A genus is a taxonomic rank used in the biological classification of living and fossil organisms, as well as viruses, in biology. In the hierarchy of biological classification, genus comes above species and below family. In binomial nomenclature, the genus name forms the first part of the binomial species name for each species within the genus.
Australopithecus ( OS-trə-lo-PITH-i-kəs; from Latin australis, meaning 'southern', and Greek πίθηκος, meaning 'ape', informal australopithecine or australopith is a 'genus' of hominins. From paleontological and archaeological evidence, the genus Australopithecus apparently evolved in eastern Africa around 4 million years ago before spreading throughout the continent and eventually becoming extinct two million years ago. Australopithecus is not literally extinct as the Kenyanthropus, Paranthropus and Homo genera probably emerged as sister of a late Australopithecus species such as A. Africanus and/or A. Sediba. During that time, a number of australopithecine species emerged, including Australopithecus afarensis, A. africanus, A. anamensis, A. bahrelghazali, A. deyiremeda, A. garhi, and A. sediba.
In taxonomy, Homo sapiens is the only extant human species. The name is Latin for "wise man" and was introduced in 1758 by Carl Linnaeus.
Homo erectus appeared about two million years ago and, in several early migrations, it spread throughout Africa (where it is dubbed Homo ergaster) and Eurasia. It was likely the first human species to live in a hunter-gatherer society and to control fire. An adaptive and successful species, Homo erectus persisted for more than a million years, and gradually diverged into new species by around 500,000 years ago.
Homo erectus is a species of archaic humans that lived throughout most of the Pleistocene geological epoch. Its earliest fossil evidence dates to 1.8 million years ago.
Homo ergaster , also Homo erectus ergaster or African Homo erectus is an extinct chronospecies of the genus Homo that lived in eastern and southern Africa during the early Pleistocene, between about 1.9 million and 1.4 million years ago.
A hunter-gatherer is a human living in a society in which most or all food is obtained by foraging. Hunter-gatherer societies stand in contrast to agricultural societies, which rely mainly on domesticated species.
In the context of the recent African origin of modern humans, the Southern Dispersal scenario refers to the early migration along the southern coast of Asia, from the Arabian peninsula via Persia and India to Southeast Asia and Oceania. Alternative names include the "southern coastal route" or "rapid coastal settlement".
Early human migrations are the earliest migrations and expansions of archaic and modern humans across continents and are believed to have begun approximately 2 million years ago with the out of Africa migration of Homo erectus. This initial migration was followed by other archaic humans including H. heidelbergensis, which lived around 500,000 years ago and was the likely ancestor of both Denisovans and Neanderthals. Early hominids were said to have "crossed land bridges that were eventually covered in water".
Eurasia is the combined continental landmass of Europe and Asia. The term is a portmanteau of its constituent continents. Located primarily in the Northern and Eastern Hemispheres, it is bordered by the Atlantic Ocean to the west, the Pacific Ocean to the east, the Arctic Ocean to the north, and by Africa, the Mediterranean Sea, and the Indian Ocean to the south. The division between Europe and Asia as two different continents is a historical social construct, with no clear physical separation between them; thus, in some parts of the world, Eurasia is recognized as the largest of the six, five, or even four continents on Earth. In geology, Eurasia is often considered as a single rigid megablock. However, the rigidity of Eurasia is debated based on paleomagnetic data.
The Latin noun homō (genitive hominis) means "human being" or "man" in the generic sense of "human being, mankind". The binomial nameHomo sapiens was coined by Carl Linnaeus (1758). Names for other species of the genus were introduced beginning in the second half of the 19th century (H. neanderthalensis 1864, H. erectus 1892).
Latin is a classical language belonging to the Italic branch of the Indo-European languages. The Latin alphabet is derived from the Etruscan and Greek alphabets and ultimately from the Phoenician alphabet.
The term "man" and words derived from it can designate any or even all of the human race regardless of their sex or age. In traditional usage, man itself refers to the species, to humanity, or "mankind", as a whole.
Binomial nomenclature, also called binominal nomenclature or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin grammatical forms, although they can be based on words from other languages. Such a name is called a binomial name, a binomen, binominal name or a scientific name; more informally it is also called a Latin name. The first part of the name – the generic name – identifies the genus to which the species belongs, while the second part – the specific name or specific epithet – identifies the species within the genus. For example, humans belong to the genus Homo and within this genus to the species Homo sapiens. Tyrannosaurus rex is probably the most widely known binomial. The formal introduction of this system of naming species is credited to Carl Linnaeus, effectively beginning with his work Species Plantarum in 1753. But Gaspard Bauhin, in as early as 1623, had introduced in his book Pinax theatri botanici many names of genera that were later adopted by Linnaeus.
