Barnesoceras

Last updated

Barnesoceras
Temporal range: Early Ordovician
Scientific classification
Kingdom: Animalia
Phylum: Mollusca
Class: Cephalopoda
Subclass: Nautiloidea
Superorder: Plectronoceratoidea
Order: Ellesmerocerida
Family: Ellesmeroceratidae
Genus:Barnesoceras
Flower, 1964

Barnesoceras is a genus of moderately large, Lower Ordovician ellesmoerceratids with a simple, but strongly cyctoconic shell that expands in the vertical dimension more rapidly than in the horizontal, resulting in its being laterally compressed. The aperture is simple, unconstricted, and straight sided from in front. Growth lines show a broad crest on either side that swings forward above the ventral hyponomic sinus. Septa are close spaced and have a shallow curvature. The body, or living, chamber takes up much of the shell. The siphuncle is ventral, close to the longitudinally concave side.

The Ellesmeroceratidae constitute a family within the cephalopod order Ellesmerocerida. They lived from the Upper Cambrian to the Lower Ordovician. They are characterized by straight and endogastric shells, often laterally compressed, so the dorso-ventral dimension is slightly greater than the lateral, with close spaced sutures having shallow lateral lobes and a generally large tubular ventro-marginal siphuncle with concave segments and irregularly spaced diaphragms. Connecting rings are thick and layered, externally straight but thickening inwardly with the maximum near the middle of the segment so as to leave concave depressions on internal siphuncle molds. Septal necks are typically orthochoanitic but vary in length from almost absent (achoanitic) to reaching halfway to the previous septum (hemichoanitic) and may even slope inwardly (loxochoanitic).

Siphuncle

The siphuncle is a strand of tissue passing longitudinally through the shell of a cephalopod mollusk. Only cephalopods with chambered shells have siphuncles, such as the extinct ammonites and belemnites, and the living nautiluses, cuttlefish, and Spirula. In the case of the cuttlefish, the siphuncle is indistinct and connects all the small chambers of that animal's highly modified shell; in the other cephalopods it is thread-like and passes through small openings in the walls dividing the chambers.

All eight species named and described, including the type, Barnsoceras clavum Flower, 1964, were found in the Threadgill Limestone of the Lower Ordovician Tanyard Formation in Texas.

Related Research Articles

The Discosorida are an order of cephalopods that lived from the beginning of the Middle Ordovician, through the Silurian, and into the Devonian. Discosorids are unique in the structure and formation of the siphuncle, the tube that runs through and connects the camerae (chambers) in cephalopods, which unlike those in other orders is zoned longitudinally along the segments rather than laterally. Siphuncle structure indicated that the Discosorida evolved directly from the Plectronoceratida rather than through the more developed Ellesmerocerida, as did the other orders. Finally and most diagnostic, discosorids developed a reinforcing, grommet-like structure in the septal opening of the siphuncle known as the bullette, formed by a thickening of the connecting ring as it draped around the folded back septal neck.

Plectronocerida order of molluscs

Plectronocerida is a primitive order from which subsequent cephalopod orders are ultimately derived.

Piloceratidae family of molluscs

The Piloceratidae are a compressed, rapidly expanding, cyrtoconic brevicones with holochoanitic ventral siphuncles and simple endocones. Most likely evolved from Clitendoceras, a narrow, slightly endogastric genus intermediate in form between straight shelled Proendoceras and the bulkier Piloceratidae. Found in shallow carbonate marine sediments of Demingian through the Cassinian age,.

Clitendoceras is a genus of cephalopods in the order Endocerida from the Lower Ordovician with an elongate shell with a slight downward, endogastric, curvature and a siphuncle that lies along the ventral margin. Common for endocerids, the chambers are short and the septa close spaced with sutures sloping forward across the back of the shell. Septal necks are short in the young, lengthening in the adult. Endocones are simple, but with the ventral side of the last formed projecting forward. The endosiphotube running down the middle is arched on top and somewhat flat on the lower side.

Actinoceras is the principal and root genus of the Actinoceratidae, a major family in the Actinocerida, that lived during the Middle and Late Ordovician.

