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Cyathea fuliginosa | |
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Scientific classification | |
Kingdom: | Plantae |
Division: | Pteridophyta |
Class: | Pteridopsida |
Order: | Cyatheales |
Family: | Cyatheaceae |
Genus: | Cyathea |
Subgenus: | Cyathea |
Section: | Alsophila |
Species: | C. fuliginosa |
Binomial name | |
Cyathea fuliginosa (Christ) Copeland, 1909 | |
Synonyms | |
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Cyathea fuliginosa is a species of tree fern native to the islands of Luzon, Biliran and Mindanao in the Philippines, where it grows in forest at an altitude of 640–2400 m. This tree fern has an erect trunk up to 5 m tall or more. Fronds are bi- or tripinnate and 1–3 m in length. The stipe is warty and may also (or instead) have short spines as well as many scattered scales towards the base. These scales are dark, glossy and have fragile edges. Sori occur near the fertile pinnule midvein and are protected by firm, dark indusia.
The order Cyatheales, which includes the tree ferns, is a taxonomic division of the fern class, Polypodiopsida. No clear morphological features characterize all of the Cyatheales, but DNA sequence data indicate the order is monophyletic. Some species in the Cyatheales have tree-like growth forms, but others have rhizomes.
Luzon is the largest and most populous island in the Philippines. It is ranked 15th largest in the world by land area. Located in the northern region of the archipelago, it is the economic and political center of the nation, being home to the country's capital city, Manila, as well as Quezon City, the country's most populous city. With a population of 53 million as of 2015,, it is the fourth most populous island in the world containing 52.5% of the country's total population.
Biliran is an island province in the Philippines located in the Eastern Visayas region. Biliran is one of the country's smallest and newest provinces. Formerly a sub-province of Leyte, it became an independent province in 1992.
Cyathea alpicola is a species of tree fern native to central Sumatra, where it grows in montane rain forest at an altitude of 2000–2750 m. The erect trunk can reach 5 m or more in height. Fronds are bi- or tripinnate and usually 2–3 m long. They have a tendency to persist on the plant after withering, forming an irregular skirt around the trunk. This species has a spiny stipe, which is covered with a woolly layer of scales. There appear to be two forms with different scales; either dark glossy brown with a broad paler margin and fragile edges, or small, brown and finely fringed. Sori are covered by thin, fragile indusia and occur near the midvein of fertile pinnules. The closest relative of C. alpicola is thought to be Cyathea polycarpa, which differs by lacking spines altogether. Cyathea macropoda and Cyathea magnifolia may also be closely allied with this species.
Cyathea amboinensis is a species of tree fern native to the Maluku Islands and possibly central and southern Sulawesi, where it grows in swamp and forest at low elevations. This species has an erect trunk, usually 2–4 m tall. Fronds are 1–2 m long and bi- or tripinnate. The stipe is warty near the base and covered with scattered scales that are dark, glossy, and have fragile edges. Sori occur near the midvein of fertile pinnules and are covered by small, dark indusia which resemble small scales.
Cyathea aneitensis is a species of tree fern native to Vanuatu and possibly New Caledonia. This species has an erect trunk up to 3 m tall. Fronds are bipinnate and may reach 2 m in length. The rachis and stipe are either very dark and smooth or have a few scales towards the base of the stipe. The scales are dark and narrow. Sori occur near the pinnule midvein and are covered by large, thin, fragile indusia. The closest relative of C. aneitensis appears to be Cyathea vieillardii. It can be distinguished from that species by its very dark stipes and frond bases.
Cyathea archboldii is a species of tree fern native to New Guinea and Bougainville, where it is common in submontane rain forest at an altitude of 1000–3000 m. The trunk is erect and up to about 3 m tall. Fronds are bipinnate and 2–3 m long. The rachis may be purplish and has short spines and scales. The scales range in colour from pale to brown, to bicoloured and have fragile edges. Sori occur in two rows along each side of the pinnule midvein and are covered by firm indusia.
Cyathea brevipinna is a species of tree fern endemic to the higher parts of Mount Gower on Lord Howe Island, where it grows in exposed areas at an altitude of about 790 m. The trunk is erect and may reach 3 m in height. It is often covered with reddish brown scales and stipe bases. This species may produce stolons at ground level. Fronds are tripinnate, densely crowded, and up to about 3 m long. The stipe is brown and sometimes warty after scales fall off. The scales are long, glossy dark brown, with a distinctly narrow apex and fragile paler edges. Sori are attached to deeply divided fertile pinnules that may uncurl over the sori. Indusia are firm and large. C. brevipinna is a stunted plant with short pinnae.
Cyathea camerooniana is a species of tree fern native to Sierra Leone, Cameroon, northern Angola and western Uganda, where it grows in montane forest at an altitude of 900–1200 m. The trunk is erect and 2–3 m tall. Fronds are pinnate and 2–3 m long. The rachis ranges in colour from dark to pale and has some hairs on the underside. The stipe is dark and is covered with scales throughout. Where scales have fallen, small warts are present. The scales are glossy brown and have a thin, dull edge. Sori occur at the forks of veins and are covered by thin indusia, which range in shape from cup- to saucer-like.
Cyathea capensis is a regionally widespread and highly variable species of tree fern. It is indigenous to Southern Africa and South America.