Even today, the genus Homo has not been properly defined. Since the early human fossil record began to slowly emerge from the earth, the boundaries and definitions of the genus Homo have been poorly defined and constantly in flux. Because there was no reason to think it would ever have any additional members, Carl Linnaeus did not even bother to define Homo when he first created it for humans in the 18th century. The discovery of Neanderthal brought the first addition.
Carl Linnaeus, also known after his ennoblement as Carl von Linné, was a Swedish botanist, physician, and zoologist who formalised binomial nomenclature, the modern system of naming organisms. He is known as the "father of modern taxonomy". Many of his writings were in Latin, and his name is rendered in Latin as Carolus Linnæus.
The genus Homo was given its taxonomic name to suggest that its member species can be classified as human. And, over the decades of the 20th century, fossil finds of pre-human and early human species from late Miocene and early Pliocene times produced a rich mix for debating classifications. There is continuing debate on delineating Homo from Australopithecus—or, indeed, delineating Homo from Pan, as one body of scientists argue that the two species of chimpanzee should be classed with genus Homo rather than Pan. Even so, classifying the fossils of Homo coincides with evidence of: 1) competent human bipedalism in Homo habilis inherited from the earlier Australopithecus of more than four million years ago, as demonstrated by the Laetoli footprints; and 2) human tool culture having begun by 2.5 million years ago.
The Miocene is the first geological epoch of the Neogene Period and extends from about 23.03 to 5.333 million years ago (Ma). The Miocene was named by Charles Lyell; its name comes from the Greek words μείων and καινός and means "less recent" because it has 18% fewer modern sea invertebrates than the Pliocene. The Miocene is preceded by the Oligocene and is followed by the Pliocene.
The Pliocene Epoch is the epoch in the geologic timescale that extends from 5.333 million to 2.58 million years BP. It is the second and youngest epoch of the Neogene Period in the Cenozoic Era. The Pliocene follows the Miocene Epoch and is followed by the Pleistocene Epoch. Prior to the 2009 revision of the geologic time scale, which placed the four most recent major glaciations entirely within the Pleistocene, the Pliocene also included the Gelasian stage, which lasted from 2.588 to 1.806 million years ago, and is now included in the Pleistocene.
The genus Pan consists of two extant species: the common chimpanzee and the bonobo. Taxonomically, these two ape species are collectively termed panins; however, both species are more commonly referred to collectively using the generalized term chimpanzees, or chimps. Together with humans, gorillas, and orangutans they are part of the family Hominidae. Native to sub-Saharan Africa, common chimpanzees and bonobos are currently both found in the Congo jungle, while only the common chimpanzee is also found further north in West Africa. Both species are listed as endangered on the IUCN Red List of Threatened Species, and in 2017 the Convention on Migratory Species selected the common chimpanzee for special protection.
From the late-19th to mid-20th centuries, a number of new taxonomic names including new generic names were proposed for early human fossils; most have since been merged with Homo in recognition that Homo erectus was a single and singular species with a large geographic spread of early migrations. Many such names are now dubbed as "synonyms" with Homo, including Pithecanthropus,Protanthropus,Sinanthropus,Cyphanthropus,Africanthropus,Telanthropus,Atlanthropus, and Tchadanthropus.
Classifying the genus Homo into species and subspecies is subject to incomplete information and remains poorly done. This has led to using common names ("Neanderthal" and "Denisovan") in even scientific papers to avoid trinomial names or the ambiguity of classifying groups as incertae sedis (uncertain placement)—for example, H. neanderthalensis vs. H. sapiens neanderthalensis, or H. georgicus vs. H. erectus georgicus. Some recently extinct species in the genus Homo are only recently discovered and do not as yet have consensus binomial names (see Denisova hominin and Red Deer Cave people). Since the beginning of the Holocene, it is likely that Homo sapiens (anatomically modern humans) has been the only extant species of Homo.
