Rhizodermis is the root epidermis (also referred to as epiblem), the outermost primary cell layer of the root.
Specialized rhisodermal cells, trichoblasts, form long tubular structures (from 5 to 17 micrometers in diameter and from 80 micrometers to 1.5 millimeters in length) almost perpendicular to the main cell axis – root hairs that absorb water and nutrients. Root hairs of the rhizodermis are always in close contact with soil particles and because of their high surface to volume ratio form an absorbing surface which is much larger than the transpiring surfaces of the plant.
With some species of the family Fabaceae, the rhizodermis participates in the recognition and the uptake of nitrogen-fixing Rhizobia bacteria – the first stage of nodulation leading to formation of root nodules. Rhizodermis plays an important role in nutrient uptake by the plant roots. [1]
In contrast with the epidermis, rhizodermis contains no stomata, and is not covered by cuticle. Its unique feature is the presence of root hairs. Root hair is the outgrowth of a single rhizodermal cell. They occur in high frequency in the adsorptive zone of the root. Root hair derives from a trichoblast as a result of an unequal division. It contains a large vacuole; its cytoplasm and nucleus are superseded to the apical region of the outgrowth. Although it does not divide, its DNA replicates so the nucleus is polyploid. Root hairs live only for few days, and die off in 1–2 days due to mechanical damages.
Plant cells are the cells present in green plants, photosynthetic eukaryotes of the kingdom Plantae. Their distinctive features include primary cell walls containing cellulose, hemicelluloses and pectin, the presence of plastids with the capability to perform photosynthesis and store starch, a large vacuole that regulates turgor pressure, the absence of flagella or centrioles, except in the gametes, and a unique method of cell division involving the formation of a cell plate or phragmoplast that separates the new daughter cells.
Skin is the layer of usually soft, flexible outer tissue covering the body of a vertebrate animal, with three main functions: protection, regulation, and sensation.
In vascular plants, the roots are the organs of a plant that are modified to provide anchorage for the plant and take in water and nutrients into the plant body, which allows plants to grow taller and faster. They are most often below the surface of the soil, but roots can also be aerial or aerating, that is, growing up above the ground or especially above water.
A mycorrhiza is a symbiotic association between a fungus and a plant. The term mycorrhiza refers to the role of the fungus in the plant's rhizosphere, its root system. Mycorrhizae play important roles in plant nutrition, soil biology, and soil chemistry.
In cell biology, the meristem is a type of tissue found in plants. It consists of undifferentiated cells capable of cell division. Cells in the meristem can develop into all the other tissues and organs that occur in plants. These cells continue to divide until they become differentiated and lose the ability to divide.
Trichomes are fine outgrowths or appendages on plants, algae, lichens, and certain protists. They are of diverse structure and function. Examples are hairs, glandular hairs, scales, and papillae. A covering of any kind of hair on a plant is an indumentum, and the surface bearing them is said to be pubescent.
Plant nutrition is the study of the chemical elements and compounds necessary for plant growth and reproduction, plant metabolism and their external supply. In its absence the plant is unable to complete a normal life cycle, or that the element is part of some essential plant constituent or metabolite. This is in accordance with Justus von Liebig's law of the minimum. The total essential plant nutrients include seventeen different elements: carbon, oxygen and hydrogen which are absorbed from the air, whereas other nutrients including nitrogen are typically obtained from the soil.
Root nodules are found on the roots of plants, primarily legumes, that form a symbiosis with nitrogen-fixing bacteria. Under nitrogen-limiting conditions, capable plants form a symbiotic relationship with a host-specific strain of bacteria known as rhizobia. This process has evolved multiple times within the legumes, as well as in other species found within the Rosid clade. Legume crops include beans, peas, and soybeans.
An arbuscular mycorrhiza (AM) is a type of mycorrhiza in which the symbiont fungus penetrates the cortical cells of the roots of a vascular plant forming arbuscules. Arbuscular mycorrhiza is a type of endomycorrhiza along with ericoid mycorrhiza and orchid mycorrhiza. They are characterized by the formation of unique tree-like structures, the arbuscules. In addition, globular storage structures called vesicles are often encountered.
Root hair, or absorbent hairs, are outgrowths of epidermal cells, specialized cells at the tip of a plant root. They are lateral extensions of a single cell and are only rarely branched. They are found in the region of maturation, of the root. Root hair cells improve plant water absorption by increasing root surface area to volume ratio which allows the root hair cell to take in more water. The large vacuole inside root hair cells makes this intake much more efficient. Root hairs are also important for nutrient uptake as they are main interface between plants and mycorrhizal fungi.
The rhizosphere is the narrow region of soil or substrate that is directly influenced by root secretions and associated soil microorganisms known as the root microbiome. Soil pores in the rhizosphere can contain many bacteria and other microorganisms that feed on sloughed-off plant cells, termed rhizodeposition, and the proteins and sugars released by roots, termed root exudates. This symbiosis leads to more complex interactions, influencing plant growth and competition for resources. Much of the nutrient cycling and disease suppression by antibiotics required by plants occurs immediately adjacent to roots due to root exudates and metabolic products of symbiotic and pathogenic communities of microorganisms. The rhizosphere also provides space to produce allelochemicals to control neighbours and relatives.