Cyathea costalisora is a species of tree fern native to western New Guinea, where it grows on the edges of forest and in moist hollows at an altitude of 1900–3225 m. The trunk is erect, up to 4 m tall and may branch near the base. Fronds are bi- or tripinnate and 1-1.5 m long. The stipe is warty, especially where scales have fallen. The scales are pale and have a distinctive dark glossy central region, with a paler dull margin. Sori are round and occur near the fertile pinnule midvein. They are covered by firm, dark indusia that are cup-like in appearance.
Cyathea costulisora is a species of tree fern native to Sumatra. The trunk is erect and usually 1–4 m tall. Fronds are bi- or tripinnate and 1–2 m long. The stipe is covered with warts and scales. The scales are either pale and glossy or dark and flat. Sori occur near the fertile pinnule midvein and are covered by large, firm, brown indusia.
Cyathea edanoi is a species of tree fern endemic to Luzon in the Philippines, where it grows in montane forest at an altitude of about 1300 m. The trunk is erect and 1–2 m tall. Fronds are bi- or tripinnate and approximately 1 m long. The stipe is covered in dark, glossy scales that have narrow, fragile edges. Sori occur near the midvein of fertile pinnules and are covered, sometimes half-covered, by large, firm, brown indusia.
Cyathea erinacea is a species of tree fern native to Mexico, Costa Rica, Panama, Venezuela, Colombia, Ecuador, Peru and Bolivia.
Cyathea gigantea is a species of tree fern native to northeastern to southern India, Sri Lanka, Nepal to Myanmar, Thailand, Laos, Vietnam, the Malay Peninsula, as well as central Sumatra and western Java. It grows in moist open areas at an altitude of 600–1000 m. The trunk of this species is erect and may be as tall as 5 m or more. Fronds are bi- or tripinnate and usually 2–3 m long. The rachis is long, dark to black in colouration and rough in appearance after the fall of scales. These scales are dark brown, glossy and have a narrow paler margin and fragile edges. Sori are round and indusia absent.
Cyathea glabra is a species of tree fern native to Borneo, western Java, Sumatra and the Malay Peninsula, where it grows in lowland swamp forest and montane forest at an elevation of up to 1500 m. The trunk of this plant is erect and 2–4 m tall. Fronds are bi- or tripinnate and 1–2 m in length. Characteristically of this species, the lowest pinnae may be significantly reduced. The stipe is very dark and bears basal scales. These scales are dark, glossy and have a paler margin and fragile edges. Sori are produced in groups of one to three on fertile pinnule veins. They lack indusia.
Cyathea hancockii is a species of tree fern native to the Ryukyu Islands, Japan, Taiwan and Hong Kong. The specific epithet hancockii commemorates William Hancock (1847-1914), who collected numerous plants in Japan, China and Southeast Asia. It grows in forest, on stream banks, and in forest margins at an elevation of about 600 m or higher.
Cyathea heterochlamydea is a little-known species of tree fern native to the islands of Luzon, Panay, Negros and Mindanao in the Philippines, where it grows in montane forest. The trunk of this plant is erect and usually up to 4 m tall or more. Fronds may be bi- or tripinnate and 1–2 m in length. The stipe is warty and/or bears short spines and scales. These scales are dark, glossy and have a narrow pale margin. Sori are borne near the fertile pinnule midvein and are protected by firm, brown indusia.
Cyathea khasyana is a species of tree fern. Its natural distribution extends from India to Myanmar, although it is absent from Sri Lanka. C. khasyana grows in forest at an altitude of 1400–1700 m. The trunk of this plant is erect and 5–7 m tall. Fronds may be bi- or tripinnate and 2–3 m in length. C. khasyana has a long, dark stipe that is covered in numerous scales. These scales are dark and have broad, pale, fringed edges. Sori are borne near the midvein of fertile pinnules and lack indusia.
Cyathea hymenodes is a species of tree fern native to the Malay Peninsula, Sumatra, and possibly Java, where it grows in montane forest at an altitude of 900–2000 m. The trunk of this plant is erect and 2–4 m tall. Fronds may be bi- or tripinnate and 1–2 m in length. The stipe may have a pair of reduced pinnae towards the base and is covered with scattered scales. These scales are dark and have very fragile edges. Sori are borne near the fertile pinnule midvein and are protected by firm, brown indusia that are saucer-like in appearance.
Cyathea imbricata is a species of tree fern endemic to Western New Guinea, where it grows in open forest at an altitude of 3240 m. The trunk of this plant is erect and approximately 2 m tall. Fronds may be bi- or tripinnate and are usually less than 1 m in length. The stipe is dark, spiny, and covered with caducous scales. These scales are glossy brown in colouration and have a paler margin and fragile edges. Sori are borne in groups of one to four per pinnule lobe. They are protected by firm indusia.
Cyathea javanica is a species of tree fern native to western Java and Sumatra, in Indonesia.
Cyathea latebrosa is a common and widespread species of tree fern native to Indochina. Its natural range covers Cambodia and Thailand, and stretches from the Malay Peninsula to Indonesia, where it is present on the islands of Borneo and Sumatra. Plants reported from India and Sri Lanka have thinner indusia and may represent a separate, as-yet undescribed, species. C. latebrosa grows in a wide range of habitats, including forest, secondary forest, and plantations, from sea level up to an elevation of about 1500 m.
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