John Edward Gray (1825) was an early advocate of classifying taxa by designating tribes and families. Wood and Richmond (2000) proposed that Hominini ("hominins") be designated as a tribe that comprised all species of early humans and pre-humans ancestral to humans back to after the chimpanzee-human last common ancestor; and that Hominina be designated a subtribe of Hominini to include only the genus Homo—that is, not including the earlier upright walking hominins of the Pliocene such as Australopithecus, Orrorin tugenensis, Ardipithecus, or Sahelanthropus. Designations alternative to Hominina existed, or were offered: Australopithecinae (Gregory & Hellman 1939) and Preanthropinae (Cela-Conde & Altaba 2002); and later, Cela-Conde and Ayala (2003) proposed that the four genera Australopithecus, Ardipithecus, Praeanthropus, and Sahelanthropus be grouped with Homo within Hominina.[not in citation given]
Especially since the 2010s, the delineation of Homo from Australopithecus has become more contentious. Traditionally, the advent of Homo has been taken to coincide with the first use of stone tools (the Oldowan industry), and thus by definition with the beginning of the Lower Palaeolithic. But in 2010, evidence was presented that seems to attribute the use of stone tools to Australopithecus afarensis around 3.3 million years ago, close to a million years before the first appearance of Homo.LD 350-1, a fossil mandible fragment dated to 2.8 Mya, discovered in 2015 in Afar, Ethiopia, was described as combining "primitive traits seen in early Australopithecus with derived morphology observed in later Homo. Some authors would push the development of Homo close to or even past 3 Mya. Others have voiced doubt as to whether Homo habilis should be included in Homo, proposing an origin of Homo with Homo erectus at roughly 1.9 Mya instead. 
The most salient physiological development between the earlier australopithecine species and Homo is the increase in endocranial volume (ECV), from about 460cm3 (28cuin) in A. garhi to 660cm3 (40cuin) in H. habilis and further to 760cm3 (46cuin) in H. erectus, 1,250cm3 (76cuin) in H. heidelbergensis and up to 1,760cm3 (107cuin) in H. neanderthalensis. However, a steady rise in cranial capacity is observed already in Autralopithecina and does not terminate after the emergence of Homo, so that it does not serve as an objective criterion to define the emergence of the genus.
Homo habilis emerged about 2.1 Mya. Already before 2010, there were suggestions that H. habilis should not be placed in genus Homo but rather in Australopithecus. The main reason to include H. habilis in Homo, its undisputed tool use, has become obsolete with the discovery of Australopithecus tool use at least a million years before H. habilis. Furthermore, H. habilis was long thought to be the ancestor of the more gracile Homo ergaster (Homo erectus). In 2007, it was discovered that H. habilis and H. erectus coexisted for a considerable time, suggesting that H. erectus is not immediately derived from H. habilis but instead from a common ancestor. With the publication of Dmanisi skull 5 in 2013, it has become less certain that Asian H. erectus is a descendant of African H. ergaster which was in turn derived from H. habilis. Instead, H. ergaster and H. erectus appear to be variants of the same species, which may have originated in either Africa or Asia and widely dispersed throughout Eurasia (including Europe, Indonesia, China) by 0.5 Mya.
Homo erectus has often been assumed to have developed anagenetically from Homo habilis from about 2 million years ago. This scenario was strengthened with the discovery of Homo erectus georgicus, early specimens of H. erectus found in the Caucasus, which seemed to exhibit transitional traits with H. habilis. As the earliest evidence for H. erectus was found outside of Africa, it was considered plausible that H. erectus developed in Eurasia and then migrated back to Africa. Based on fossils from the Koobi Fora Formation, east of Lake Turkana in Kenya, Spoor et al. (2007) argued that H. habilis may have survived beyond the emergence of H. erectus, so that the evolution of H. erectus would not have been anagenetically, and H. erectus would have existed alongside H. habilis for about half a million years (1.9to1.4million years ago), during the early Calabrian.
A separate South African species Homo gautengensis has been postulated as contemporary with Homo erectus in 2010.
A taxonomy of the Homo within the great apes is assessed as follows, with Paranthropus and Homo emerging within Australopithecus (shown here cladistically granting Paranthropus, Kenyanthropus, and Homo). The exact phylogeny within Australopithecus is still highly controversial. Approximate radiation dates of daughter clades are shown in Millions of years ago (Mya). Sahelanthropus, Orrorin, and Ardipithecus, possibly sisters to Australopithecus, are not shown here. Note that the naming of groupings is sometimes muddled as often certain groupings are presumed before a cladistic analyses is performed.
