List of hornbills

Last updated

Female great hornbill Female Great Hornbill carrying food.jpg
Female great hornbill

Hornbills are birds in the families Bucerotidae and Bucorvidae. There are currently 62 extant species of hornbills recognised by the International Ornithologists' Union, two in Bucorvidae and 60 in Bucerotidae. [1] Many species of fossil hornbills are known from the Miocene onwards; however, their exact number and taxonomy are unsettled due to ongoing discoveries.

Contents

Conventions

IUCN Red List categories
Conservation status
 EX  Extinct (0 species)
 EW  Extinct in the wild (0 species)
 CR  Critically Endangered (3 species)
 EN  Endangered (4 species)
 VU  Vulnerable (18 species)
 NT  Near threatened (5 species)
 LC  Least concern (17 species)

Conservation status codes listed follow the International Union for Conservation of Nature (IUCN) Red List of Threatened Species. Range maps are provided wherever possible; if a range map is not available, a description of the hornbill's range is provided. Ranges are based on the IOC World Bird List for that species unless otherwise noted. Population estimates are of the number of mature individuals and are taken from the IUCN Red List.

This list follows the taxonomic treatment (designation and order of species) and nomenclature (scientific and common names) of version 13.2 of the IOC World Bird List. [1] Where the taxonomy proposed by the IOC World Bird List conflicts with the taxonomy followed by the IUCN [lower-alpha 1] or the 2023 edition of The Clements Checklist of Birds of the World, [3] the disagreement is noted next to the species's common name (for nomenclatural disagreements) or scientific name (for taxonomic disagreements).

Classification

The International Ornithologists' Union (IOU) recognises 62 species of hornbills in across two families and fifteen genera. [1] This list does not include hybrid species, extinct prehistoric species, or putative species not yet accepted by the IOU.

Hornbills

Bucorvidae

Genus Bucorvus Lesson, 1830 – 2 species
Common nameScientific name and subspeciesRangeIUCN status and estimated population
Abyssinian ground hornbill

Bucorvus abyssinicus 177517713.jpg

B. abyssinicus
(Boddaert, 1783)
Africa  VU 


Unknown Decrease2.svg [4]

Southern ground hornbill

Southern Ground Hornbill (Bucorvus leadbeateri) male, Kruger NP (51889201069).jpg

B. leadbeateri
(Vigors, 1825)
Southern and East Africa
Bucorvus leadbeateri distribution map.png
 VU 


Unknown Steady2.svg [5]

Bucerotidae

Genus Tockus Lesson, R. P., 1830 – 10 species
Common nameScientific name and subspeciesRangeIUCN status and estimated population
Tanzanian red-billed hornbill

Tockus ruahae 59157578.jpg

T. ruahae
Kemp and Delport, 2002
Central Tanzania  NE 


Unknown Blue question mark (italic).svg

Western red-billed hornbill

Western red-billed hornbill (Tockus kempi) male.jpg

T. kempi
Tréca & Érard, 2000
West Africa  NE 


Unknown Blue question mark (italic).svg

Damara red-billed hornbill

DSC09456 - NAMIBIA 2013 (31739376283).jpg

T. damarensis
(Shelley, 1888)
Southern Africa  LC 


Unknown Steady2.svg [6]

Southern red-billed hornbill

Southern red-billed hornbill (Tockus rufirostris) male, crop.jpg

T. rufirostris
(Sundevall, 1850)
Southern Africa  NE 


Unknown Blue question mark (italic).svg

Northern red-billed hornbill

Hornbill, Ethiopia (47971177468), crop.jpg

T. erythrorhynchus
(Temminck, 1823)
Africa
Tockus erythrorhynchus - Distribution.png
 NE 


Unknown Blue question mark (italic).svg

Monteiro's hornbill

Tockus monteiri 309485879.jpg

T. monteiri
Hartlaub, 1865
Angola and Namibia
Tockus monteiri - Distribution.png
 LC 


Unknown Steady2.svg [7]

Von der Decken's hornbill

Von der Decken's Hornbill, Tarangire NP (2015).jpg

T. deckeni
(Cabanis, 1868)
East Africa  LC 


Unknown Decrease2.svg [8]

Jackson's hornbill

Tockus jacksoni 136201865.jpg

T. jacksoni
(Ogilvie-Grant, 1891)
East Africa  LC 


Unknown Decrease2.svg [9]

