Valve (mollusc)

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Loose shell plates or valves of Chiton tuberculatus from the beach drift Chiton tuberculatus plates.jpg
Loose shell plates or valves of Chiton tuberculatus from the beach drift

A valve is each articulating part of the shell of a mollusc or another multi-shelled animal such as brachiopods and some crustaceans. Each part is known as a valve or in the case of chitons, a "plate". Members of two classes of molluscs, the Bivalvia (clams) and the Polyplacophora (chitons), have valves.

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Species within one family of very unusual small sea snails, marine opisthobranch gastropods in the family Juliidae, also have two articulating shells or valves, which resemble those of a bivalve. This exceptional family is commonly known as the bivalved gastropods. Gastropods in general are sometimes called "univalves", because in those that have a shell, the shell is usually in one part.

Chitons

Live individual of the lined chiton Tonicella lineata, head end towards the right Tonicella lineata 1.jpg
Live individual of the lined chiton Tonicella lineata , head end towards the right

The valves of chitons are eight dorsal, articulated shell plates, which are frequently coloured and sculpted. After death the girdle that holds the plates together disintegrates and the plates separate. Thus individual plates can be found washed up in beach drift, as shown in the image at the top of this article.

Bivalves

Paired valves of the bivalve Abra alba Abra alba 2.jpg
Paired valves of the bivalve Abra alba

Bivalve molluscs (e.g. clams and oysters) have a shell which is composed of two separate but articulating parts. Each one of these two parts is known as a "valve".

The two valves are known as the "right valve" and the "left valve"; these are labeled with respect to the anterior end of the bivalve, when the umbones are facing upward. In many, but not all, bivalves, the two valves are more or less symmetrical and thus (other than the hinge line) look like mirror images of one another.

Bivalved gastropods

A live individual of an unidentified species of bivalved gastropod in the family Juliidae Juliidae.png
A live individual of an unidentified species of bivalved gastropod in the family Juliidae

The great majority of shelled gastropods or snails have a shell in one part, hence the older name "univalve". The gastropod operculum, when present, even when it is composed mostly of calcium carbonate, is not considered to be a valve.

In contrast, species within one family of small sea snails, the Juliidae, opisthobranch gastropod molluscs, have a hinged shell which is composed of two parts joined by a ligament. These two parts very closely resemble the two valves of a bivalve. This group of species are often referred to as "bivalved gastropods". These are sacoglossans in several genera including Julia , Berthelinia , Midorigai , Edenttellina , Tamanovalva , and Candinia .

Drawing of the interior of the left valve of the shell of "Julia borbonica" Julia borbonica shell 2.jpg
Drawing of the interior of the left valve of the shell of " Julia borbonica "

These bivalve gastropods were for a long time known only from fossils and dead material. Because of this, they had been described as being somewhat atypical bivalves. It was not until living individuals were found in the late 20th century that it was understood that these are in fact very unusual gastropods.

Other animals

Valves are present in any animal with two or more shell portions connecting at a hinge. Brachiopods are bivalved animals with two valves stacked vertically (rather than laterally). The dorsal, or brachial valve hosts the brachial system which supports feeding structures, while the rear end of the ventral, or pedicle valve sends out a fleshy holdfast known as a pedicle. Though each valve has its own symmetrical structure, the two valves have different overall forms and are not mirror images of each other. This distinguishes their valves from those of actual bivalves, which form an identical, but mirrored structure as they are derived from the side of the body.

Some crustaceans have a pair of valves which meet at a hinge. These "bivalved crustaceans" include various fossil and living forms, such as ostracods (seed shrimp), conchostracans (clam shrimp), and phyllocarids.

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<span class="mw-page-title-main">Seashell</span> Hard, protective outer layers created by an animal that lives in the sea

A seashell or sea shell, also known simply as a shell, is a hard, protective outer layer usually created by an animal or organism that lives in the sea. The shell is part of the body of the animal. Empty seashells are often found washed up on beaches by beachcombers. The shells are empty because the animal has died and the soft parts have decomposed or been eaten by another animal.

<span class="mw-page-title-main">Bivalvia</span> Class of molluscs

Bivalvia, in previous centuries referred to as the Lamellibranchiata and Pelecypoda, is a class of marine and freshwater molluscs that have laterally compressed bodies enclosed by a shell consisting of two hinged parts. As a group, bivalves have no head and they lack some usual molluscan organs, like the radula and the odontophore. The class includes the clams, oysters, cockles, mussels, scallops, and numerous other families that live in saltwater, as well as a number of families that live in freshwater. The majority are filter feeders. The gills have evolved into ctenidia, specialised organs for feeding and breathing. Most bivalves bury themselves in sediment, where they are relatively safe from predation. Others lie on the sea floor or attach themselves to rocks or other hard surfaces. Some bivalves, such as the scallops and file shells, can swim. The shipworms bore into wood, clay, or stone and live inside these substances.