Several of the Homo lineages appear to have surviving progeny through introgression into other lines. An archaic lineage separating from the other human lineages 1.5 million years ago, perhaps H. erectus, may have interbred into the Denisovans about 55,000 years ago, although H. erectus is generally regarded as being extinct by then. However, the thigh bone, dated at 14,000 years, found in a Maludong cave (Red Deer Cave people) strongly resembles very ancient species like early Homo erectus or the even more archaic lineage, Homo habilis, which lived around 1.5 million year ago. There is evidence for introgression of H. Heidelbergensis into H. sapiens. The genomes of non-sub-Saharan African humans show what appear to be numerous independent introgression events involving Neanderthal and in some cases also Denisovans around 45,000 years ago. Likewise the genetic structure of sub-Saharan Africans seems to be indicative of introgression from a distinct, as yet unidentified archaic human lineage such as H. heidelbergensis.
Among extant populations of Homo sapiens, the deepest temporal division is found in the San people of Southern Africa, estimated at close to 130,000 years, or possibly more than 300,000 years ago. Temporal division among non-Africans is of the order of 60,000 years in the case of Australo-Melanesians. Division of Europeans and East Asians is of the order of 50,000 years, with repeated and significant admixture events throughout Eurasia during the Holocene.
There has historically been a trend to postulate "new human species" based on as little as an individual fossil. A "minimalist" approach to human taxonomy recognizes at most three species, Homo habilis (2.1–1.5 Mya, membership in Homo questionable), Homo erectus (1.8–0.1 Mya, including the majority of the age of the genus, and the majority of archaic varieties as subspecies, including H. heidelbergensis as a late or transitional variety) and Homo sapiens (300 kya to present, including H. neanderthalensis and other varieties as subspecies).
Human evolution is the evolutionary process that led to the emergence of anatomically modern humans, beginning with the evolutionary history of primates—in particular genus Homo—and leading to the emergence of Homo sapiens as a distinct species of the hominid family, the great apes. This process involved the gradual development of traits such as human bipedalism and language, as well as interbreeding with other hominins, which indicate that human evolution was not linear but a web.
Homininae, also called "African hominids" or "African apes", is a subfamily of Hominidae. It includes two tribes, with their extant as well as extinct species: 1) the Hominini tribe ―and 2) the Gorillini tribe (gorillas). Alternatively, the genus Pan is sometimes considered to belong to its own third tribe, Panini. Homininae comprises all hominids that arose after orangutans split from the line of great apes. The Homininae cladogram has three main branches, which lead to gorillas, and to humans and chimpanzees via the tribe Hominini and subtribes Hominina and Panina. There are two living species of Panina and two living species of gorillas, but only one extant human species. Traces of hypothetical Homo species, including Homo floresiensis and Homo denisova, have been found with dates as recent as 40,000 years ago. Organisms in this subfamily are described as hominine or hominines.
Homo habilis is a proposed archaic species of Homo, which lived between roughly 2.1 and 1.5 million years ago, during the Gelasian and early Calabrian stages of the Pleistocene geological epoch.
Homo rudolfensis is an extinct species of the Hominini tribe, on the morphological boundary between the genera Homo and Australopithecus. Its oldest fossil is dated to 2.4 million years ago, at the very beginning of the Pleistocene, with the possible exception of the LD 350-1 representing the oldest fossil evidence of the emergence of archaic humans from their australopithecine ancestors.
Homo heidelbergensis is an extinct species or subspecies of archaic humans in the genus Homo, which radiated in the Middle Pleistocene from about 700,000 to 300,000 years ago, known from fossils found in Southern Africa, East Africa and Europe. African H. heidelbergensis has several subspecies. The subspecies are Homo heidelbergensis heidelbergensis, Homo heidelbergensis daliensis, Homo rhodesiensis, and Homo heidelbergensis steinheimensi. The derivation of Homo sapiens from Homo rhodesiensis has often been proposed, but is obscured by a fossil gap from 400–260 kya. The species was originally named Homo heidelbergensis due to the skeleton's first discovery near Heidelberg, Germany.
Homo floresiensis is an extinct species in the genus Homo.
The Lower Paleolithic is the earliest subdivision of the Paleolithic or Old Stone Age. It spans the time from around 3.3 million years ago when the first evidence for stone tool production and use by hominins appears in the current archaeological record, until around 300,000 years ago, spanning the Oldowan and Acheulean lithics industries.
Homo antecessor is a proposed archaic human species of the Lower Paleolithic, known to have been present in Western Europe between about 1.2 million and 0.8 million years ago (Mya). It was described in 1997 by Eudald Carbonell, Juan Luis Arsuaga and J. M. Bermúdez de Castro, who based on its "unique mix of modern and primitive traits" classified it as a previously unknown archaic human species.
