Biostratigraphy

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Biostratigraphy is the branch of stratigraphy which focuses on correlating and assigning relative ages of rock strata by using the fossil assemblages contained within them. [1] The primary objective of biostratigraphy is correlation, demonstrating that a particular horizon in one geological section represents the same period of time as another horizon at a different section. Fossils within these strata are useful because sediments of the same age can look completely different, due to local variations in the sedimentary environment. For example, one section might have been made up of clays and marls, while another has more chalky limestones. However, if the fossil species recorded are similar, the two sediments are likely to have been laid down around the same time. Ideally these fossils are used to help identify biozones, as they make up the basic biostratigraphy units, and define geological time periods based upon the fossil species found within each section.

Contents

Basic concepts of biostratigraphic principles were introduced many centuries ago, going as far back as the early 1800s. A Danish scientist and bishop by the name of Nicolas Steno was one of the first geologists to recognize that rock layers correlate to the Law of Superposition. With advancements in science and technology, by the 18th century it began to be accepted that fossils were remains left by species that had become deceased and were then preserved within the rock record. [2] The method was well-established before Charles Darwin explained the mechanism behind it—evolution. [3] Scientists William Smith, George Cuvier, and Alexandre Brongniart came to the conclusion that fossils then indicated a series of chronological events, establishing layers of rock strata as some type of unit, later termed biozone. [4] From here on, scientists began relating the changes in strata and biozones to different geological eras, establishing boundaries and time periods within major faunal changes. By the late 18th century the Cambrian and Carboniferous periods were internationally recognized due to these findings. During the early 20th century, advancements in technology gave scientists the ability to study radioactive decay. Using this methodology, scientists were able to establish geological time, the boundaries of the different eras (Paleozoic, Mesozoic, Cenozoic), as well as Periods (Cambrian, Ordovician, Silurian) through the isotopes found within fossils via radioactive decay. [2] Current 21st century uses of biostratigraphy involve interpretations of age for rock layers, which are primarily used by oil and gas industries for drilling workflows and resource allocations. [5]

The first reef builder is a worldwide index fossil for the Lower Cambrian Archeocyathids.JPG
The first reef builder is a worldwide index fossil for the Lower Cambrian

Fossils as a basis for stratigraphic subdivision

Fossil assemblages were traditionally used to designate the duration of periods. Since a large change in fauna was required to make early stratigraphers create a new period, most of the periods we recognize today are terminated by a major extinction event or faunal turnover.

Concept of stage

A stage is a major subdivision of strata, each systematically following the other each bearing a unique assemblage of fossils. Therefore, stages can be defined as a group of strata containing the same major fossil assemblages. French palaeontologist Alcide d'Orbigny is credited for the invention of this concept. He named stages after geographic localities with particularly good sections of rock strata that bear the characteristic fossils on which the stages are based.

Concept of zone

In 1856 German palaeontologist Albert Oppel introduced the concept of zone (also known as biozones or Oppel zone). A zone includes strata characterized by the overlapping range of fossils. They represent the time between the appearance of species chosen at the base of the zone and the appearance of other species chosen at the base of the next succeeding zone. Oppel's zones are named after a particular distinctive fossil species, called an index fossil. Index fossils are one of the species from the assemblage of species that characterize the zone.

Biostratigraphy uses zones for the most fundamental unit of measurement. The thickness and range of these zones can be a few meters, up to hundreds of meters. They can also range from local to worldwide, as the extent of which they can reach in the horizontal plane relies on tectonic plates and tectonic activity. Two of the tectonic processes that run the risk of changing these zones' ranges are metamorphic folding and subduction. Furthermore, biostratigraphic units are divided into six principal kinds of biozones: Taxon range biozone, [6] Concurrent range biozone, [6] Interval biozone, Lineage biozone, Assemblage biozone, and Abundance biozone.