Lambeoceras is a genus of rather large actinocerids with a convexly lenticular cross section from the Upper Ordovician of North America, and the sole representative of the family Lambeoceratidae.

Clarkoceras is a genus of breviconic ellesmerocerid cephalopods, one of only two genera known to have crossed from the Late Cambrian, Trempealeauan, into the Early Ordovician, Gasconadian. ; the other being Ectenolites.

The Reudemannoceratidae are the ancestral and most primitive of the Discosorida, an order of cephalopods from the early Paleozoic. The Reudemannoceratidae produced generally medium-sized endogastric and almost straight shells with the siphuncle slightly ventral from the center.

Baltoceratidae is an extinct family of orthoconic cephalopods belonging to the subclass Nautiloidea endemic to what would be Asia, Australia, Europe, North America, and South America during the Ordovician living from about 480–460 mya, existing for approximately 20 million years.

Cyrtobaltoceras is an extinct cephalopod genus known from the upper Lower Ordovician Fort Cassin Formation at Valcour, N.Y. that is included in the Nautiloid family Baltoceratidae

Oncoceratidae family of molluscs

Oncoceratidae is a family of nauatiloid cephalopods in the order Oncocerida established by Hyatt, 1884, that range from the Middle Ordovician to the Upper Silurian.

The Bassleroceratidae is a family of gradually expanding, smooth ellesmerocerids with a slight to moderate exogastric curvature, subcircular to strongly compressed cross section, and ventral orthochaonitc siphuncle. The ventral side is typically more sharply rounded than the dorsal side and septa are close spaced. Connecting rings are thick and slightly expanded into the siphuncle, making the segments slightly concave; characteristic of the Ellesmerocerida.

The Actinoceriatidae are a family of actinocerids named by Saemann in 1853 for those that grew to have large shells with blunt apecies and large siphuncles with widely expanded segments and a generally arcuate endosiphucular canal system. Their range is from the upper Middle Ordovician to the Lower Silurian. Actinocerids are generally straight-shelled nautiloid cephalopods with a siphuncle composed of expanded segments, typically with thin connecting rings, in which the internal deposits are penetrated by a system of canals

Nybyoceras is an actinocerid genus assigned to the Armenoceratidae and similar to Armenoceras except for having a siphuncle close to the ventral side of the shell.

Protcycloceratidae is an extinct family of slender, commonly annulate, members of the cephalopod order Ellesmerocerida that lived during the Early Ordovician.

Endoceratidae is a family of large to very large straight shelled nautiloid cephalopods belonging to the order Endocerida that lived during the Middle and Late Ordovician. They include the largest known Paleozoic invertebrates, represented by Endoceras and Cameroceras.

The Cyrtogomphoceratidae are a family in the cephalopod order Discosorida that comprises genera commonly with compressed, endogastrically curved shells. Siphuncles lie close to the ventral side, segments are broadly inflated, connecting rings thick and apically expanded thick bullettes. Chambers are short, separated by shallow, dish shaped septa. Apertures are generally simple.

Murrayoceras is a nautilid cephalopod included in the orthocerid family Baltoceratidae, widespread in the Middle Ordovician of North America, characterized by a depressed orthoconic shell with a subtriangular cross section and flattened venter and a proportionally large verntral siphuncle, 0.15 to 0.3 the dorso-ventral shell diameter. Septa are close spaced with sutures forming broad lobes on the upper flanks and ventral surface.

Ellesmeroceras is the type genus for the Ellesmeroceratidae, a family of primitive nautiloid cephalopods, that is characterized by its small, generally compressed, gradually expanded, orthoconic shell, found in Lower Ordovician marine sediments. The septa are close spaced and the siphuncle is ventral, about 0.2 the diameter of the shell. Septal necks are typically orthochoanitic but may slant inwardly (loxochoanitic) or reach halfway to the previous septum (hemichoanitic). Connecting rings are thick. As common for the Ellesmerocerida, Ellesmeroceras has diaphragms within the siphuncle tube.

Eothinoceratidae is a family of Lower Ordovician nautiloid cephalopods included in the Ellesmerocerida originally established for the genus Eothinoceras.

References