Kirengellida

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Kirengellida
Temporal range: Late Cambrian–Mid Ordovician
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Brachiopoda (?)
Order: Kirengellida
Family: Kirengellidae
Rozov 1975

The Kirengellids are a group of problematic Cambrian fossil shells of marine organisms. The shells bear a number of paired muscle scars on the inner surface of the valve.

These fossils have conventionally been regarded as monoplacophoran molluscs, and possibly ancestral to gastropods or cephalopods. [1] They were presumed to be exogastric on the presumption that their larger muscle scars were anterior, [2] [3] but it may be dangerous to compare these scars with molluscan musculature. [4] In any case, they coiled in the opposite direction to Romaniella . [4] However, their calcitic shells, the position of the muscle scars, and putative association with secondary shell elements, make a brachiopod affinity possible, by analogy with the mobergellans: a group of phosphatic shells from the same time period, with a similar set of muscle scars. [4] There is also strong similarity to the contemporary brachiopod group, the Craniopsids. In the case of this diagnosis, a simple lophophore apparatus is postulated to sit between the muscle scars and the edges of the shell. [4] On the other hand, Vendrasco (2012) reaffirmed the interpretation of the kirengellids as molluscs, noting that the Kirengella muscle scar pattern is also similar to what occurs in monoplacophorans. [5] [6] Bouchet et al. (2017) classified Kirengellida as an order within the subclass Tergomya. [7]

Included taxa

After [4] :fig. 5

Related Research Articles

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<span class="mw-page-title-main">Monoplacophora</span> Superclass of molluscs

Monoplacophora, meaning "bearing one plate", is a polyphyletic superclass of molluscs with a cap-like shell inhabiting deep sea environments. Extant representatives were not recognized as such until 1952; previously they were known only from the fossil record, and were thought to have become extinct 375 million years ago.

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Strepsodiscus is an extinct genus of very primitive fossil snail-like molluscs from the early part of the Late Cambrian of North America. The coiled, slightly asymmetrical shells are about 3 cm in height. It is not known whether these are shells of gastropods or monoplacophorans, which are more primitive mollusks.

The Bellerophontida is a taxonomic order of extinct marine mollusks that are found in the fossil record from the Lower Cambrian to the Lower Triassic. They are considered by some experts to be primitive sea snails with primarily symmetrically coiled shells, marine gastropod mollusks.

<span class="mw-page-title-main">Bellerophontidae</span> Extinct family of gastropods

The Bellerophontidae are an extinct family of specialized globose bellerophontids, Paleozoic and early Triassic mollusks of the class Gastropoda.

<span class="mw-page-title-main">Evolution of molluscs</span> The origin and diversification of molluscs through geologic time

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<i>Yochelcionella</i> Extinct genus of molluscs

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<i>Scenella</i> Genus of molluscs (fossil)

Scenella is an extinct genus of fossil invertebrate animal which is generally considered to be a mollusc; at various times it has been suggested that this genus belongs with the gastropods, the monoplacophorans, or the helcionellids, although no firm association with any of these classes has been established. An affinity with the hydrozoa has been considered, although some authors oppose this hypothesis. A gastropod affinity is defended on the basis of six pairs of internal muscle scars, whilst the serially-repeated nature of these scars suggests to other authors a monoplacophoran affinity. However the specimens showing this scarring have not been convincingly shown to belong to the genus Scenella. A similarity to the Ediacaran Ovatoscutum has also been drawn.

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<i>Knightoconus</i> Extinct genus of molluscs

Knightoconus antarcticus is an extinct species of fossil monoplacophoran from the Cambrian Minaret Formation of Antarctica. It is thought to represent an ancestor to the cephalopods. It had a chambered conical shell, but lacked a siphuncle.

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<span class="mw-page-title-main">Acrotretida</span> Extinct order of brachiopods

Acrotretides (Acrotretida) are an extinct order of linguliform brachiopods in the class Lingulata. Acrotretida contains 8 families within the sole superfamily Acrotretoidea. They lived from the Lower Cambrian to the Middle Devonian, rapidly diversifying during the middle Cambrian. In the upper Cambrian, linguliforms reached the apex of their diversity: acrotretides and their relatives the lingulides together comprised nearly 70% of brachiopod genera at this time. Though acrotretides continued to diversify during the Ordovician, their proportional dominance declined, as rhynchonelliforms took on a larger role in brachiopod faunas.

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Mobergella is a millimetric Lower Cambrian shelly fossil of unknown affinity, usually preserved in phosphate and particularly well known from Swedish strata, where it is diagnostic of lowermost Cambrian rocks. Originally interpreted as a monoplacophoran, the circular, cap-shaped shell resembles a hyolith operculum, with concentric rings on its upper surface, and seven pairs of internal muscle scars. It is never found in association with a conch, and its affinity therefore remains undetermined. Nevertheless, its heavy musculature does seem to indicate that it functioned as an operculum.

<span class="mw-page-title-main">Siphonotretida</span> Extinct order of marine lamp shells

Siphonotretida is an extinct order of linguliform brachiopods in the class Lingulata. The order is equivalent to the sole superfamily Siphonotretoidea, itself containing the sole family Siphonotretidae. Siphonotretoids were originally named as a superfamily of Acrotretida, before being raised to their own order.

References

  1. e.g. Yochelson, E. L.; Flower, R. H.; Webers, G. F. (1973), "The bearing of the new Late Cambrian monoplacophoran genus Knightoconus upon the origin of the Cephalopoda", Lethaia, 6 (3): 275–309, doi:10.1111/j.1502-3931.1973.tb01199.x
  2. Yochelson, E.L. (1978). "An alternative approach to the interpretation of the phylogeny of ancient mollusks". Malacologia. 17 (2): 165–191.
  3. Yochelson, E.L; Webers, G.F. (2006). "A restudy of the Late Cambrian Molluscan fauna of Berkey (1898) from Taylors Falls, Minnesota" (PDF). Minnesota Geological Survey Report of Investigations. 64: 60.
  4. 1 2 3 4 5 Dzik, Jerzy (2010). "BRACHIOPOD IDENTITY OF THE ALLEGED MONOPLACOPHORAN ANCESTORS OF CEPHALOPODS" (PDF). Malacologia. 52 (1): 97–113.
  5. Vendrasco, M. J. (2012). "Early evolution of molluscs". In Averkii Fyodorov; Havrila Yakovlev (eds.). Mollusks: Morphology, Behavior and Ecology. Nova Science Publishers. pp. 1–43. ISBN   9781621009870.
  6. Cope, J. C. W.; Ebbestad, J. O. R. (2023). "Tergomyan molluscs from the Early Ordovician of the Llangynog Inlier, South Wales, UK". PalZ. doi: 10.1007/s12542-023-00667-5 .
  7. Bouchet, P.; Rocroi, J.-P.; Hausdorf, B.; Kaim, A.; Kano, Y.; Nützel, A.; Parkhaev, P.; Schrödl, M.; Strong, E.E. (2017). "Revised classification, nomenclator and typification of gastropod and monoplacophoran families". Malacologia. 61 (1–2): 1–526. doi:10.4002/040.061.0201.