Dmanisi skull 3

Last updated
Dmanisi skull 3
Dmanisi cranium D2700 (B).jpg
Catalog no.D2700
Common nameDmanisi skull 3
Species Homo erectus georgicus
Agec. 1.81 million years
Place discovered Dmanisi, Kvemo Kartli, Georgia
Date discovered2001
Discovered byAbesalom Vekua and David Lordkipanidze

D2700, also known as Dmanisi skull 3, is one of five skulls discovered in Dmanisi, Georgia in 2001 and classified as early Homo erectus . It is an almost complete skull and is in an exceptionally good condition. It was dated stratigraphically as about 1.8 million years old.

Contents

Since the publication of the Dmanisi skull 5 in 2013, all of the five Dmanisi skulls (Skull 3 included) have contributed to the ongoing debate on human taxonomy, with some experts[ who? ][ citation needed ] proposing the re-categorization of Homo ergaster , and possibly even[ citation needed ] Homo habilis , as morphologically diverse subspecies of H. erectus.

Discovery

D2700 and D2735 were found in 2001, just a decade after the first discovery of an early hominin mandible D211 at Dmanisi on September 24, 1991. In 1999, partial crania D2280 and D2282 were discovered. Cranium D2282 is likely the accompanying skull to mandible D211 and to be the remains of a young adult around 18–20 years old. Skull D2280 is inferred to have been an adult around 25–30 years old at the time of death. [1] Moreover, in 2005, a new complete skull cranium was found: D4500. This cranium is inferred to be an adult skull and is more commonly known as "Skull 5." D4500's features are very rare compared to early Homo in that it had a small braincase yet an unusually large prognathic face. [2] "Skull 5" has an accompanying mandible, D2600, which was found in 2000.

In 1999, two other skulls had been found at the same site—D2280 and D2282. D2280 was a near-complete brain-case with 780 cc brain-size. D2282 was a cranium and it included many of the facial and upper jaw bones. Its brain size was about 650 cc. In 1991 and 2000, two more lower jaws D211 and D2600 were discovered from the same site.

The five Dmanisi skulls are: D2280 (skull 1), D2282/D211 (skull 2), D2700/D2735 (skull 3), D3444/D3900 (skull 4) and D4500/D2600 (skull 5)

Morphology

Dmanisi skull 3, Fossils skull D2700 and D2735 jaw, two of several found in Dmanisi in the Georgian Caucasus Homo Georgicus IMG 2921.JPG
Dmanisi skull 3, Fossils skull D2700 and D2735 jaw, two of several found in Dmanisi in the Georgian Caucasus

The skull was found in an exceptionally good condition including a lower jaw (D2735) found about a meter away which is considered to be of the same person. D2700 is smaller than D2282. The fossil was dated stratigraphically, as from a site that was occupied between c. 1.85 and 1.77 million years ago. Its brain size is estimated at 600 cc.[ citation needed ]

There are many characteristics in which it resembles H. ergaster (or H. erectus) and also a number that resembles the ER1813.

Vekua et al. (2002) concluded: [3]

In overall shape, D2700 is similar to D2280 and D2282, and D2735 resembles D211. Despite certain differences among these Dmanisi individuals, we do not see sufficient grounds for assigning them to more than one hominid [now, "hominin"] taxon. We view the new specimen as a member of the same population as the other fossils, and we here assign the new skull provisionally to Homo erectus.

All Dmanisi hominin remains were suggested to have been buried quickly after death. D2700 was found with four upper teeth, while D2735 contained eight teeth. As both fossil hominin remains are jointed, it is found that they ultimately fit one another. D2700's narrow nasal bones were found inferiorly broken from the orbital margins. Because the cranium's upper molars were partially erupted, D2700 is believed to have been the skull of a 13 to 15-year-old individual. [1] It is considered that the skull was of a teenager.

