor and the Chair of the Department of Anatomical Sciences at State University of New York at Stony Brook on Long Island, New York. He is best known for his work on the biomechanics of bipedal locomotion in hominids such as the 3.4-million-year-old Lucy, and the 6.1- to 5.8-million-year-old Millennium Man Orrorin tugenensis. He devoted much of his career to the study of the lemurs of Madagascar, especially giant extinct subfossil forms such as Megaladapis. More recently, Jungers has been a subject of media attention due to his analysis of the remains of Homo floresiensis, which he believed to be legitimate members of a newly discovered species based on remains of the shoulder, the wrist, and the feet.
Vertical clinging and leaping (VCL) is a type of arboreal locomotion seen most commonly among the strepsirrhine primates and haplorrhine tarsiers. The animal begins at rest with its torso upright and elbows fixed, with both hands clinging to a vertical support, such as the side of a tree or bamboo stalk. To move from one support to another, it pushes off from one vertical support with its hindlimbs, landing on another vertical support after an extended period of free flight. Vertical clinging and leaping primates have evolved a specialized anatomy to compensate for the physical implications of this form of locomotion. These key morphological specializations have been identified in prosimian fossils from as early as the Eocene.
The Lukeino Formation is a geologic formation located in Kenya and it dates to the Late Miocene (Messinian).