Dmanisi skull 5

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Dmanisi skull 5
D4500.jpg
Skull 5 in National Museum
Catalog no.D4500
Common nameDmanisi skull 5
Species Homo erectus
Age1.8 million years
Place discovered Dmanisi in Georgia
Date discovered2005
Discovered by David Lordkipanidze

The Dmanisi skull, also known as Skull 5 or D4500, is one of five skulls discovered in Dmanisi, Georgia and classified as early Homo erectus . Described in a publication in October 2013, it is estimated to be about 1.8 million years old and is the most complete skull of a Pleistocene Homo species, [1] [2] and the first complete adult hominin skull of that degree of antiquity. [3] [4] According to researchers, the discovery "provides the first evidence that early Homo comprised adult individuals with small brains but body mass, stature and limb proportions reaching the lower range limit of modern variation." [3]

Contents

The skull has been the cause of a paleontological controversy that is still ongoing as of 2017: many hominin fossils thought to be from different species such as Homo rudolfensis or Homo habilis may not have been separate species at all. Rather, they may have been a single evolving lineage. [2]

Discovery of the skull

Excavation. Skull in dmanisi archaeological site.JPG
Excavation.

In 1991, Georgian scientist David Lordkipanidze found traces of early human occupation in the cave at Dmanisi in Georgia: a hamlet and an archaeological site about 90 kilometres (56 mi) southwest of the country's capital, Tbilisi. Since then, five early hominin skulls have been discovered at the site. Skull 5, found in 2005, is the most complete specimen of them all. After further analysis, it was coupled with its mandible (D2600) found 5 years earlier. The final analysis of the discovery took many years and was only published in 2013. [1]

Description of the skull

David Lordkipanidze (centre) on the archeological site at Dmanisi, 2010 Amb. Bass-with D.Lordkipanidze at Dmanisi.JPG
David Lordkipanidze (centre) on the archeological site at Dmanisi, 2010

Despite a brain volume the size of a large Australopithecus male, (546 cubic centimetres (33.3 in3)) all the characteristics of Homo erectus are clearly visible on Skull 5. The braincase sizes of the other Dmanisi skulls are between 601 cubic centimetres (36.7 in3) and 730 cubic centimetres (45 in3). [1]

The cranial superstructures of Skull 5, which are prominent and massive, suggest that it was probably a male. Its face is one of the largest of any human ancestor discovered to date. [1] The zygomatic arches are massive with the right one displaying many traces of deformations, probably due to fractures and injuries. Researchers estimate that the height of the individual was between 146 centimetres (4.79 ft) and 166 centimetres (5.45 ft) and his weight between 47 kilograms (104 lb) and 50 kilograms (110 lb). [1]

Dating

Stone tools found on the Dmanisi paleontological site Dmanisi stone tool 1.jpg
Stone tools found on the Dmanisi paleontological site

Until the 1980s, scientists assumed that hominins had been restricted to the African continent for the whole of the Early Pleistocene (until about 0.8 Ma), only migrating out during a phase named Out of Africa I. Thus, the vast majority of archaeological effort was disproportionately focused on Africa. [5] The Dmanisi archeological site is the earliest hominin site out of Africa and the analysis of its artifacts showed that some hominins, chiefly the Homo erectus georgicus had left Africa as far back as 1.85 million years ago. All of the 5 skulls are roughly the same age. [1]

Consequently, the paleontological significance of the dating of the Dmanisi skulls is major. It is worth noting that it was not associated with the discovery of skull 5, but with another specimen found at the same site, the Dmanisi Skull 3, also known as D2700.

Controversy and significance of the discovery

The Dmanisi skulls, especially Skull 5 with its comparatively tiny 546 cubic centimetres (33.3 in3) brain compared to other skulls found at the site, suggests that the earliest species of the genus Homo were actually subspecies of the species erectus. [1] The variation in morphology of all the Dmanisi skulls is so large that had they been discovered on different archaeological sites, they most likely would have been classified as different species. However, all Dmanisi skulls have the same age and have been found at exactly the same place.