Southern yellow-billed hornbill

Southern Yellow-billed Hornbill, Tockus leucomelas at Mapungubwe National Park, Limpopo, South Africa (18305273221).jpg

T. leucomelas
(Lichtenstein, M. H. C., 1842)

Two subspecies
  • T. l. elegans
  • T. l. leucomelas
Southern Africa
Tockus leucomelas - Distribution.png
 LC 


Unknown Decrease2.svg [10]

Eastern yellow-billed hornbill

Eastern Yellow-billed Hornbill - Shaba Kenya.jpg

T. flavirostris
(Rüppell, 1835)
East Africa
Tockus flavirostris - Distribution.png
 LC 


Unknown Decrease2.svg [11]

Genus Lophoceros Hemprich and Ehrenberg, 1833 – 8 species
Common nameScientific name and subspeciesRangeIUCN status and estimated population
Bradfield's hornbill

Lophoceros bradfieldi 312096352.jpg

L. bradfieldi
(Roberts, 1930)
Southern Africa
Tockus bradfieldi - Distribution.png
 LC 


Unknown Decrease2.svg [12]

Crowned hornbill

Crowned Hornbill - Kenya S4E8074 (16382513056).jpg

L. alboterminatus
Büttikofer, 1889
Africa
Tockus alboterminatus - Distribution.png
 LC 


Unknown Decrease2.svg [13]

West African pied hornbill

African pied hornbill (Tockus fasciatus semifasciatus) male.jpg

L. semifasciatus
(Hartlaub, 1855)
West Africa  LC 


Unknown Blue question mark (italic).svg [14]

Congo pied hornbill L. fasciatus
(Shaw, 1812)
Central Africa  LC 


Unknown Blue question mark (italic).svg [15]

Hemprich's hornbill

Hemprich's hornbill, Lake Langano, Ethiopia (51601917326).jpg

L. hemprichii
(Ehrenberg, 1833)
East Africa
Tockus hemprichii - Distribution.png
 LC 


Unknown Decrease2.svg [16]

African grey hornbill

African grey hornbill (Tockus nasutus nasutus) male.jpg

L. nasutus
(Linnaeus, 1766)

Two subspecies
  • L. n. nasutus
  • L. n. epirhinus
Africa
Tockus nasutus - Distribution, subspecies.png
 LC 


Unknown Steady2.svg [17]

Red-billed dwarf hornbill

A monograph of the Bucerotidae, or family of the hornbills (Plate LIX) (6944171846).jpg

L. camurus
(Cassin, 1857)
East Africa  LC 


Unknown Decrease2.svg [18]

Pale-billed hornbill

Tockus pallidirostris subsp pallidirostris, oog van Cuitorivier, Birding Weto, a.jpg

L. pallidirostris
(Hartlaub and Finsch, 1870)

Two subspecies
  • L. p. pallidirostris
  • L. p. neumanni
Southern Africa
Tockus pallidirostris - Distribution.png
 LC 


Unknown Decrease2.svg [19]

Genus Bycanistes Cabanis and Heine, 1860 – 6 species
Common nameScientific name and subspeciesRangeIUCN status and estimated population
Piping hornbill

Piping Hornbill RWD1.jpg

B. fistulator
(Cassin, 1850)

Three subspecies
  • B. f. fistulator
  • B. f. sharpii
  • B. f. duboisi
West and Central Africa  LC 


Unknown Decrease2.svg [20]

Trumpeter hornbill

Trumpeter Hornbill (Bycanistes bucinator) male captive ... (32861014451).jpg

B. bucinator
(Temminck, 1824)
Southern and East Africa  LC 


Unknown Decrease2.svg [21]

Brown-cheeked hornbill

Kalao-trubach.jpg

B. cylindricus
(Temminck, 1831)
West Africa  VU 


Unknown Decrease2.svg [22]

White-thighed hornbill

A monograph of the Bucerotidae, or family of the hornbills (Plate XXXI) (7090229887) (cropped).jpg

B. albotibialis
(Cassin and Reichenow, 1877)
Central Africa  LC 


Unknown Decrease2.svg [23]

Black-and-white-casqued hornbill

Black-and-White Casqued Hornbill - Kakamega Kenya 06 1778 (16407565342).jpg

B. subcylindricus
(Sclater, P. L., 1871)

Two subspecies
  • B. s. subcylindricus
  • B. s. subquadratus
Central and West Africa  LC 


