| Xiaotingia | |
|---|---|
 | |
| Type specimen | |
Scientific classification  | |
| Domain: | Eukaryota |
| Kingdom: | Animalia |
| Phylum: | Chordata |
| Clade: | Dinosauria |
| Clade: | Saurischia |
| Clade: | Theropoda |
| Family: | † Anchiornithidae |
| Genus: | † Xiaotingia Xu et al., 2011 |
| Type species | |
| Xiaotingia zhengi Xu et al., 2011 | |
Xiaotingia is a genus of anchiornithid theropod dinosaur from Middle Jurassic or early Late Jurassic deposits of western Liaoning, China, containing a single species, Xiaotingia zhengi. [2] [3]
Xiaotingia is known from the holotype STM 27-2, an articulated and almost complete skeleton including the skull. It was probably collected in the Linglongta area, Jianchang, from the Tiaojishan Formation. [2]
Xiaotingia was first named by Xu Xing, You Hailu, Du Kai and Han Fenglu in 2011 and the type species is Xiaotingia zhengi. The generic name and specific name together honour paleontologist Zheng Xiaoting. [2]
Xiaotingia was morphologically similar to other anchiornithids. It was about 60 cm long and weighed an estimated 0.82 kg. It was a small feathered dinosaur that lived in an arboreal environment. Like Archaeopteryx it had long forelimbs. Its femur was longer than its humerus, 84 mm compared to 71 mm, which might indicate that it stood on its hind limbs and could flap its forelimbs to achieve flight. [2]
Xiaotingia had feathers on its head, body, forelimbs and hind limbs. The feathers on the femur were quite long, measuring 55 mm. It also had long pennaceous feathers on its tibia and metatarsus. If Xiaotingia could fly short distances it might also have used its hind limbs as wings. [4]
Xiaotingia had a dentary tooth count probably less than 10 and teeth similar in morphology to those of basal avians. [2]
The initial analysis by Xu et al. showed that Xiaotingia formed a clade with Archaeopteryx , Dromaeosauridae and Troodontidae to the exclusion of other groups traditionally seen as birds. Xu et al. therefore (re)defined the concepts of Deinonychosauria and Avialae to the extent that Archaeopteryx and Xiaotingia belonged to the Deinonychosauria in the clade Archaeopterygidae. [2] This led to popular reports that "Archaeopteryx is no longer a bird", [5] although Xu et al. noted that there are several competing definitions of the clade Aves currently in use, pointing out that their definitions are compatible with a traditional Aves with Archaeopteryx as a specifier. [2] This phylogenetic hypothesis was challenged by an analysis using different methods published several months later however, in which Archaeopteryx was again recovered as an avialan, while Xiaotingia remained closely allied to Anchiornis within the Troodontidae. [6] In 2012, an expanded and revised version of the initial analysis also found Archaeopteryx to be avialan and Anchiornis to be troodontid, but recovered Xiaotingia as the most primitive member of the clade Dromaeosauridae rather than a close relative of Anchiornis within Troodontidae. [7]
Cladogram following the results of a phylogenetic study by Lefèvre et al., 2017. [4]
In the 2017 re-evaluation of the Harlem Archaeopteryx specimen, Xiaotingia was found to be an anchiornithid, with this group being avialan. [3] Hartman et al. (2019), which aimed to improve the state of theropod phylogenetic research, placed both Xiaotingia and Archaeopteryx in Deinonychosauria, the former as a troodontid and the latter a close relative of anchiornithids. [8]
Archaeopteryx, sometimes referred to by its German name, "Urvogel" is a genus of bird-like dinosaurs. The name derives from the ancient Greek ἀρχαῖος (archaīos), meaning "ancient", and πτέρυξ (ptéryx), meaning "feather" or "wing". Between the late 19th century and the early 21st century, Archaeopteryx was generally accepted by palaeontologists and popular reference books as the oldest-known bird. Older potential avialans have since been identified, including Anchiornis, Xiaotingia, and Aurornis.