<span class="mw-page-title-main">Chiton</span> Class (Polyplacophora) of marine molluscs

Chitons are marine molluscs of varying size in the class Polyplacophora, formerly known as Amphineura. About 940 extant and 430 fossil species are recognized.

<span class="mw-page-title-main">Clam shrimp</span> Suborder of arthropods

Clam shrimp are a group of bivalved branchiopod crustaceans that resemble the unrelated bivalved molluscs. They are extant and also known from the fossil record, from at least the Devonian period and perhaps before. They were originally classified in the former order Conchostraca, which later proved to be paraphyletic, due to the fact that water fleas are nested within clam shrimps. Clam shrimp are now divided into three orders, Cyclestherida, Laevicaudata, and Spinicaudata, in addition to the fossil family Leaiidae.

<span class="mw-page-title-main">Rhynchonellida</span> Order of brachiopods

The taxonomic order Rhynchonellida is one of the two main groups of living articulate brachiopods, the other being the order Terebratulida. They are recognized by their strongly ribbed wedge-shaped or nut-like shells, and the very short hinge line.

<span class="mw-page-title-main">Periostracum</span>

The periostracum is a thin, organic coating that is the outermost layer of the shell of many shelled animals, including molluscs and brachiopods. Among molluscs, it is primarily seen in snails and clams, i.e. in gastropods and bivalves, but it is also found in cephalopods such as Allonautilus scrobiculatus. The periostracum is an integral part of the shell, and it forms as the shell forms, along with the other shell layers. The periostracum is used to protect the organism from corrosion.

A molluscivore is a carnivorous animal that specialises in feeding on molluscs such as gastropods, bivalves, brachiopods and cephalopods. Known molluscivores include numerous predatory molluscs,, arthropods such as crabs and firefly larvae, and, vertebrates such as fish, birds and mammals. Molluscivory is performed in a variety ways with some animals highly adapted to this method of feeding behaviour. A similar behaviour, durophagy, describes the feeding of animals that consume hard-shelled or exoskeleton bearing organisms, such as corals, shelled molluscs, or crabs.

<span class="mw-page-title-main">Bivalve shell</span>

A bivalve shell is part of the body, the exoskeleton or shell, of a bivalve mollusk. In life, the shell of this class of mollusks is composed of two hinged parts or valves. Bivalves are very common in essentially all aquatic locales, including saltwater, brackish water, and freshwater. The shells of bivalves commonly wash up on beaches and along the edges of lakes, rivers, and streams. Bivalves by definition possess two shells or valves, a "right valve" and a "left valve", that are joined by a ligament. The two valves usually articulate with one another using structures known as "teeth" which are situated along the hinge line. In many bivalve shells, the two valves are symmetrical along the hinge line—when truly symmetrical, such an animal is said to be equivalved; if the valves vary from each other in size or shape, inequivalved. If symmetrical front-to-back, the valves are said to be equilateral, and are otherwise considered inequilateral.

<span class="mw-page-title-main">Brachiopod</span> Phylum of marine animals also known as lamp shells

Brachiopods, phylum Brachiopoda, are a phylum of trochozoan animals that have hard "valves" (shells) on the upper and lower surfaces, unlike the left and right arrangement in bivalve molluscs. Brachiopod valves are hinged at the rear end, while the front can be opened for feeding or closed for protection. Two major categories are traditionally recognized, articulate and inarticulate brachiopods. The word "articulate" is used to describe the tooth-and-groove structures of the valve-hinge which is present in the articulate group, and absent from the inarticulate group. This is the leading diagnostic skeletal feature, by which the two main groups can be readily distinguished as fossils. Articulate brachiopods have toothed hinges and simple, vertically-oriented opening and closing muscles. Conversely, inarticulate brachiopods have weak, untoothed hinges and a more complex system of vertical and oblique (diagonal) muscles used to keep the two valves aligned. In many brachiopods, a stalk-like pedicle projects from an opening near the hinge of one of the valves, known as the pedicle or ventral valve. The pedicle, when present, keeps the animal anchored to the seabed but clear of sediment which would obstruct the opening.