There is some uncertainty as to the sex of the individual. The smaller size of the D2700 skull compared to skulls D2280 and D2282 suggests as well as its more slender feature suggest female sex, [1] but the very large size of huge upper canines and large crowns have been argued to support classification as male. [4]

Classification

Detailed view of Dmanisi skull 3 Paleolithic skull from Dmanisi.jpg
Detailed view of Dmanisi skull 3
View of Dmanisi skull 3 Dmanisi fossils D2700 + D2735.JPG
View of Dmanisi skull 3

It was conjectured that the categorical differences seen in for instance the mandibular and dental features of the crania from Dmanisi, may be ascribed to taxonomic diversity, as opposed to all five deriving from the same H. erectus lineage. [5] The rather small braincase of D2700 was believed to be at odds with the characteristics of H. erectus. H. erectus georgicus was besides small in size compared to H. erectus . [1] For these reasons, D2700, along with the other Dmanisi fossil remains, has been variously[ by whom? ][ year needed ] classified as a new subspecies, H. erectus georgicus, or as a new species in its own right, H. georgicus .

See also

Related Research Articles

<i>Homo habilis</i> Archaic human species from 2.8 to 1.65 mya

Homo habilis is an extinct species of archaic human from the Early Pleistocene of East and South Africa about 2.3 million years ago to 1.65 million years ago (mya). Upon species description in 1964, H. habilis was highly contested, with many researchers recommending it be synonymised with Australopithecus africanus, the only other early hominin known at the time, but H. habilis received more recognition as time went on and more relevant discoveries were made. By the 1980s, H. habilis was proposed to have been a human ancestor, directly evolving into Homo erectus, which directly led to modern humans. This viewpoint is now debated. Several specimens with insecure species identification were assigned to H. habilis, leading to arguments for splitting, namely into "H. rudolfensis" and "H. gautengensis" of which only the former has received wide support.

<i>Homo ergaster</i> Extinct species or subspecies of archaic human

Homo ergaster is an extinct species or subspecies of archaic humans who lived in Africa in the Early Pleistocene. Whether H. ergaster constitutes a species of its own or should be subsumed into H. erectus is an ongoing and unresolved dispute within palaeoanthropology. Proponents of synonymisation typically designate H. ergaster as "African Homo erectus" or "Homo erectus ergaster". The name Homo ergaster roughly translates to "working man", a reference to the more advanced tools used by the species in comparison to those of their ancestors. The fossil range of H. ergaster mainly covers the period of 1.7 to 1.4 million years ago, though a broader time range is possible. Though fossils are known from across East and Southern Africa, most H. ergaster fossils have been found along the shores of Lake Turkana in Kenya. There are later African fossils, some younger than 1 million years ago, that indicate long-term anatomical continuity, though it is unclear if they can be formally regarded as H. ergaster specimens. As a chronospecies, H. ergaster may have persisted to as late as 600,000 years ago, when new lineages of Homo arose in Africa.

<i>Homo rudolfensis</i> Extinct hominin from the Early Pleistocene of East Africa

Homo rudolfensis is an extinct species of archaic human from the Early Pleistocene of East Africa about 2 million years ago (mya). Because H. rudolfensis coexisted with several other hominins, it is debated what specimens can be confidently assigned to this species beyond the lectotype skull KNM-ER 1470 and other partial skull aspects. No bodily remains are definitively assigned to H. rudolfensis. Consequently, both its generic classification and validity are debated without any wide consensus, with some recommending the species to actually belong to the genus Australopithecus as A. rudolfensis or Kenyanthropus as K. rudolfensis, or that it is synonymous with the contemporaneous and anatomically similar H. habilis.

<i>Homo</i> Genus of hominins that includes humans and their closest extinct relatives

Homo is a genus of great ape that emerged from the genus Australopithecus and encompasses the extant species Homo sapiens and a number of extinct species classified as either ancestral or closely related to modern humans. These include Homo erectus and Homo neanderthalensis. The oldest member of the genus is Homo habilis, with records of just over 2 million years ago. Homo, together with the genus Paranthropus, is probably most closely related to the species Australopithecus africanus within Australopithecus. The closest living relatives of Homo are of the genus Pan, with the ancestors of Pan and Homo estimated to have diverged around 5.7-11 million years ago during the Late Miocene.

<span class="mw-page-title-main">Dmanisi</span> Town in Kvemo Kartli, Georgia

Dmanisi is a town and archaeological site in the Kvemo Kartli region of Georgia approximately 93 km southwest of the nation’s capital Tbilisi in the river valley of Mashavera. The hominin site is dated to 1.8 million years ago. It was the earliest known evidence of hominins outside Africa before stone tools dated to 2.1 million years were discovered in 2018 in Shangchen, China.