Troodontidae is a clade of bird-like theropod dinosaurs. During most of the 20th century, troodontid fossils were few and incomplete and they have therefore been allied, at various times, with many dinosaurian lineages. More recent fossil discoveries of complete and articulated specimens, have helped to increase understanding about this group. Anatomical studies, particularly studies of the most primitive troodontids, like Sinovenator, demonstrate striking anatomical similarities with Archaeopteryx and primitive dromaeosaurids, and demonstrate that they are relatives comprising a clade called Paraves.
Deinonychosauria is a clade of paravian dinosaurs which lived from the Late Jurassic to the Late Cretaceous periods. Fossils have been found across the globe in North America, Europe, Africa, Asia, South America, and Antarctica, with fossilized teeth giving credence to the possibility that they inhabited Australia as well. This group of dinosaurs are known for their sickle-shaped toe claws and features in the shoulder bones.
Dromaeosauridae is a family of feathered coelurosaurian theropod dinosaurs. They were generally small to medium-sized feathered carnivores that flourished in the Cretaceous Period. The name Dromaeosauridae means 'running lizards', from Greek δρομαῖος (dromaîos), meaning 'running at full speed', 'swift', and σαῦρος (saûros), meaning 'lizard'. In informal usage, they are often called raptors, a term popularized by the film Jurassic Park; several genera include the term "raptor" directly in their name, and popular culture has come to emphasize their bird-like appearance and speculated bird-like behavior.
Maniraptora is a clade of coelurosaurian dinosaurs which includes the birds and the non-avian dinosaurs that were more closely related to them than to Ornithomimus velox. It contains the major subgroups Avialae, Dromaeosauridae, Troodontidae, Oviraptorosauria, and Therizinosauria. Ornitholestes and the Alvarezsauroidea are also often included. Together with the next closest sister group, the Ornithomimosauria, Maniraptora comprises the more inclusive clade Maniraptoriformes. Maniraptorans first appear in the fossil record during the Jurassic Period, and survive today as living birds.
Scansoriopterygidae is an extinct family of climbing and gliding maniraptoran dinosaurs. Scansoriopterygids are known from five well-preserved fossils, representing four species, unearthed in the Tiaojishan Formation fossil beds of Liaoning and Hebei, China.
Archaeopterygidae is a group of paravian dinosaurs, known from the latest Jurassic and earliest Cretaceous of Europe. In most current classifications, it contains only the genera Archaeopteryx and Wellnhoferia. As its name suggests, Protarchaeopteryx was also once referred to this group, but most paleontologists now consider it an oviraptorosaur. Other referred genera, like Jurapteryx, Wellnhoferia, and "Proornis", are probably synonymous with Archaeopteryx or do not belong into this group. Jinfengopteryx was originally described as an archaeopterygid, though it was later shown to be a troodontid. A few studies have recovered Anchiornis and Xiaotingia to also be members of the Archaeopterygidae, though most subsequent analyses have failed to arrive at the same result. Uncertainties still exist, however, and it may not be possible to confidently state whether archaeopterygids are more closely related to modern birds or to deinonychosaurs barring new and better specimens of relevant species. Teeth attributable to archaeopterygids are known from the earliest Cretaceous (Berriasian) Cherves-de-Cognac locality and the Angeac-Charente bonebed of France.
Maniraptoriformes is a clade of dinosaurs with pennaceous feathers and wings that contains ornithomimosaurs and maniraptorans. This group was named by Thomas Holtz, who defined it as "the most recent common ancestor of Ornithomimus and birds, and all descendants of that common ancestor."
Avialae is a clade containing the only living dinosaurs, the birds. It is usually defined as all theropod dinosaurs more closely related to birds (Aves) than to deinonychosaurs, though alternative definitions are occasionally used.