Faced with the wide variation between Skull 5 and the other Dmanisi skulls, Georgian and University of Zurich researchers were prompted to examine normal variations in modern human skulls and chimpanzees skulls. They found that while they looked different from one another, the great variations among all Dmanisi skulls were no greater than those seen among modern people and among chimpanzees. [1] Consequently, it was entirely possible that such a discrepancy could be found in Homo erectus.

This discovery led the scientists to suggest that two species of early hominins, Homo rudolfensis and Homo habilis , were actually both Homo erectus . [2] This is an ongoing debate among the scientific community. [6]

List of Dmanisi 5 specimens

The five Dmanisi skulls are named:

See also

Related Research Articles

<i>Homo habilis</i> Archaic human species from 2.1 to 1.5 mya

Homo habilis is an extinct species of archaic human from the Early Pleistocene of East and South Africa about 2.31 million years ago to 1.65 million years ago (mya). Upon species description in 1964, H. habilis was highly contested, with many researchers recommending it be synonymised with Australopithecus africanus, the only other early hominin known at the time, but H. habilis received more recognition as time went on and more relevant discoveries were made. By the 1980s, H. habilis was proposed to have been a human ancestor, directly evolving into Homo erectus which directly led to modern humans. This viewpoint is now debated. Several specimens with insecure species identification were assigned to H. habilis, leading to arguments for splitting, namely into "H. rudolfensis" and "H. gautengensis" of which only the former has received wide support.

<i>Kenyanthropus</i> Oldest-known tool-making hominin

Kenyanthropus is a hominin genus identified from the Lomekwi site by Lake Turkana, Kenya, dated to 3.3 to 3.2 million years ago during the Middle Pliocene. It contains one species, K. platyops, but may also include the 2 million year old Homo rudolfensis, or K. rudolfensis. Before its naming in 2001, Australopithecus afarensis was widely regarded as the only australopithecine to exist during the Middle Pliocene, but Kenyanthropus evinces a greater diversity than once acknowledged. Kenyanthropus is most recognisable by an unusually flat face and small teeth for such an early hominin, with values on the extremes or beyond the range of variation for australopithecines in regard to these features. Multiple australopithecine species may have coexisted by foraging for different food items, which may be reason why these apes anatomically differ in features related to chewing.

<i>Homo ergaster</i> Extinct species or subspecies of archaic human

Homo ergaster is an extinct species or subspecies of archaic humans who lived in Africa in the Early Pleistocene. Whether H. ergaster constitutes a species of its own or should be subsumed into H. erectus is an ongoing and unresolved dispute within palaeoanthropology. Proponents of synonymisation typically designate H. ergaster as "African Homo erectus" or "Homo erectus ergaster". The name Homo ergaster roughly translates to "working man", a reference to the more advanced tools used by the species in comparison to those of their ancestors. The fossil range of H. ergaster mainly covers the period of 1.7 to 1.4 million years ago, though a broader time range is possible. Though fossils are known from across East and Southern Africa, most H. ergaster fossils have been found along the shores of Lake Turkana in Kenya. There are later African fossils, some younger than 1 million years ago, that indicate long-term anatomical continuity, though it is unclear if they can be formally regarded as H. ergaster specimens. As a chronospecies, H. ergaster may have persisted to as late as 600,000 years ago, when new lineages of Homo arose in Africa.

<i>Homo rudolfensis</i> Extinct hominin from the Early Pleistocene of East Africa

Homo rudolfensis is an extinct species of archaic human from the Early Pleistocene of East Africa about 2 million years ago (mya). Because H. rudolfensis coexisted with several other hominins, it is debated what specimens can be confidently assigned to this species beyond the lectotype skull KNM-ER 1470 and other partial skull aspects. No bodily remains are definitively assigned to H. rudolfensis. Consequently, both its generic classification and validity are debated without any wide consensus, with some recommending the species to actually belong to the genus Australopithecus as A. rudolfensis or Kenyanthropus as K. rudolfensis, or that it is synonymous with the contemporaneous and anatomically similar H. habilis.