Human taxonomy is the classification of the human species within zoological taxonomy. The systematic genus, Homo, is designed to include both anatomically modern humans and extinct varieties of archaic humans. Current humans have been designated as subspecies Homo sapiens sapiens, differentiated from the direct ancestor, Homo sapiens idaltu.
A number of varieties of Homo are grouped into the broad category of archaic humans in the period contemporary to and predating the emergence of the earliest anatomically modern humans over 315 ka. The term typically includes Homo neanderthalensis (430+–25 ka), Denisovans, Homo rhodesiensis (300–125 ka), Homo heidelbergensis (600–200 ka), and Homo antecessor.
Paleolithic Europe, the Lower or Old Stone Age in Europe encompasses the era from the arrival of the first archaic humans, about 1.4 million years ago until the beginning of the Mesolithic around 10,000 years ago. This period thus covers over 99% of the total human presence on the European continent. The early arrival and disappearance of Homo erectus and Homo heidelbergensis, the appearance, complete evolution and eventual demise of Homo neanderthalensis and the immigration and successful settlement of Homo sapiens all have taken place during the European Paleolithic.
The Hominidae, whose members are known as great apes or hominids, are a taxonomic family of primates that includes eight extant species in four genera: Pongo, the Bornean, Sumatran and Tapanuli orangutan; Gorilla, the eastern and western gorilla; Pan, the common chimpanzee and the bonobo; and Homo, which includes modern humans and their extinct relatives, and ancestors, such as Homo erectus.
Australopithecus sediba is a species of Australopithecus of the early Pleistocene, identified based on fossil remains dated to about 2 million years ago. The species is known from six skeletons discovered in the Malapa Fossil Site at the Cradle of Humankind World Heritage Site in South Africa, including a juvenile male, an adult female, an adult male, and three infants. The fossils were found together at the bottom of the Malapa Cave, where they apparently fell to their death, and have been dated to between 1.980 and 1.977 million years ago.
Homo gautengensis is a hominin species proposed by biological anthropologist Darren Curnoe in 2010. The species is composed of South African hominin fossils previously attributed to Homo habilis, Homo ergaster, or, in some cases, Australopithecus, and is argued by Curnoe to be the earliest species in the genus Homo.
Several expansions of populations of archaic humans out of Africa and throughout Eurasia took place in the course of the Lower Paleolithic, and into the beginning Middle Paleolithic, between about 2.1 million and 0.2 million years ago (Ma). These expansions are collectively dubbed as Out of Africa I, in contrast to the expansion of Homo sapiens (anatomically modern humans) into Eurasia, which may have begun shortly after 0.2 million years ago.
The diet of known human ancestors varies dramatically over time. Strictly speaking, according to evolutionary anthropologists and archaeologists, there is not a single hominin Paleolithic diet. The Paleolithic covers roughly 2.8 million years, concurrent with the Pleistocene, and includes multiple human ancestors with their own evolutionary and technological adaptations living in a wide variety of environments. This fact with the difficulty of finding conclusive of evidence often makes broad generalizations of the earlier human diets very difficult. Our pre-hominin primate ancestors were broadly herbivorous, relying on either foliage or fruits and nuts and the shift in dietary breadth during the Paleolithic is often considered a critical point in hominin evolution. A generalization between Paleolithic diets of the various human ancestors that many anthropologists do make is that they are all to one degree or another omnivorous and are inextricably linked with tool use and new technologies.
↑ H. erectus in the narrow sense (the Asian species) was extinct by 140,000 years ago, Homo erectus soloensis, found in Java, is considered the latest known survival of H. erectus. Formerly dated to as late as 50,000 to 40,000 years ago, a 2011 study pushed back the date of its extinction of H. e. soloensis to 143,000 years ago at the latest, more likely before 550,000 years ago. Indriati E, Swisher CC III, Lepre C, Quinn RL, Suriyanto RA, et al. 2011 The Age of the 20 Meter Solo River Terrace, Java, Indonesia and the Survival of Homo erectus in Asia.PLoS ONE 6(6): e21562. doi:10.1371/journal.pone.0021562.
