| Shirley poppy | |
|---|---|
 Shirley poppy | |
| Species | Papaver rhoeas |
| Origin | Shirley, London in England. |
| Cultivar group members | single forms semi-double forms |
The Shirley poppy is an annual ornamental cultivar group derived from the European wild field poppy ( Papaver rhoeas ).
The Shirley poppy was created from 1880 onwards by the Reverend William Wilks, vicar of the parish of Shirley in England. Wilks found in a corner of his garden where it adjoined arable fields, a variant of the field poppy that had a narrow white border around the petals. By careful selection and hybridization over many years he obtained a strain of poppies ranging in colour from white and pale lilac to pink and red, and unlike the wild poppies these had no dark blotches at the base of the petals. Further selection has given rise to semi-double and double forms, as well as flowers with a ring of contrasting colour around the edge: the picotee form. [1] [2]
The striking example of deviation from a wild type under artificial selection provided by the Shirley poppy attracted the attention of pioneer geneticists and biometricians. The biometrician Karl Pearson used the Shirley poppy to study his ideas of homotyposis, which he defined as “the quantitative degree of resemblance to be found on the average between the like parts of organisms”. [3] [4] However Pearson's work on Shirley poppy was ridiculed by the pioneer geneticist William Bateson for ignoring the recently discovered analytical methods of Mendelian genetics. Bateson wrote: "Misconception of the nature and significance of intermediates has deprived the work of the biometrical school of scientific value as a contribution to the study of heredity. This is well seen in the case of the colours of the Shirley Poppies, one of the subjects with reference to which copious statistics have been amassed and published". [5] The rancorous debate between genetics and biometrics, in which the Shirley poppy became embroiled, was only resolved through the work of Ronald Fisher who showed that the two schools of thought were actually compatible.
Sir Cedric Morris created another form such as 'Mother of Pearl'. Later 'Cedric Morris', 'Fairy Wings' and 'Angels' Choir' were found These are taller-growing semi-doubles and doubles, as well as 'Shirley Double Mixed'. [1] They can grow to between 18in and 2 ft tall, on lightish soil in sun or light shade. They grow quickly from seeds scattered in April. [1]
The Mayor of Croydon’s has as his staff of office a silver mace, on which the head is decorated with the 'Shirley poppy'. In 1935 there was a public house called ‘The Shirley Poppy’ pub, although it has been converted into a burger outlet. [2]
Gregor Johann MendelOSA was a German-Czech biologist, meteorologist, mathematician, Augustinian friar and abbot of St. Thomas' Abbey in Brno (Brünn), Margraviate of Moravia. Mendel was born in a German-speaking family in the Silesian part of the Austrian Empire and gained posthumous recognition as the founder of the modern science of genetics. Though farmers had known for millennia that crossbreeding of animals and plants could favor certain desirable traits, Mendel's pea plant experiments conducted between 1856 and 1863 established many of the rules of heredity, now referred to as the laws of Mendelian inheritance.
Heredity, also called inheritance or biological inheritance, is the passing on of traits from parents to their offspring; either through asexual reproduction or sexual reproduction, the offspring cells or organisms acquire the genetic information of their parents. Through heredity, variations between individuals can accumulate and cause species to evolve by natural selection. The study of heredity in biology is genetics.
Mendelian inheritance is a type of biological inheritance following the principles originally proposed by Gregor Mendel in 1865 and 1866, re-discovered in 1900 by Hugo de Vries and Carl Correns, and later popularized by William Bateson. These principles were initially controversial. When Mendel's theories were integrated with the Boveri–Sutton chromosome theory of inheritance by Thomas Hunt Morgan in 1915, they became the core of classical genetics. Ronald Fisher combined these ideas with the theory of natural selection in his 1930 book The Genetical Theory of Natural Selection, putting evolution onto a mathematical footing and forming the basis for population genetics within the modern evolutionary synthesis.
The modern synthesis was the early 20th-century synthesis of Charles Darwin's theory of evolution and Gregor Mendel's ideas on heredity into a joint mathematical framework. Julian Huxley coined the term in his 1942 book, Evolution: The Modern Synthesis. The synthesis combined the ideas of natural selection, Mendelian genetics, and population genetics. It also related the broad-scale macroevolution seen by palaeontologists to the small-scale microevolution of local populations.
A poppy is a flowering plant in the subfamily Papaveroideae of the family Papaveraceae. Poppies are herbaceous plants, often grown for their colourful flowers. One species of poppy, Papaver somniferum, is the source of the narcotic drug mixture opium which contains powerful medicinal alkaloids such as morphine and has been used since ancient times as an analgesic and narcotic medicinal and recreational drug. It also produces edible seeds. Following the trench warfare in the poppy fields of Flanders, Belgium during World War I, poppies have become a symbol of remembrance of soldiers who have died during wartime, especially in the UK, Canada, Australia, New Zealand and other Commonwealth realms.