Unknown Blue question mark (italic).svg [24]

Silvery-cheeked hornbill

Silvery-cheeked Hornbill RWD3.jpg

B. brevis
Friedmann, 1929
East Africa  LC 


Unknown Decrease2.svg [25]

Genus Ceratogymna Bonaparte, 1854 – 2 species
Common nameScientific name and subspeciesRangeIUCN status and estimated population
Black-casqued hornbill

Ceratogymna atrata - Jardin d'oiseaux tropicaux - DSC04947.JPG

C. atrata
(Temminck, 1835)
West and Central Africa  VU 


8,000–9,000 Decrease2.svg [26]

Yellow-casqued hornbill

Ceratogymna elata -Hong Kong Zoo-8a crop.jpg

C. elata
(Temminck, 1831)
West and Central Africa  LC 


Unknown Decrease2.svg [27]

Genus Horizocerus Oberholser, 1899 – 2 species
Common nameScientific name and subspeciesRangeIUCN status and estimated population
Black dwarf hornbill

A monograph of the Bucerotidae, or family of the hornbills (Plate LVIII) BHL38534751.jpg

H. hartlaubi
(Gould, 1861)

Two subspecies
  • H. h. hartlaubi
  • H. h. granti
Central and West Africa
Tockus hartlaubi - Distribution.png
 LC 


Unknown Decrease2.svg [28]

White-crested hornbill

Closeup Bird (125447445).jpeg

H. albocristatus
(Cassin, 1848)

Three subspecies
  • H. a. albocristatus
  • H. a. macrourus
  • H. a. cassini
Central and West Africa  LC 


Unknown Decrease2.svg [29]

Genus Berenicornis Bonaparte, 1850 – 1 species
Common nameScientific name and subspeciesRangeIUCN status and estimated population
White-crowned hornbill

White-crowned Hornbill (14839739067).jpg

B. comatus
(Raffles, 1822)
Southeast Asia  EN 


Unknown Decrease2.svg [30]

Genus Buceros Linnaeus, 1758 – 3 species
Common nameScientific name and subspeciesRangeIUCN status and estimated population
Rhinoceros hornbill

Buceros rhinoceros rhinoceros 215483344.jpg

B. rhinoceros
Linnaeus, 1758

Three subspecies
  • B. r. borneoensis
  • B. r. rhinoceros
  • B. r. silvestris
Southeast Asia  VU 


Unknown Decrease2.svg [31]

Great hornbill

Great hornbill Photograph by Shantanu Kuveskar.jpg

B. bicornis
Linnaeus, 1758
South and Southeast Asia  VU 


13,000–27,000 Decrease2.svg [32]

Rufous hornbill

Buceros hydrocorax - Weltvogelpark Walsrode 2011-07.jpg

B. hydrocorax
Linnaeus, 1766

Three subspecies
  • B. h. hydrocorax
  • B. h. semigaleatus
  • B. h. mindanensis
Philippines  VU 


Unknown Decrease2.svg [33] [34]

Genus Rhinoplax Gloger, 1841 – 1 species
Common nameScientific name and subspeciesRangeIUCN status and estimated population
Helmeted hornbill

Helmeted Hornbill.jpg

R. vigil
(Pennant, 1781)
Southeast Asia  CR 


Unknown Decrease2.svg [35]

Genus Anthracoceros Reichenbach, 1849 – 5 species
Common nameScientific name and subspeciesRangeIUCN status and estimated population
Palawan hornbill

Palawan Hornbill.jpg

A. marchei
Oustalet, 1885
Palawan and surrounding islands  VU 


2,500–9,999 Decrease2.svg [36]

Oriental pied hornbill

Anthracoceros albirostris (Brunei) 2023-08-18.jpg

A. albirostris
(Shaw, 1808)

Two subspecies
  • A. a. albirostris
  • A. a. convexus
South and Southeast Asia  LC 


Unknown Steady2.svg [37]

Malabar pied hornbill

Malabar pied hornbill (Anthracoceros coronatus) male.jpg

A. coronatus
(Boddaert, 1783)
India and Sri Lanka  NT 


Unknown Decrease2.svg [38]

Sulu hornbill

A monograph of the Bucerotidae, or family of the hornbills (Plate XXV) BHL38534609.jpg

A. montani
(Oustalet, 1880)
Sulu Archipelago  CR 


1–49 Decrease2.svg [39]