<i>Homo rhodesiensis</i> Species of primate (fossil)

Homo rhodesiensis is the species name proposed by Arthur Smith Woodward (1921) to classify Kabwe 1, a Middle Stone Age fossil recovered from Broken Hill mine in Kabwe, Northern Rhodesia. In 2020, the skull was dated to 324,000 to 274,000 years ago. Other similar older specimens also exist.

Human taxonomy is the classification of the human species within zoological taxonomy. The systematic genus, Homo, is designed to include both anatomically modern humans and extinct varieties of archaic humans. Current humans have been designated as subspecies Homo sapiens sapiens, differentiated, according to some, from the direct ancestor, Homo sapiens idaltu.

<span class="mw-page-title-main">Lantian Man</span> Subspecies of the genus Homo (fossil)

Lantian Man, Homo erectus lantianensis) is a subspecies of Homo erectus known from an almost complete mandible from Chenchiawo (陈家窝) Village discovered in 1963, and a partial skull from Gongwangling (公王岭) Village discovered in 1964, situated in Lantian County on the Loess Plateau. The former dates to about 710–684 thousand years ago, and the latter 1.65–1.59 million years ago. This makes Lantian Man the second-oldest firmly dated H. erectus beyond Africa, and the oldest in East Asia. The fossils were first described by Woo Ju-Kan in 1964, who considered the subspecies an ancestor to Peking Man.

<span class="mw-page-title-main">KNM ER 3733</span> Hominin fossil

KNM ER 3733 is a fossilized hominid cranium of the extinct hominid Homo ergaster, alternatively referred to as African Homo erectus. It was discovered in 1975 in Koobi Fora, Kenya, right next to Lake Turkana, in a survey led by Richard Leakey, by a field worker called Bernard Ngeneo.

<span class="mw-page-title-main">Post-orbital constriction</span>

In physical anthropology, post-orbital constriction is the narrowing of the cranium (skull) just behind the eye sockets found in most non-human primates and early hominins. This constriction is very noticeable in non-human primates, slightly less so in Australopithecines, even less in Homo erectus and completely disappears in modern Homo sapiens. Post-orbital constriction index in non-human primates and hominin range in category from increased constriction, intermediate, reduced constriction and disappearance. The post-orbital constriction index is defined by either a ratio of minimum frontal breadth (MFB), behind the supraorbital torus, divided by the maximum upper facial breadth (BFM), bifrontomalare temporale, or as the maximum width behind the orbit of the skull.

<span class="mw-page-title-main">David Lordkipanidze</span>

David Otaris dze Lordkipanidze is a Georgian anthropologist and archaeologist, Professor (2004), Dr.Sc. (2002), Corresponding Member of the Georgian National Academy of Sciences (2009), and since 2004 the General Director of the Georgian National Museum (GNM). He is a son of the archaeologist Otar Lordkipanidze.

<i>Homo erectus</i> Extinct species of archaic human

Homo erectus is an extinct species of archaic human from the Pleistocene, with its earliest occurrence about 2 million years ago. Its specimens are among the first recognizable members of the genus Homo.

<i>Homo gautengensis</i> Name proposed for an extinct species of hominin from South Africa

Homo gautengensis is a species name proposed by anthropologist Darren Curnoe in 2010 for South African hominin fossils otherwise attributed to H. habilis, H. ergaster, or, in some cases, Australopithecus or Paranthropus. The fossils assigned to the species by Curnoe cover a vast temporal range, from about 1.8 million years ago to potentially as late as 0.8 million years ago, meaning that if the species is considered valid, H. gautengensis would be both one of the earliest and one of the longest lived species of Homo.

Givi Maisuradze is a Georgian geologist, professor and doctor of science. He and his wife Nina Klopotovskaia, a paleontologist, were part of the research team that discovered early hominin skulls and later skeletons dating back to 1.8 million years old in Dmanisi, Georgia.

<span class="mw-page-title-main">Dmanisi skull 5</span> Hominin fossil

The Dmanisi skull, also known as Skull 5 or D4500, is one of five skulls discovered in Dmanisi, Georgia and classified as early Homo erectus. Described in a publication in October 2013, it is estimated to be about 1.8 million years old and is the most complete skull of a Pleistocene Homo species, and the first complete adult hominin skull of that degree of antiquity. According to researchers, the discovery "provides the first evidence that early Homo comprised adult individuals with small brains but body mass, stature and limb proportions reaching the lower range limit of modern variation."