The epidermis is a single layer of cells that covers the leaves, flowers, roots and stems of plants. It forms a boundary between the plant and the external environment. The epidermis serves several functions: it protects against water loss, regulates gas exchange, secretes metabolic compounds, and absorbs water and mineral nutrients. The epidermis of most leaves shows dorsoventral anatomy: the upper (adaxial) and lower (abaxial) surfaces have somewhat different construction and may serve different functions. Woody stems and some other stem structures such as potato tubers produce a secondary covering called the periderm that replaces the epidermis as the protective covering.
A primordium in embryology, is an organ or tissue in its earliest recognizable stage of development. Cells of the primordium are called primordial cells. A primordium is the simplest set of cells capable of triggering growth of the would-be organ and the initial foundation from which an organ is able to grow. In flowering plants, a floral primordium gives rise to a flower.
Lateral roots, emerging from the pericycle, extend horizontally from the primary root (radicle) and over time makeup the iconic branching pattern of root systems. They contribute to anchoring the plant securely into the soil, increasing water uptake, and facilitate the extraction of nutrients required for the growth and development of the plant. Lateral roots increase the surface area of a plant's root system and can be found in great abundance in several plant species. In some cases, lateral roots have been found to form symbiotic relationships with rhizobia (bacteria) and mycorrhizae (fungi) found in the soil, to further increase surface area and increase nutrient uptake.
Rhizobacteria are root-associated bacteria that can have a detrimental, neutral or beneficial effect on plant growth. The name comes from the Greek rhiza, meaning root. The term usually refers to bacteria that form symbiotic relationships with many plants (mutualism). Rhizobacteria are often referred to as plant growth-promoting rhizobacteria, or PGPRs. The term PGPRs was first used by Joseph W. Kloepper in the late 1970s and has become commonly used in scientific literature.
The Hartig net is the network of inward-growing hyphae, that extends into the plant host root, penetrating between plant cells in the root epidermis and cortex in ectomycorrhizal symbiosis. This network is the internal component of fungal morphology in ectomycorrhizal symbiotic structures formed with host plant roots, in addition to a hyphal mantle or sheath on the root surface, and extramatrical mycelium extending from the mantle into the surrounding soil. The Hartig net is the site of mutualistic resource exchange between the fungus and the host plant. Essential nutrients for plant growth are acquired from the soil by exploration and foraging of the extramatrical mycelium, then transported through the hyphal network across the mantle and into the Hartig net, where they are released by the fungi into the root apoplastic space for uptake by the plant. The hyphae in the Hartig net acquire sugars from the plant root, which are transported to the external mycelium to provide a carbon source to sustain fungal growth.
The exodermis is a physiological barrier that has a role in root function and protection. The exodermis is a membrane of variable permeability responsible for the radial flow of water, ions, and nutrients. It is the outer layer of a plant's cortex. The exodermis serves a double function as it can protect the root from invasion by foreign pathogens and ensures that the plant does not lose too much water through diffusion through the root system and can properly replenish its stores at an appropriate rate.
Biomass partitioning is the process by which plants divide their energy among their leaves, stems, roots, and reproductive parts. These four main components of the plant have important morphological roles: leaves take in CO2 and energy from the sun to create carbon compounds, stems grow above competitors to reach sunlight, roots absorb water and mineral nutrients from the soil while anchoring the plant, and reproductive parts facilitate the continuation of species. Plants partition biomass in response to limits or excesses in resources like sunlight, carbon dioxide, mineral nutrients, and water and growth is regulated by a constant balance between the partitioning of biomass between plant parts. An equilibrium between root and shoot growth occurs because roots need carbon compounds from photosynthesis in the shoot and shoots need nitrogen absorbed from the soil by roots. Allocation of biomass is put towards the limit to growth; a limit below ground will focus biomass to the roots and a limit above ground will favor more growth in the shoot.
Plant root exudates are fluids emitted through the roots of plants. These secretions influence the rhizosphere around the roots to inhibit harmful microbes and promote the growth of self and kin plants.
Seventeen elements or nutrients are essential for plant growth and reproduction. They are carbon (C), hydrogen (H), oxygen (O), nitrogen (N), phosphorus (P), potassium (K), sulfur (S), calcium (Ca), magnesium (Mg), iron (Fe), boron (B), manganese (Mn), copper (Cu), zinc (Zn), molybdenum (Mo), nickel (Ni) and chlorine (Cl). Nutrients required for plants to complete their life cycle are considered essential nutrients. Nutrients that enhance the growth of plants but are not necessary to complete the plant's life cycle are considered non-essential, although some of them, such as silicon (Si), have been shown to improve nutrent availability, hence the use of stinging nettle and horsetail macerations in Biodynamic agriculture. With the exception of carbon, hydrogen and oxygen, which are supplied by carbon dioxide and water, and nitrogen, provided through nitrogen fixation, the nutrients derive originally from the mineral component of the soil. The Law of the Minimum expresses that when the available form of a nutrient is not in enough proportion in the soil solution, then other nutrients cannot be taken up at an optimum rate by a plant. A particular nutrient ratio of the soil solution is thus mandatory for optimizing plant growth, a value which might differ from nutrient ratios calculated from plant composition.