Black hornbill

Anthracoceros malayanus 260745794 (cropped).jpg

A. malayanus
(Raffles, 1822)
Southeast Asia  VU 


Unknown Decrease2.svg [40]

Genus Ocyceros Hume, 1873 – 3 species
Common nameScientific name and subspeciesRangeIUCN status and estimated population
Malabar grey hornbill

Hello hornbills (51757754297).jpg

O. griseus
(Latham, 1790)
Southwestern India  VU 


Unknown Decrease2.svg [41]

Sri Lanka grey hornbill

Ocyceros gingalensis 31966300.jpg

O. gingalensis
(Shaw, 1812)
Sri Lanka  LC 


Unknown Steady2.svg [42]

Indian grey hornbill

Grey hornbill on tree.jpg

O. birostris
(Scopoli, 1786)
India and Nepal  LC 


Unknown Steady2.svg [43]

Genus Anorrhinus Reichenbach, 1849 – 3 species
Common nameScientific name and subspeciesRangeIUCN status and estimated population
Tickell's brown hornbill

Tickell's brown hornbill (33419252811).jpg

A. tickelli
(Blyth, 1855)
Myanmar and Thailand  NT 


2,500–9,999 Decrease2.svg [44]

Austen's brown hornbill

Anorrhinus austeni 266231698.jpg

A. austeni
Jerdon, 1872
Indochina  NT 


Unknown Decrease2.svg [45]

Bushy-crested hornbill

Anorrhinus galeritus 260745728.jpg

A. galeritus
(Temminck, 1831)
Southeast India  NT 


Unknown Decrease2.svg [46]

Genus Aceros Hodgson, 1844 – 1 species
Common nameScientific name and subspeciesRangeIUCN status and estimated population
Rufous-necked hornbill

Rufous necked hornbill0462-1.jpg

A. nipalensis
(Vigors, 1825)
Northwest India and Indochina  VU 


7,000–10,000 Decrease2.svg [47]

Genus Rhyticeros Reichenbach, 1849 – 6 species
Common nameScientific name and subspeciesRangeIUCN status and estimated population
Blyth's hornbill

Rhyticeros plicatus -Lincoln Park Zoo-8a-3c.jpg

R. plicatus
(Pennant, 1781)
Moluccas and Oceania  VU 


Unknown Decrease2.svg [48]

Narcondam hornbill

Narcondam Hornbill DSCN1992 03.jpg

R. narcondami
Hume, 1873
Narcondam Island  VU 


300–650 Steady2.svg [49]

Wreathed hornbill

Wreathed hornbill1.jpg

R. undulatus
(Shaw, 1811)
South and Southeast Asia  VU 


Unknown Decrease2.svg [50]

Sumba hornbill

Stavenn Sumba Hornbill Wiki.jpg

R. everetti
Rothschild, 1898
Sumba Island  VU 


1,000–2,000 Decrease2.svg [51]

Plain-pouched hornbill

A monograph of the Bucerotidae, or family of the hornbills (Plate XXXVI) (7090232911).jpg

R. subruficollis
(Blyth, 1843)
Southeast Asia  VU 


1,500–7,000 Decrease2.svg [52]

Knobbed hornbill

Helmhornvogel O (258326127).jpeg

R. cassidix
(Temminck, 1823)
Sulawesi and surrounding islands  VU 


Unknown Decrease2.svg [53]

Genus Rhabdotorrhinus Meyer, A. B. and Wiglesworth, 1898 – 4 species
Common nameScientific name and subspeciesRangeIUCN status and estimated population
Walden's hornbill

Ee writhead hornbill zebu zoo.jpg

R. waldeni
(Sharpe, 1877)
Panay and Negros  CR 


1,000–2,499 Decrease2.svg [54]

Writhed hornbill

Aceros leucocephalus (Mindanaohornvogel - Writhed Hornbill) - Weltvogelpark Walsrode 2013-06.jpg

R. leucocephalus
(Vieillot, 1816)
Mindanao  NT 


Unknown Decrease2.svg [55]

Sulawesi hornbill

Zoborozec celebesky (Penelopides exarhatus) - ZOO Plzen 01.jpg

R. exarhatus
(Temminck, 1823)

Two subspecies
  • R. e. exarhatus
  • R. e. sanfordi
Sulawesi  VU 


Unknown Decrease2.svg [56]