↑ Lowery, R.K.; Uribe, G.; Jimenez, E.B.; Weiss, M.A.; Herrera, K.J.; Regueiro, M.; Herrera, R.J. (2013). "Neanderthal and Denisova genetic affinities with contemporary humans: Introgression versus common ancestral polymorphisms". Gene. 530 (1): 83–94. doi:10.1016/j.gene.2013.06.005. PMID23872234. This study raises the possibility of observed genetic affinities between archaic and modern human populations being mostly due to common ancestral polymorphisms.
↑ The word "human" itself is from Latin humanus, an adjective formed on the root of homo, thought to derive from a Proto-Indo-European word for "earth" reconstructed as *dhǵhem-. dhghem The American Heritage Dictionary of the English Language: Fourth Edition. 2000.
↑ Linné, Carl von (1758). Systema naturæ. Regnum animale (10 ed.). Sumptibus Guilielmi Engelmann. pp.18, 20. Retrieved 19 November 2012.. Note: In 1959, Linnaeus was designated as the lectotype for Homo sapiens (Stearn, W T. 1959. "The background of Linnaeus's contributions to the nomenclature and methods of systematic biology", Systematic Zoology 8 (1): 4–22, p. 4) which means that following the nomenclatural rules, Homo sapiens was validly defined as the animal species to which Linnaeus belonged.
↑ "ape-man", from Pithecanthropus erectus (Java Man), Eugène Dubois, Pithecanthropus erectus: eine menschenähnliche Übergangsform aus Java (1894), identified with the Pithecanthropus alalus (i.e. "non-speaking ape-man") hypothesized earlier by Ernst Haeckel
↑ "African man", used by T.F. Dreyer (1935) for the Florisbad Skull he found in 1932 (also Homo florisbadensis or Homo helmei). Also the genus suggested for a number of archaic human skulls found at Lake Eyasi by Weinert (1938). Leaky, Journal of the East Africa Natural History Society' (1942), p. 43.
↑ "remote man"; from Telanthropus capensis (Broom and Robinson 1949), see (1961), p. 487.
↑ from Atlanthropus mauritanicus, name given to the species of fossils (three lower jaw bones and a parietal bone of a skull) discovered in 1954 to 1955 by Camille Arambourg in Tighennif, Algeria. Arambourg, C. (1955). "A recent discovery in human paleontology: Atlanthropus of ternifine (Algeria)". American Journal of Physical Anthropology. 13 (2): 191–201. doi:10.1002/ajpa.1330130203.
↑ Y. Coppens, "L'Hominien du Tchad", Actes V Congr. PPEC I (1965), 329f.; "Le Tchadanthropus", Anthropologia 70 (1966), 5–16.
↑ J.E. Gray, "An outline of an attempt at the disposition of Mammalia into Tribes and Families, with a list of genera apparently appertaining to each Tribe", Annals of Philosophy', new series (1825), pp. 337–344.
1 2 McPherron, S.P.; Alemseged, Z.; Marean, C.W.; Wynn, J.G.; Reed, D.; Geraads, D.; Bobe, R.; Bearat, H.A. (2010). "Evidence for stone-tool-assisted consumption of animal tissues before 3.39 million years ago at Dikika, Ethiopia". Nature. 466 (7308): 857–860. Bibcode:2010Natur.466..857M. doi:10.1038/nature09248. PMID20703305. "The oldest direct evidence of stone tool manufacture comes from Gona (Ethiopia) and dates to between 2.6 and 2.5 million years (Myr) ago. [...] Here we report stone-tool-inflicted marks on bones found during recent survey work in Dikika, Ethiopia [... showing] unambiguous stone-tool cut marks for flesh removal [..., dated] to between 3.42 and 3.24 Myr ago [...] Our discovery extends by approximately 800,000 years the antiquity of stone tools and of stone-tool-assisted consumption of ungulates by hominins; furthermore, this behaviour can now be attributed to Australopithecus afarensis."
↑ "the adaptive coherence of Homo would be compromised if H. habilis is included in Homo. Thus, if these arguments are accepted the origins of the genus Homo are coincident in time and place with the emergence of H. erectus, not H. habilis" Bernard Wood, "Did early Homo migrate 'out of' or 'in to' Africa?", PNAS vol. 108, no.26 (28 June 2011), 10375–10376.
↑ " A fresh look at brain size, hand morphology and earliest technology suggests that a number of key Homo attributes may already be present in generalized species of Australopithecus, and that adaptive distinctions in Homo are simply amplifications or extensions of ancient hominin trends. [...] the adaptive shift represented by the ECV of Australopithecus is at least as significant as the one represented by the ECV of early Homo, and that a major ‘grade-level’ leap in brain size with the advent of H. erectus is probably illusory" William H. Kimbel, Brian Villmoare, "From Australopithecus to Homo: the transition that wasn't", Philosophical Transactions of the Royal Society B, 13 June 2016, DOI: 10.1098/rstb.2015.0248.