<i>Homo</i> Genus of hominins that includes humans and their closest extinct relatives

Homo is the genus that emerged in the genus Australopithecus that encompasses the extant species Homo sapiens, along with several extinct species classified as either ancestral to or closely related to modern humans, including Homo erectus and Homo neanderthalensis. The oldest member of Homo is Homo habilis with records of just over 2 million years ago. Homo, together with the genus Paranthropus, is probably sister to Australopithecus africanus, which itself had previously split from the lineage of Pan, the chimpanzees.

<span class="mw-page-title-main">Dmanisi</span> Town in Kvemo Kartli, Georgia

Dmanisi is a town and archaeological site in the Kvemo Kartli region of Georgia approximately 93 km southwest of the nation’s capital Tbilisi in the river valley of Mashavera. The hominin site is dated to 1.8 million years ago. It was the earliest known evidence of hominins outside Africa before stone tools dated to 2.1 million years were discovered in 2018 in Shangchen, China.

Marc R. Meyer is an archaeologist and anthropologist who is notable for his excavation of, and research into, the remains of fossil hominids such as Australopithecines and early genus Homo. He currently lectures at Chaffey College, Rancho Cucamonga, CA.

Human taxonomy is the classification of the human species within zoological taxonomy. The systematic genus, Homo, is designed to include both anatomically modern humans and extinct varieties of archaic humans. Current humans have been designated as subspecies Homo sapiens sapiens, differentiated, according to some, from the direct ancestor, Homo sapiens idaltu.

<span class="mw-page-title-main">OH 24</span> Hominin fossil

OH 24 is a fossilized skull of the species Homo habilis. It was discovered in Olduvai Gorge, Tanzania by Peter Nzube in 1968. The skull was found crushed almost flat and was therefore named after the famously skinny model of the time Twiggy. Estimated at 1.8 mya, the cranium was found crushed flat and cemented together with a mass coating of limestone. It is now a Smithsonian exhibit item.

<span class="mw-page-title-main">David Lordkipanidze</span>

David Otaris dze Lordkipanidze is a Georgian anthropologist and archaeologist, Professor (2004), Dr.Sc. (2002), Corresponding Member of the Georgian National Academy of Sciences (2009), since 2004 General Director of the Georgian National Museum (GNM). He is a son of the archaeologist Otar Lordkipanidze.

<span class="mw-page-title-main">Ubeidiya prehistoric site</span> Prehistoric site in the Jordan Valley

'Ubeidiya, some 3 km south of the Sea of Galilee, in the Jordan Rift Valley, Israel, is an archaeological site of the early Pleistocene, c. 1.5 million years ago, preserving traces of one of the earliest migrations of Homo erectus out of Africa, with only the site of Dmanisi in Georgia being older. The site yielded hand axes of the Acheulean type, but very few human remains. The animal remains include a hippopotamus' femur bone, and an immensely large pair of horns belonging to a species of extinct bovid.

<i>Homo erectus</i> Extinct species of archaic human

Homo erectus is an extinct species of archaic human from the Pleistocene, with its earliest occurrence about 2 million years ago. Its specimens are among the first recognizable members of the genus Homo.

<i>Homo gautengensis</i> Name proposed for an extinct species of hominin from South Africa

Homo gautengensis is a species name proposed by anthropologist Darren Curnoe in 2010 for South African hominin fossils otherwise attributed to H. habilis, H. ergaster, or, in some cases, Australopithecus or Paranthropus. The fossils assigned to the species by Curnoe cover a vast temporal range, from about 1.8 million years ago to potentially as late as 0.8 million years ago, meaning that if the species is considered valid, H. gautengensis would be both one of the earliest and one of the longest lived species of Homo.

<span class="mw-page-title-main">Red Deer Cave people</span> Prehistoric humans from 12,500 BCE in southwest China

The Red Deer Cave people were a prehistoric population of modern humans known from bones dated to between about 17,830 to c. 11,500 years ago, found in Red Deer Cave and Longlin Cave, Yunnan Province, in Southwest China.

