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The following tables give an overview of notable finds of hominin fossils and remains relating to human evolution, beginning with the formation of the tribe Hominini (the divergence of the human and chimpanzee lineages) in the late Miocene, roughly 7 to 8 million years ago.
As there are thousands of fossils, mostly fragmentary, often consisting of single bones or isolated teeth with complete skulls and skeletons rare, this overview is not complete, but shows some of the most important findings. The fossils are arranged by approximate age as determined by radiometric dating and/or incremental dating and the species name represents current consensus; if there is no clear scientific consensus the other possible classifications are indicated.
The early fossils shown are not considered ancestors to Homo sapiens but are closely related to ancestors and are therefore important to the study of the lineage. After 1.5 million years ago (extinction of Paranthropus ), all fossils shown are human (genus Homo ). After 11,500 years ago (11.5 ka, beginning of the Holocene), all fossils shown are Homo sapiens (anatomically modern humans), illustrating recent divergence in the formation of modern human sub-populations.
The chimpanzee–human divergence likely took place during around 10 to 7 million years ago. [1] The list of fossils begins with Graecopithecus , dated some 7.2 million years ago, which may or may not still be ancestral to both the human and the chimpanzee lineage. For the earlier history of the human lineage, see Timeline of human evolution#Hominidae, Hominidae#Phylogeny .
Image | Name | Age (Ma) | Species | Year discovered | Country | Discovered by | Now located at |
---|---|---|---|---|---|---|---|
El Graeco | 7.20 [2] | Graecopithecus | 1944, 2017 | Greece, Bulgaria Site:Pyrgos Vassilissis, Azmaka | Böhme (Tübingen), Spassov (BAS) | Met, Athens; Tübingen, Germany | |
TM 266 (Toumai) | 7.00–6.00 [3] | Sahelanthropus tchadensis | 2001 | Chad Site:Djurab Desert | Michel Brunet, Alain Beauvilain, Fanone Gongdibe, Mahamat Adoum and Ahounta Djimdoumalbaye | N'Djamena (Chad), BEAC | |
BAR 1000'00 | 6.1–5.7 [4] | Orrorin tugenensis | 2000 | Kenya Site:Lukeino | Martin Pickford, Kiptalam Cheboi, Dominique Gommery, Pierre Mein, Brigitte Senut | ||
Trachilos footprints | 6.05 [5] | Made by hominin or hominin-like primate | 2002 | Greece | Gerard D. Gierliński | ||
ALA-VP 1/20 [6] | 5.65±0.150 | Ardipithecus kadabba | 1997 | Ethiopia Site:Middle Awash | Yohannes Haile-Selassie | ||
Human evolution is the evolutionary process within the history of primates that led to the emergence of Homo sapiens as a distinct species of the hominid family that includes all the great apes. This process involved the gradual development of traits such as human bipedalism, dexterity, and complex language, as well as interbreeding with other hominins, indicating that human evolution was not linear but weblike. The study of the origins of humans involves several scientific disciplines, including physical and evolutionary anthropology, paleontology, and genetics; the field is also known by the terms anthropogeny, anthropogenesis, and anthropogony.
Homininae, is a subfamily of the family Hominidae (hominids). This subfamily includes two tribes, Hominini and Gorillini, both having extant species as well as extinct species.
Homo habilis is an extinct species of archaic human from the Early Pleistocene of East and South Africa about 2.3 million years ago to 1.65 million years ago (mya). Upon species description in 1964, H. habilis was highly contested, with many researchers recommending it be synonymised with Australopithecus africanus, the only other early hominin known at the time, but H. habilis received more recognition as time went on and more relevant discoveries were made. By the 1980s, H. habilis was proposed to have been a human ancestor, directly evolving into Homo erectus, which directly led to modern humans. This viewpoint is now debated. Several specimens with insecure species identification were assigned to H. habilis, leading to arguments for splitting, namely into "H. rudolfensis" and "H. gautengensis" of which only the former has received wide support.
Paranthropus is a genus of extinct hominin which contains two widely accepted species: P. robustus and P. boisei. However, the validity of Paranthropus is contested, and it is sometimes considered to be synonymous with Australopithecus. They are also referred to as the robust australopithecines. They lived between approximately 2.9 and 1.2 million years ago (mya) from the end of the Pliocene to the Middle Pleistocene.
Australopithecus (, OS-trə-lə-PITH-i-kəs, -loh-; or is a genus of early hominins that existed in Africa during the Pliocene and Early Pleistocene. The genera Homo, Paranthropus, and Kenyanthropus evolved from some Australopithecus species. Australopithecus is a member of the subtribe Australopithecina, which sometimes also includes Ardipithecus, though the term "australopithecine" is sometimes used to refer only to members of Australopithecus. Species include A. garhi, A. africanus, A. sediba, A. afarensis, A. anamensis, A. bahrelghazali and A. deyiremeda. Debate exists as to whether some Australopithecus species should be reclassified into new genera, or if Paranthropus and Kenyanthropus are synonymous with Australopithecus, in part because of the taxonomic inconsistency.
Homo rudolfensis is an extinct species of archaic human from the Early Pleistocene of East Africa about 2 million years ago (mya). Because H. rudolfensis coexisted with several other hominins, it is debated what specimens can be confidently assigned to this species beyond the lectotype skull KNM-ER 1470 and other partial skull aspects. No bodily remains are definitively assigned to H. rudolfensis. Consequently, both its generic classification and validity are debated without any wide consensus, with some recommending the species to actually belong to the genus Australopithecus as A. rudolfensis or Kenyanthropus as K. rudolfensis, or that it is synonymous with the contemporaneous and anatomically similar H. habilis.