Karl Pearson was an English mathematician and biostatistician. He has been credited with establishing the discipline of mathematical statistics. He founded the world's first university statistics department at University College London in 1911, and contributed significantly to the field of biometrics and meteorology. Pearson was also a proponent of Social Darwinism and eugenics, and his thought is an example of what is today described as scientific racism. Pearson was a protégé and biographer of Sir Francis Galton. He edited and completed both William Kingdon Clifford's Common Sense of the Exact Sciences (1885) and Isaac Todhunter's History of the Theory of Elasticity, Vol. 1 (1886–1893) and Vol. 2 (1893), following their deaths.
Papaver rhoeas, with common names including common poppy, corn poppy, corn rose, field poppy, Flanders poppy, and red poppy, is an annual herbaceous species of flowering plant in the poppy family Papaveraceae. It is a temperate native with a very wide distribution area, from Africa to temperate and tropical Asia and Europe.
William Bateson was an English biologist who was the first person to use the term genetics to describe the study of heredity, and the chief populariser of the ideas of Gregor Mendel following their rediscovery in 1900 by Hugo de Vries and Carl Correns. His 1894 book Materials for the Study of Variation was one of the earliest formulations of the new approach to genetics.
"The Correlation between Relatives on the Supposition of Mendelian Inheritance" is a scientific paper by Ronald Fisher which was published in the Transactions of the Royal Society of Edinburgh in 1918,. In it, Fisher puts forward the "infinitesimal model", a genetics conceptual model showing that continuous variation amongst phenotypic traits could be the result of Mendelian inheritance. The paper also contains the first use of the statistical term variance.
Papaver is a genus of 70–100 species of frost-tolerant annuals, biennials, and perennials native to temperate and cold regions of Eurasia, Africa and North America. It is the type genus of the poppy family, Papaveraceae.
Papaver somniferum, commonly known as the opium poppy or breadseed poppy, is a species of flowering plant in the family Papaveraceae. It is the species of plant from which both opium and poppy seeds are derived and is also a valuable ornamental plant grown in gardens. Its native range was east of the Mediterranean Sea, but now is obscured by ancient introductions and cultivation, being naturalized across much of Europe and Asia.
Walter Frank Raphael Weldon FRS, was an English evolutionary biologist and a founder of biometry. He was the joint founding editor of Biometrika, with Francis Galton and Karl Pearson.
Charles Benedict Davenport was a biologist and eugenicist influential in the American eugenics movement.
Blending inheritance is an obsolete theory in biology from the 19th century. The theory is that the progeny inherits any characteristic as the average of the parents' values of that characteristic. As an example of this, a crossing of a red flower variety with a white variety of the same species would yield pink-flowered offspring.
Mutationism is one of several alternatives to evolution by natural selection that have existed both before and after the publication of Charles Darwin's 1859 book On the Origin of Species. In the theory, mutation was the source of novelty, creating new forms and new species, potentially instantaneously, in sudden jumps. This was envisaged as driving evolution, which was thought to be limited by the supply of mutations.
Papaver nudicaule, the Iceland poppy, is a boreal flowering plant. Native to subpolar regions of Asia and North America, and the mountains of Central Asia as well as temperate China, Iceland poppies are hardy but short-lived perennials, often grown as biennials. They yield large, papery, bowl-shaped, lightly fragrant flowers supported by hairy, 1 foot (30 cm) curved stems among feathery blue-green foliage 1-6 inches long. They were first described by botanists in 1759. The wild species blooms in white or yellow, and is hardy from USDA Zones 3a-10b.
Reverend William Wilks (1843–1923) was a notable British horticulturalist and clergyman. He served as secretary of the Royal Horticultural Society from 1888 to 1919 and as editor of its journal, he was instrumental in getting the work of Gregor Mendel translated and published in English for the first time in 1901. This translation of Mendel's 1865 paper was made by Charles Thomas Druery with the involvement of William Bateson.
Charles Chamberlain Hurst (1870–1947) was an English geneticist.
Muriel Wheldale Onslow was a British biochemist, born in Birmingham, England. She studied the inheritance of flower colour in the common snapdragon Antirrhinum and the biochemistry of anthocyanin pigment molecules. She attended the King Edward VI High School in Birmingham and then matriculated at Newnham College, Cambridge in 1900. At Cambridge she majored in botany. Onslow later worked within Bateson's genetic group and then Frederick Gowland Hopkins biochemical group in Cambridge, providing her with expertise in biochemical genetics for investigating the inheritance and biosynthesis of petal colour in Antirrhinum. She was one of the first women appointed as a lecturer at Cambridge, after moving to the Biochemistry department.
Florence Margaret Durham was a British geneticist at Cambridge in the early 1900s and an advocate of the theory of Mendelian inheritance, at a time when it was still controversial. She was part of an informal school of genetics at Cambridge led by her brother-in-law William Bateson. Her work on the heredity of coat colours in mice and canaries helped to support and extend Mendel's law of heredity. It is also one of the first examples of epistasis.