<span class="mw-page-title-main">Dmanisi skull 3</span> Hominin fossil

D2700, also known as Dmanisi skull 3, is one of five skulls discovered in Dmanisi, Georgia in 2001 and classified as early Homo erectus. It is an almost complete skull and is in an exceptionally good condition. It was dated stratigraphically as about 1.8 million years old.

<span class="mw-page-title-main">Early expansions of hominins out of Africa</span> First hominin expansion into Eurasia (2.1–0.1 Ma)

Several expansions of populations of archaic humans out of Africa and throughout Eurasia took place in the course of the Lower Paleolithic, and into the beginning Middle Paleolithic, between about 2.1 million and 0.2 million years ago (Ma). These expansions are collectively known as Out of Africa I, in contrast to the expansion of Homo sapiens (anatomically modern humans) into Eurasia, which may have begun shortly after 0.2 million years ago.

<span class="mw-page-title-main">SK 847</span> Hominin fossil

SK 847 is the abbreviated designation for the fossilized fragments of a Homo habilis cranium, discovered in South Africa, which was dated to an age between 1.8 and 1.5 million years. This fossil shares morphological traits with the early African Homo erectus, sometimes known as Homo ergaster.

<i>Homo naledi</i> South African archaic human species

Homo naledi is an extinct species of archaic human discovered in 2013 in the Rising Star Cave, Cradle of Humankind, South Africa dating to the Middle Pleistocene 335,000–236,000 years ago. The initial discovery comprises 1,550 specimens, representing 737 different elements, and at least 15 different individuals. Despite this exceptionally high number of specimens, their classification with other Homo remains unclear.

<span class="mw-page-title-main">Dmanisi hominins</span> Hominid species or subspecies discovered in Dmanisi, Georgia

The Dmanisi hominins, Dmanisi people, or Dmanisi man were a population of Early Pleistocene hominins whose fossils have been recovered at Dmanisi, Georgia. The fossils and stone tools recovered at Dmanisi range in age from 1.85–1.77 million years old, making the Dmanisi hominins the earliest well-dated hominin fossils in Eurasia and the best preserved fossils of early Homo from a single site so early in time, though earlier fossils and artifacts have been found in Asia. Though their precise classification is controversial and disputed, the Dmanisi fossils are highly significant within research on early hominin migrations out of Africa. The Dmanisi hominins are known from over a hundred postcranial fossils and five famous well-preserved skulls, referred to as Dmanisi Skulls 1–5.

<span class="mw-page-title-main">Dmanisi historic site</span> Historic site in Kvemo Kartli, Georgia

The Dmanisi historic site is a historic and archaeological site in Georgia, located north of the village of Patara Dmanisi, Dmanisi Municipality, in south-central region of Kvemo Kartli, some 85 km southwest of Tbilisi, Georgia's capital.

References

  1. 1 2 3 4 5 6 7 8 David Lordkipanidze, Marcia S. Ponce de Leòn, Ann Margvelashvili, Yoel Rak, G. Philip Rightmire, Abesalom Vekua, Christoph P. E. Zollikofer (18 October 2013). "A Complete Skull from Dmanisi, Georgia, and the Evolutionary Biology of Early Homo". Science. 342 (6156): 326–331. Bibcode:2013Sci...342..326L. doi:10.1126/science.1238484. PMID   24136960. S2CID   20435482.{{cite journal}}: CS1 maint: multiple names: authors list (link)
  2. 1 2 3 Skull suggests three early human species were one : Nature News & Comment
  3. 1 2 Wilford, John Noble (October 17, 2013). "Skull Fossil Suggests Simpler Human Lineage". The New York Times . Retrieved October 18, 2013.
  4. Watson, Traci (October 17, 2013). "Skull discovery sheds light on human species". USA Today . Retrieved October 18, 2013.
  5. Lahr, M.M., 2010. “Saharan Corridors and Their Role in the Evolutionary Geography of ‘Out of Africa I’”. In: A. Baden et al. (Eds.), Out of Africa I: The First Hominin Colonization of Eurasia. Springer Netherlands, 27–46.
  6. Switek, Brian (17 October 2013). "Beautiful Skull Spurs Debate on Human History". National Geographic. Retrieved 22 September 2014.

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