Paraves are a widespread group of theropod dinosaurs that originated in the Middle Jurassic period. In addition to the extinct dromaeosaurids, troodontids, anchiornithids, and possibly the scansoriopterygids, the group also contains the avialans, which include diverse extinct taxa as well as the over 10,000 species of living birds. Basal members of Paraves are well known for the possession of an enlarged claw on the second digit of the foot, which was held off the ground when walking in some species. A number of differing scientific interpretations of the relationships between paravian taxa exist. New fossil discoveries and analyses make the classification of Paraves an active subject of research.
Epidexipteryx is a genus of small paravian dinosaurs, known from one fossil specimen in the collection of the Institute of Vertebrate Paleontology and Paleoanthropology in Beijing. Epidexipteryx represents the earliest known example of ornamental feathers in the fossil record.
Anchiornis is a genus of small, four-winged paravian dinosaurs, with only one known species, the type species Anchiornis huxleyi, named for its similarity to modern birds. The Latin name Anchiornis derives from a Greek word meaning "near bird", and huxleyi refers to Thomas Henry Huxley, a contemporary of Charles Darwin.
Jinfengopteryginae is a subfamily of bird-like theropod dinosaurs known from the Cretaceous of Eurasia. This group includes relatively few genera, with members discovered in 2005 but the name erected in 2012. Like other troodontids, this group of dinosaurs resided in the Paraves potentially close to the Avialae.
Eosinopteryx is an extinct genus of theropod dinosaurs known to the Late Jurassic epoch of China. It contains a single species, Eosinopteryx brevipenna.
Aurornis is an extinct genus of anchiornithid theropod dinosaurs from the Jurassic period of China. The genus Aurornis contains a single known species, Aurornis xui. Aurornis xui may be the most basal ("primitive") avialan dinosaur known to date, and it is one of the earliest avialans found to date. The fossil evidence for the animal pre-dates that of Archaeopteryx lithographica, often considered the earliest bird species, by about 10 million years.
This timeline of troodontid research is a chronological listing of events in the history of paleontology focused on the troodontids, a group of bird-like theropod dinosaurs including animals like Troodon. Troodontid remains were among the first dinosaur fossils to be reported from North America after paleontologists began performing research on the continent, specifically the genus Troodon itself. Since the type specimen of this genus was only a tooth and Troodon teeth are unusually similar to those of the unrelated thick-headed pachycephalosaurs, Troodon and its relatives would be embroiled in taxonomic confusion for over a century. Troodon was finally recognized as distinct from the pachycephalosaurs by Phil Currie in 1987. By that time many other species now recognized as troodontid had been discovered but had been classified in the family Saurornithoididae. Since these families were the same but the Troodontidae named first, it carries scientific legitimacy.
Jianianhualong is a genus of troodontid theropod dinosaur from the Early Cretaceous of China. It contains a single species, Jianianhualong tengi, named in 2017 by Xu Xing and colleagues based on an articulated skeleton preserving feathers. The feathers at the middle of the tail of Jianianhualong are asymmetric, being the first record of asymmetrical feathers among the troodontids. Despite aerodynamic differences from the flight feathers of modern birds, the feathers in the tail vane of Jianianhualong could have functioned in drag reduction whilst the animal was moving. The discovery of Jianianhualong supports the notion that asymmetrical feathers appeared early in the evolutionary history of the Paraves.
Serikornis is a genus of small, feathered anchiornithid dinosaur from the Upper Jurassic Tiaojishan Formation of Liaoning, China. It is represented by the type species Serikornis sungei.
Anchiornithidae is a family of small paravian dinosaurs. Anchiornithids have been classified at varying positions in the paravian tree, with some scientists classifying them as a distinct family, a basal subfamily of Troodontidae, members of Archaeopterygidae, or an assemblage of dinosaurs that are an evolutionary grade within Avialae or Paraves.
The biogeography of Paravian dinosaurs is the study of the global distribution of Paraves through geological history. Paraves is a clade that includes all of the Theropoda that are more closely related to birds than to oviraptorosaurs. These include Dromaeosauridae and Troodontidae and Avialae. The distribution of paraves is closely related to the evolution of the clade. Understanding the changes in their distributions may shed light on problems like how and why paraves evolve, eventually gaining the ability to fly.