↑ by W. Schnaubelt & N. Kieser (Atelier WILD LIFE ART); see Westfalen_in_der-Alt-und_Mittelsteinzeit, Landschaftsverband Westfalen-Lippe, Münster (2013), fig. 42.
↑ F. Spoor; M.G. Leakey; P.N. Gathogo; F.H. Brown; S.C. Antón; I. McDougall; C. Kiarie; F.K. Manthi; L.N. Leakey (2007-08-09). "Implications of new early Homo fossils from Ileret, east of Lake Turkana, Kenya". Nature. 448 (7154): 688–691. Bibcode:2007Natur.448..688S. doi:10.1038/nature05986. PMID17687323.F. Spoor; M.G. Leakey; P.N. Gathogo; F.H. Brown; S.C. Antón; I. McDougall; C. Kiarie; F.K. Manthi; L.N. Leakey (2007-08-09). "Implications of new early Homo fossils from Ileret, east of Lake Turkana, Kenya". Nature. 448 (7154): 688–691. Bibcode:2007Natur.448..688S. doi:10.1038/nature05986. PMID17687323. "A partial maxilla assigned to H. habilis reliably demonstrates that this species survived until later than previously recognized, making an anagenetic relationship with H. erectus unlikely"
↑ "A partial maxilla assigned to H. habilis reliably demonstrates that this species survived until later than previously recognized, making an anagenetic relationship with H. erectus unlikely. The discovery of a particularly small calvaria of H. erectus indicates that this taxon overlapped in size with H. habilis, and may have shown marked sexual dimorphism. The new fossils confirm the distinctiveness of H. habilis and H. erectus, independently of overall cranial size, and suggest that these two early taxa were living broadly sympatrically in the same lake basin for almost half a million years." Spoor, F; Leakey, M.G; Gathogo, P.N; Brown, F.H; Antón, S.C; McDougall, I; Kiarie, C; Manthi, F.K.; Leakey, L.N. (2007). "Implications of new early Homo fossils from Ileret, east of Lake Turkana, Kenya". Nature. 448 (7154): 688–691. Bibcode:2007Natur.448..688S. doi:10.1038/nature05986. PMID17687323.
↑ Curnoe, D (2010). "A review of early Homo in southern Africa focusing on cranial, mandibular and dental remains, with the description of a new species (Homo gautengensis sp. nov.)". HOMO – Journal of Comparative Human Biology. 61 (3): 151–177. doi:10.1016/j.jchb.2010.04.002. PMID20466364.
↑ H. erectus in the narrow sense (the Asian species) was extinct by 140,000 years ago, Homo erectus soloensis, found in Java, is considered the latest known survival of H. erectus. Formerly dated to as late as 50,000 to 40,000 years ago, a 2011 study pushed back the date of its extinction of H. e. soloensis to 143,000 years ago at the latest, more likely before 550,000 years ago. Indriati E, Swisher CC III, Lepre C, Quinn RL, Suriyanto RA, et al. 2011 The Age of the 20 Meter Solo River Terrace, Java, Indonesia and the Survival of Homo erectus in Asia.PLoS ONE 6(6): e21562.doi:10.1371/journal.pone.0021562.
1 2 Zeitoun, Valery (2003). "High occurrence of a basicranial feature in Homo erectus: Anatomical description of the preglenoid tubercle". The Anatomical Record Part B: The New Anatomist. 274B (1): 148–156. doi:10.1002/ar.b.10028. ISSN1552-4914. PMID12964205.
↑ Confirmed H. habilis fossils are dated to between 2.1 and 1.5 million years ago. This date range overlaps with the emergence of Homo erectus. Schrenk, Friedemann; Kullmer, Ottmar; Bromage, Timothy (2007). "The Earliest Putative Homo Fossils". In Henke, Winfried; Tattersall, Ian (eds.). Handbook of Paleoanthropology. 1. In collaboration with Thorolf Hardt. Berlin, Heidelberg: Springer. pp.1611–1631. doi:10.1007/978-3-540-33761-4_52. ISBN978-3-540-32474-4.DiMaggio, Erin N.; Campisano, Christopher J.; Rowan, John; et al. (March 20, 2015). "Late Pliocene fossiliferous sedimentary record and the environmental context of early Homo from Afar, Ethiopia". Science. 347 (6228): 1355–1359. Bibcode:2015Sci...347.1355D. doi:10.1126/science.aaa1415. ISSN0036-8075. PMID25739409. Hominins with "proto-Homo" traits may have lived as early as 2.8 million years ago, as suggested by a fossil jawbone classified as transitional between Australopithecus and Homo discovered in 2015.