<span class="mw-page-title-main">Early expansions of hominins out of Africa</span> First hominin expansion into Eurasia (2.1–0.1 Ma)

Several expansions of populations of archaic humans out of Africa and throughout Eurasia took place in the course of the Lower Paleolithic, and into the beginning Middle Paleolithic, between about 2.1 million and 0.2 million years ago (Ma). These expansions are collectively known as Out of Africa I, in contrast to the expansion of Homo sapiens (anatomically modern humans) into Eurasia, which may have begun shortly after 0.2 million years ago.

<span class="mw-page-title-main">Dmanisi hominins</span> Hominid species or subspecies discovered in Dmanisi, Georgia

The Dmanisi hominins, Dmanisi people, or Dmanisi man were a population of Early Pleistocene hominins whose fossils have been recovered at Dmanisi, Georgia. The fossils and stone tools recovered at Dmanisi range in age from 1.85 to 1.77 million years old, making the Dmanisi hominins the earliest well-dated hominin fossils in Eurasia and the best preserved fossils of early Homo from a single site so early in time, though earlier fossils and artifacts have been found in Asia. Though their precise classification is controversial and disputed, the Dmanisi fossils are highly significant within research on early hominin migrations out of Africa. The Dmanisi hominins are known from over a hundred postcranial fossils and five famous well-preserved skulls, referred to as Dmanisi Skulls 1–5.

<span class="mw-page-title-main">Dmanisi historic site</span> Historic site in Kvemo Kartli, Georgia

The Dmanisi historic site is a historic and archaeological site in Georgia, located north of the village of Patara Dmanisi, Dmanisi Municipality, in south-central region of Kvemo Kartli, some 85 km southwest of Tbilisi, Georgia's capital.

<span class="mw-page-title-main">Salé cranium</span> Enigmatic Middle Pleistocene hominin specimen

The Salé cranium is a pathological specimen of enigmatic Middle Pleistocene hominin discovered from Salé, Morocco by quarrymen in 1971. Since its discovery, the specimen has variously been classified as Homo sapiens, Homo erectus, Homo rhodesiensis/bodoensis, or Homo heidelbergensis. Its pathological condition and mosaic anatomy has proved difficult to classify. It was discovered with few faunal fossils and no lithics, tentatively dated to 400 ka by some sources.

<span class="mw-page-title-main">Kocabaş cranium</span> Damaged Homo erectus calvarium.

The Kocabaş cranium is the damaged calvarium fossil of a young Homo erectus discovered near the village of Kocabaş, located in the Denizli Province of Turkey by quarry workers in 2002.

References

  1. 1 2 3 4 Lumley (2006). "Human Remains from the Upper Pliocene- Early Pleistocene Dmanisi site, Georgia (1991-2000). Part 1 Fossil Skulls D2280, D2282, and D2700". L'Anthropologie. 110 (1): 1–110.
  2. Lordkipanidze, D.; Ponce de Leon, M. S.; Margvelashvili, A.; Rak, Y.; Rightmire, G. P.; Vekua, A.; Zollikofer, C. P. E. (17 October 2013). "A Complete Skull from Dmanisi, Georgia, and the Evolutionary Biology of Early Homo". Science. 342 (6156): 326–331. doi:10.1126/science.1238484. PMID   24136960. S2CID   20435482.
  3. Gabunia L., de Lumley M.-A., Vekua A., Lordkipanidze D., and de Lumley H. (2002): Découverte d'un nouvel hominidé à Dmanissi (Transcaucasie, Georgie). C.R.Pale vol 1, 2002:243-53, doi : 10.1016/S1631-0683(02)00032-5, cited after: "Dmanisi's Paleoanthropological Importance". Anthropology.net. 2007. Archived from the original on 1 October 2007. Retrieved 15 October 2012.
  4. Vekua, A. (5 July 2002). "A New Skull of Early Homo from Dmanisi, Georgia". Science. 297 (5578): 85–89. doi:10.1126/science.1072953. PMID   12098694. S2CID   32726786.
  5. Schwartz, J. H.; Tattersall, I.; Chi, Z. (24 April 2014). "Comment on "A Complete Skull from Dmanisi, Georgia, and the Evolutionary Biology of Early Homo"". Science. 344 (6182): 360. doi: 10.1126/science.1250056 . PMID   24763572.