In mycology (the branch of biology that includes the study of mushrooms and other fungi), the pileus is the cap or cap-like part of a basidiocarp or ascocarp (fungal fruiting body) that supports a spore-bearing surface, the hymenium. [1] The hymenium (hymenophore) may consist of lamellae, tubes, or teeth, on the underside of the pileus. A pileus is characteristic of agarics, boletes, some polypores, tooth fungi, and some ascomycetes.
The word pileus comes from the Latin for a type of felt cap.
Pilei can be formed in various shapes, and the shapes can change over the course of the developmental cycle of a fungus. The most familiar pileus shape is hemispherical or convex. Convex pilei often continue to expand as they mature until they become flat. Many well-known species have a convex pileus, including the button mushroom, various Amanita species and boletes.
Some, such as the parasol mushroom, have distinct bosses or umbos and are described as umbonate . An umbo is a knobby protrusion at the center of the cap. Some fungi, such as chanterelles have a funnel- or trumpet-shaped appearance. In these cases the pileus is termed infundibuliform.