↑ Curnoe, Darren (June 2010). "A review of early Homo in southern Africa focusing on cranial, mandibular and dental remains, with the description of a new species (Homo gautengensis sp. nov.)". HOMO – Journal of Comparative Human Biology. 61 (3): 151–177. doi:10.1016/j.jchb.2010.04.002. ISSN0018-442X. PMID20466364. A species proposed in 2010 based on the fossil remains of three individuals dated between 1.9 and 0.6 million years ago. The same fossils were also classified as H. habilis, H. ergaster or Australopithecus by other anthropologists.
↑ Haviland, William A.; Walrath, Dana; Prins, Harald E.L.; McBride, Bunny (2007). Evolution and Prehistory: The Human Challenge (8th ed.). Belmont, CA: Thomson Wadsworth. p.162. ISBN978-0-495-38190-7.H. erectus may have appeared some 2 million years ago. Fossils dated to as much as 1.8 million years ago have been found both in Africa and in Southeast Asia, and the oldest fossils by a narrow margin (1.85 to 1.77 million years ago) were found in the Caucasus, so that it is unclear whether H. erectus emerged in Africa and migrated to Eurasia, or if, conversely, it evolved in Eurasia and migrated back to Africa.
↑ Formerly dated to as late as 50,000 to 40,000 years ago, a 2011 study pushed back the date of its extinction of H. e. soloensis to 143,000 years ago at the latest, more likely before 550,000 years ago. Indriati E, Swisher CC III, Lepre C, Quinn RL, Suriyanto RA, et al. 2011 The Age of the 20 Meter Solo River Terrace, Java, Indonesia and the Survival of Homo erectus in Asia.PLoS ONE 6(6): e21562. doi:10.1371/journal.pone.0021562.
↑ The type fossil is Mauer 1, dated to ca. 0.6 million years ago. The transition from H.heidelbergensis to H.neanderthalensis between 300 and 243 thousand years ago is conventional, and makes use of the fact that there is no known fossil in this period. Examples of H. heidelbergensis are fossils found at Bilzingsleben (also classified as Homo erectus bilzingslebensis).
↑ Muttoni, Giovanni; Scardia, Giancarlo; Kent, Dennis V.; Swisher, Carl C.; Manzi, Giorgio (2009). "Pleistocene magnetochronology of early hominin sites at Ceprano and Fontana Ranuccio, Italy". Earth and Planetary Science Letters. 286 (1–2): 255–268. Bibcode:2009E&PSL.286..255M. doi:10.1016/j.epsl.2009.06.032.
↑ Bischoff, James L.; Shamp, Donald D.; Aramburu, Arantza; et al. (March 2003). "The Sima de los Huesos Hominids Date to Beyond U/Th Equilibrium (>350 kyr) and Perhaps to 400–500 kyr: New Radiometric Dates". Journal of Archaeological Science. 30 (3): 275–280. doi:10.1006/jasc.2002.0834. ISSN0305-4403. The first humans with "proto-Neanderthal traits" lived in Eurasia as early as 0.6 to 0.35 million years ago (classified as H. heidelbergensis, also called a chronospecies because it represents a chronological grouping rather than being based on clear morphological distinctions from either H. erectus or H. neanderthalensis). There is a fossil gap in Europe between 300 and 243 kya, and by convention, fossils younger than 243 kya are called "Neanderthal". D. Dean; J.-J. Hublin; R. Holloway; R. Ziegler (1998). "On the phylogenetic position of the pre-Neandertal specimen from Reilingen, Germany". Journal of Human Evolution. 34 (5). pp.485–508. doi:10.1006/jhev.1998.0214.
↑ Détroit, F.; Mijares, A. S.; Corny, J.; Daver, G.; Zanolli, C.; Dizon, E.; Robles, E.; Grün, R.; Piper, P. J. (2019). "A new species of Homo from the Late Pleistocene of the Philippines". Nature. 568 (7751): 181–186. doi:10.1038/s41586-019-1067-9.