Wrinkled hornbill

Wrinkled hornbill (Rhabdotorrhinus corrugatus).jpg

R. corrugatus
(Temminck, 1832)
Southeast Asia  EN 


Unknown Decrease2.svg [57]

Genus Penelopides Reichenbach, 1849 – 5 species
Common nameScientific name and subspeciesRangeIUCN status and estimated population
Luzon hornbill

A male Luzon Hornbill gazes out the forest in the Philippines.jpg

P. manillae
(Boddaert, 1783)

Two subspecies
  • P. m. manillae
  • P. m. sanfordi
Luzon and surrounding islands  LC 


Unknown Decrease2.svg [58]

Mindoro hornbill

MIndoro Hornbill (male) -1.jpg

P. mindorensis
Steere, 1890
Mindoro  EN 


250–999 Decrease2.svg [59]

Mindanao hornbill

A monograph of the Bucerotidae, or family of the hornbills (Plate XXII) (7090226895) (cropped).jpg

P. affinis
Tweeddale, 1877

Two subspecies
  • P. a. affinis
  • P. a. basilanicus
Mindanao and surrounding islands  LC 


Unknown Decrease2.svg [60]

Samar hornbill P. corrugatus
Steere, 1890
Eastern Visayas  NE 


Unknown Blue question mark (italic).svg

Visayan hornbill

Visayan Tarictic Hornbill.jpg

P. panini
(Boddaert, 1783)

Two subspecies
  • P. p. ticaensis
  • P. p. panini
Western Visayas  EN 


1,200 Decrease2.svg [61]

Notes

  1. The IUCN follows the taxonomy proposed by the HBW and BirdLife Taxonomic Checklist. [2]

Related Research Articles

This article is a list of biological species, subspecies, and evolutionary significant units that are known to have become extinct during the Holocene, the current geologic epoch, ordered by their known or approximate date of disappearance from oldest to most recent.

<span class="mw-page-title-main">Red-billed dwarf hornbill</span> Species of bird

The red-billed dwarf hornbill is a species of hornbill in the family Bucerotidae. It is distributed across the African tropical rainforest.

<span class="mw-page-title-main">Pale-billed hornbill</span> Species of bird

The pale-billed hornbill is a species of hornbill in the family Bucerotidae. It is found in Angola, DRC, Kenya, Malawi, Mozambique, Tanzania, and Zambia.