<span class="mw-page-title-main">Evolution of brachiopods</span> The origin and diversification of brachiopods through geologic time

The origin of the brachiopods is uncertain; they either arose from reduction of a multi-plated tubular organism, or from the folding of a slug-like organism with a protective shell on either end. Since their Cambrian origin, the phylum rose to a Palaeozoic dominance, but dwindled during the Mesozoic.

<span class="mw-page-title-main">Juliidae</span> Family of gastropods

Juliidae, common name the bivalved gastropods, is a family of minute sea snails, marine gastropod mollusks or micromollusks in the superfamily Oxynooidea, an opisthobranch group.

<span class="mw-page-title-main">Julia (gastropod)</span> Genus of gastropods

Julia is a minute sea snails genus, marine gastropod mollusks or micromollusks in the superfamily Oxynooidea.

<span class="mw-page-title-main">Rhynchonelliformea</span> Subphylum of brachiopods

Rhynchonelliformea is a major subphylum and clade of brachiopods. It is roughly equivalent to the former class Articulata, which was used previously in brachiopod taxonomy up until the 1990s. These so-called articulated brachiopods have many anatomical differences relative to "inarticulate" brachiopods of the subphyla Linguliformea and Craniformea. Articulates have hard calcium carbonate shells with tongue-and-groove hinge articulations and separate sets of simple opening and closing muscles.

<span class="mw-page-title-main">Umbo (bivalve)</span>

The umbo is the vaguely defined, often most prominent, highest part of each valve of the shell of a bivalve or univalve mollusc. It usually contains the valve's beak, the oldest point of the valve, and its degree of prominence and position relative to the hinge line are sometimes helpful in distinguishing bivalve taxa. The umbo forms while the animal is a juvenile, and radial growth subsequently proceeds around that area. The umbo is situated above the hinge line. In those bivalves where the umbones do not protrude, as is the case for example in some mussels, the umbones can nonetheless usually be readily identified by examining the concentric growth lines of the shell.

<span class="mw-page-title-main">Hinge teeth</span>

Hinge teeth are part of the anatomical structure of the inner surface of a bivalve shell, i.e. the shell of a bivalve mollusk. Bivalves by definition have two valves, which are joined together by a strong and flexible ligament situated on the hinge line at the dorsal edge of the shell. In life, the shell needs to be able to open slightly to allow the foot and siphons to protrude, and then close again, without the valves moving out of alignment with one another. To make this possible, in most cases the two valves are articulated using an arrangement of structures known as hinge teeth. Like the ligament, the hinge teeth are also situated along the hinge line of the shell, in most cases.

<span class="mw-page-title-main">Prodissoconch</span>

A prodissoconch is an embryonic or larval shell which is present in the larva of a bivalve mollusk. The prodissoconch is often but not always smooth, and has no growth lines. It is sometimes still present and visible in the adult shell, if there has been no erosion of the shell in that area.

<span class="mw-page-title-main">Adductor muscles (bivalve)</span>

The adductor muscles are the main muscular system in bivalve mollusks. In many parts of the world, when people eat scallops, the adductor muscles are the only part of the animal which is eaten. Adductor muscles leave noticeable scars or marks on the interior of the shell's valves. Those marks are often used by scientists who are in the process of identifying empty shells to determine their correct taxonomic placement.

<span class="mw-page-title-main">Paterinata</span> Extinct class of marine lamp shells

Paterinata is an extinct class of linguliform brachiopods which lived from the lower Cambrian ("Tommotian") to the Upper Ordovician (Hirnantian). It contains the single order Paterinida and the subfamily Paterinoidea. Despite being some of the earliest brachiopods to appear in the fossil record, paterinides stayed as a relatively subdued and low-diversity group even as other brachiopods diversified later in the Cambrian and Ordovician. Paterinides are notable for their high degree of convergent evolution with rhynchonelliform (articulate) brachiopods, which have a similar set of muscles and hinge-adjacent structures.

<i>Argyrotheca</i>

Argyrotheca is a genus of very small to minute lampshells. All species share a large pedicel opening, one ridge on the inside of the pedunculate valve, pits in a diamond pattern on the inside of both valves, and without radial ridges that end in tubercles. It occurs in depths between 6 and 1300 m. It is known since the latest Cretaceous.

<span class="mw-page-title-main">Kutorginata</span> Extinct genus of shelled animals

Kutorginates (Kutorginata) are an extinct class of early rhynchonelliform ("articulate") brachiopods. The class contains only a single order, Kutorginida (kutorginides). Kutorginides were among the earliest rhynchonelliforms, restricted to the lower-middle part of the Cambrian Period.

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