Homo is a genus of great ape that emerged from the genus Australopithecus and encompasses only a single extant species, Homo sapiens, along with a number of extinct species classified as either ancestral or closely related to modern humans; these include Homo erectus and Homo neanderthalensis. The oldest member of the genus is Homo habilis, with records of just over 2 million years ago. Homo, together with the genus Paranthropus, is probably most closely related to the species Australopithecus africanus within Australopithecus. The closest living relatives of Homo are of the genus Pan, with the ancestors of Pan and Homo estimated to have diverged around 5.7-11 million years ago during the Late Miocene.
Homo floresiensis(), also known as "Flores Man" or "Hobbit", is an extinct species of small archaic humans that inhabited the island of Flores, Indonesia, until the arrival of modern humans about 50,000 years ago.
Dmanisi is a town and archaeological site in the Kvemo Kartli region of Georgia approximately 93 km southwest of the nation’s capital Tbilisi in the river valley of Mashavera. The hominin site is dated to 1.8 million years ago. It was the earliest known evidence of hominins outside Africa before stone tools dated to 2.1 million years were discovered in 2018 in Shangchen, China.
Homo rhodesiensis is the species name proposed by Arthur Smith Woodward (1921) to classify Kabwe 1, a Middle Stone Age fossil recovered from Broken Hill mine in Kabwe, Northern Rhodesia. In 2020, the skull was dated to 324,000 to 274,000 years ago. Other similar older specimens also exist.
Paranthropus boisei is a species of australopithecine from the Early Pleistocene of East Africa about 2.5 to 1.15 million years ago. The holotype specimen, OH 5, was discovered by palaeoanthropologist Mary Leakey in 1959 at Olduvai Gorge, Tanzania and described by her husband Louis a month later. It was originally placed into its own genus as "Zinjanthropus boisei", but is now relegated to Paranthropus along with other robust australopithecines. However, it is also argued that Paranthropus is an invalid grouping and synonymous with Australopithecus, so the species is also often classified as Australopithecus boisei.
The Omo remains are a collection of hominin bones discovered between 1967 and 1974 at the Omo Kibish sites near the Omo River, in Omo National Park in south-western Ethiopia. The bones were recovered by a scientific team from the Kenya National Museums directed by Richard Leakey and others. The remains from Kamoya's Hominid Site (KHS) were called Omo I and those from Paul I. Abell's Hominid Site (PHS) were called Omo II.
The Hominini (hominins) form a taxonomic tribe of the subfamily Homininae (hominines). They comprise two extant genera: Homo (humans) and Pan, but in standard usage exclude the genus Gorilla (gorillas), which is grouped separately within subfamily Homininae.
Human taxonomy is the classification of the human species within zoological taxonomy. The systematic genus, Homo, is designed to include both anatomically modern humans and extinct varieties of archaic humans. Current humans have been designated as subspecies Homo sapiens sapiens, differentiated, according to some, from the direct ancestor, Homo sapiens idaltu.
Archaic humans is a broad category denoting all species of the genus Homo that are not Homo sapiens. Among the earliest modern human remains are those from Jebel Irhoud in Morocco, Florisbad in South Africa (259 ka), Omo-Kibish I in southern Ethiopia, and Apidima Cave in Southern Greece. Some examples of archaic humans include H. antecessor (1200–770 ka), H. bodoensis (1200–300 ka), H. heidelbergensis (600–200 ka), Neanderthals, H. rhodesiensis (300–125 ka) and Denisovans.
Dali man is the remains of a late Homo erectus or archaic Homo sapiens who lived in the late-mid Pleistocene epoch. The remains comprise a complete fossilized skull, which was discovered by Liu Shuntang in 1978 in Dali County, Shaanxi Province, China.
Nanjing Man is a specimen of Homo erectus found in China. Large fragments of one male and one female skull and a molar tooth were discovered in 1993 in Hulu Cave on the Tangshan (汤山) hills in Jiangning District, Nanjing. The specimens were found in the Hulu limestone cave at a depth of 60–97 cm by Liu Luhong, a local worker. Dating the fossils yielded an estimated age of 580,000 to 620,000 years old.
Homo erectus is an extinct species of archaic human from the Pleistocene, with its earliest occurrence about 2 million years ago. Its specimens are among the first recognizable members of the genus Homo.
Several expansions of populations of archaic humans out of Africa and throughout Eurasia took place in the course of the Lower Paleolithic, and into the beginning Middle Paleolithic, between about 2.1 million and 0.2 million years ago (Ma). These expansions are collectively known as Out of Africa I, in contrast to the expansion of Homo sapiens (anatomically modern humans) into Eurasia, which may have begun shortly after 0.2 million years ago.
The Dmanisi hominins, Dmanisi people, or Dmanisi man were a population of Early Pleistocene hominins whose fossils have been recovered at Dmanisi, Georgia. The fossils and stone tools recovered at Dmanisi range in age from 1.85 to 1.77 million years old, making the Dmanisi hominins the earliest well-dated hominin fossils in Eurasia and the best preserved fossils of early Homo from a single site so early in time, though earlier fossils and artifacts have been found in Asia. Though their precise classification is controversial and disputed, the Dmanisi fossils are highly significant within research on early hominin migrations out of Africa. The Dmanisi hominins are known from over a hundred postcranial fossils and five famous well-preserved skulls, referred to as Dmanisi Skulls 1–5.
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