References

  1. 1 2 3 Gill, F.; Donsker, D.; Rasmussen, P., eds. (July 2023). "Mousebirds, cuckoo roller, trogons, hoopoes, hornbills". IOC World Bird List. v 13.2. Retrieved 7 September 2023.
  2. "Handbook of the Birds of the World and BirdLife International digital checklist of the birds of the world. Version 7". HBW and BirdLife International. 2022. Archived from the original on 25 September 2019. Retrieved 7 September 2023.
  3. Clements, James F.; Schulenberg, T. S.; Iliff, M. J.; Fredericks, T. A.; Gerbracht, J. A.; Lepage, Denis; Billerman, S. M.; Sullivan, B. L.; Wood, C. L. (2022). "The eBird/Clements checklist of Birds of the World: v2022". Clements Checklist. Retrieved 7 September 2023.
  4. BirdLife International (2018). "Bucorvus abyssinicus". IUCN Red List of Threatened Species . 2018: e.T22682632A132204438. doi: 10.2305/IUCN.UK.2018-2.RLTS.T22682632A132204438.en . Retrieved 19 November 2021.
  5. BirdLife International (2016). "Bucorvus leadbeateri". IUCN Red List of Threatened Species . 2016: e.T22682638A92955067. doi: 10.2305/IUCN.UK.2016-3.RLTS.T22682638A92955067.en . Retrieved 12 November 2021.
  6. BirdLife International (2016). "Tockus damarensis". IUCN Red List of Threatened Species . 2016: e.T22725940A94906853. doi: 10.2305/IUCN.UK.2016-3.RLTS.T22725940A94906853.en . Retrieved 7 September 2023.
  7. BirdLife International (2016). "Tockus monteiri". IUCN Red List of Threatened Species . 2016: e.T22682367A92942465. doi: 10.2305/IUCN.UK.2016-3.RLTS.T22682367A92942465.en . Retrieved 12 November 2021.
  8. BirdLife International (2016). "Tockus deckeni". IUCN Red List of Threatened Species . 2016: e.T22682392A92943252. doi: 10.2305/IUCN.UK.2016-3.RLTS.T22682392A92943252.en . Retrieved 12 November 2021.
  9. BirdLife International (2016). "Tockus jacksoni". IUCN Red List of Threatened Species . 2016: e.T22682388A92943069. doi: 10.2305/IUCN.UK.2016-3.RLTS.T22682388A92943069.en . Retrieved 12 November 2021.
  10. BirdLife International (2016). "Tockus leucomelas". IUCN Red List of Threatened Species . 2016: e.T22682384A92942849. doi: 10.2305/IUCN.UK.2016-3.RLTS.T22682384A92942849.en . Retrieved 12 November 2021.
  11. BirdLife International (2016). "Tockus flavirostris". IUCN Red List of Threatened Species . 2016: e.T22682379A92942633. doi: 10.2305/IUCN.UK.2016-3.RLTS.T22682379A92942633.en . Retrieved 13 November 2021.
  12. BirdLife International (2016). "Lophoceros bradfieldi". IUCN Red List of Threatened Species . 2016: e.T22682400A92943713. doi: 10.2305/IUCN.UK.2016-3.RLTS.T22682400A92943713.en . Retrieved 11 November 2021.
  13. BirdLife International (2016). "Lophoceros alboterminatus". IUCN Red List of Threatened Species . 2016: e.T22682396A92943440. doi: 10.2305/IUCN.UK.2016-3.RLTS.T22682396A92943440.en . Retrieved 12 November 2021.
  14. BirdLife International (2017). "Lophoceros semifasciatus". IUCN Red List of Threatened Species . 2017: e.T61612971A118859362. doi: 10.2305/IUCN.UK.2017-3.RLTS.T61612971A118859362.en . Retrieved 16 July 2023.
  15. BirdLife International (2016). "Lophoceros fasciatus". IUCN Red List of Threatened Species . 2016: e.T61612961A95171708. doi: 10.2305/IUCN.UK.2016-3.RLTS.T61612961A95171708.en . Retrieved 12 November 2021.
  16. BirdLife International (2018). "Lophoceros hemprichii". IUCN Red List of Threatened Species . 2018: e.T22682409A130080934. doi: 10.2305/IUCN.UK.2018-2.RLTS.T22682409A130080934.en . Retrieved 12 November 2021.
  17. BirdLife International (2016). "Lophoceros nasutus". IUCN Red List of Threatened Species . 2016: e.T22682413A92944101. doi: 10.2305/IUCN.UK.2016-3.RLTS.T22682413A92944101.en . Retrieved 12 November 2021.
  18. BirdLife International (2018). "Lophoceros camurus". IUCN Red List of Threatened Species . 2018: e.T22682363A130080398. doi: 10.2305/IUCN.UK.2018-2.RLTS.T22682363A130080398.en . Retrieved 13 November 2021.
  19. BirdLife International (2016). "Lophoceros pallidirostris". IUCN Red List of Threatened Species . 2016: e.T22682417A92944515. doi: 10.2305/IUCN.UK.2016-3.RLTS.T22682417A92944515.en . Retrieved 11 November 2021.
  20. BirdLife International (2018). "Bycanistes fistulator". IUCN Red List of Threatened Species . 2018: e.T22725962A132234122. doi: 10.2305/IUCN.UK.2018-2.RLTS.T22725962A132234122.en . Retrieved 12 November 2021.
  21. BirdLife International (2018). "Bycanistes bucinator". IUCN Red List of Threatened Species . 2018: e.T22682548A130081383. doi: 10.2305/IUCN.UK.2018-2.RLTS.T22682548A130081383.en . Retrieved 13 November 2021.
  22. BirdLife International (2018). "Bycanistes cylindricus". IUCN Red List of Threatened Species . 2018: e.T22682585A129918692. doi: 10.2305/IUCN.UK.2018-2.RLTS.T22682585A129918692.en . Retrieved 12 November 2021.
  23. BirdLife International (2016). "Bycanistes albotibialis". IUCN Red List of Threatened Species . 2016: e.T22682609A92953893. doi: 10.2305/IUCN.UK.2016-3.RLTS.T22682609A92953893.en . Retrieved 12 November 2021.
  24. BirdLife International (2017). "Bycanistes subcylindricus". IUCN Red List of Threatened Species . 2017: e.T22682577A111897019. doi: 10.2305/IUCN.UK.2017-1.RLTS.T22682577A111897019.en . Retrieved 12 November 2021.
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