The Taxon range biozone represents the known stratigraphic and geographic range of occurrence of a single taxon. Concurrent range biozone includes the concurrent, coincident, or overlapping part of the range of two specified taxa. Interval biozones include the strata between two specific biostratigraphic surfaces and can be based on lowest or highest occurrences. Lineage biozones are strata containing species representing a specific segment of an evolutionary lineage. Assemblage biozones are strata that contain a unique association of three or more taxa within it. Abundance biozones are strata in which the abundance of a particular taxon or group of taxa is significantly greater than in the adjacent part of the section.

Index fossils

Amplexograptus, a graptolite index fossil, from the Ordovician near Caney Springs, Tennessee. DiplograptusCaneySprings.jpg
Amplexograptus, a graptolite index fossil, from the Ordovician near Caney Springs, Tennessee.

Index fossils (also known as guide fossils, indicator fossils, or dating fossils) are the fossilized remains or traces of particular plants or animals that are characteristic of a particular span of geologic time or environment, and can be used to identify and date the containing rocks. To be practical, index fossils must have a limited vertical time range, wide geographic distribution, and rapid evolutionary trends. Rock formations separated by great distances but containing the same index fossil species are thereby known to have both formed during the limited time that the species lived.

Index fossils were originally used to define and identify geologic units, then became a basis for defining geologic periods, and then for faunal stages and zones.

Ammonites, graptolites, archeocyathids, inoceramids, and trilobites are groups of animals from which many species have been identified as index fossils that are widely used in biostratigraphy. Species of microfossils such as acritarchs, chitinozoans, conodonts, dinoflagellate cysts, ostracods, pollen, spores and foraminiferans are also frequently used. Different fossils work well for sediments of different ages; trilobites, for example, are particularly useful for sediments of Cambrian age. A long series of ammonite and inoceramid species are particularly useful for correlating environmental events around the world during the super-greenhouse of the Late Cretaceous. [7] [8]

To work well, the fossils used must be widespread geographically, so that they can be found in many different places. They must also be short-lived as a species, so that the period of time during which they could be incorporated in the sediment is relatively narrow. The longer lived the species, the poorer the stratigraphic precision, so fossils that evolve rapidly, such as ammonites, are favored over forms that evolve much more slowly, like nautiloids.

Often biostratigraphic correlations are based on a faunal assemblage, rather than an individual species — this allows greater precision as the time span in which all of the species in the assemblage existed together is narrower than the time spans of any of the members. Furthermore, if only one species is present in a sample, it can mean either that (1) the strata were formed in the known fossil range of that organism; or (2) that the fossil range of the organism was incompletely known, and the strata extend the known fossil range. For instance, the presence of the trace fossil Treptichnus pedum was used to define the base of the Cambrian period, but it has since been found in older strata. [9] If the fossil is easy to preserve and easy to identify, more precise time estimating of the stratigraphic layers is possible.

Faunal succession

Image displaying newly discovered fossil being introduced into the succession sequence. Faunal sucession.jpg
Image displaying newly discovered fossil being introduced into the succession sequence.

The concept of faunal succession was theorized at the beginning of the 19th century by William Smith. When William was studying rock strata he began to recognize that rock outcrops contained a unique collection of fossils. [10] The idea that these distant rock outcrops contained similar fossils allowed for Smith to order rock formations throughout England. With Smith's work on these rock outcrops and mapping around England, he began to notice some beds of rock may contain mostly similar species, however there were also subtle differences within or between these fossil groups. This difference in assemblages that appeared identical at first, lead to the principle of faunal succession, where fossil organisms succeed one another in a definite and determinable order, and therefore any time period can be categorized by its fossil extent. [11]

See also

Related Research Articles

In chronostratigraphy, a stage is a succession of rock strata laid down in a single age on the geologic timescale, which usually represents millions of years of deposition. A given stage of rock and the corresponding age of time will by convention have the same name, and the same boundaries.

<span class="mw-page-title-main">Stratigraphy</span> Study of rock layers and their formation

Stratigraphy is a branch of geology concerned with the study of rock layers (strata) and layering (stratification). It is primarily used in the study of sedimentary and layered volcanic rocks. Stratigraphy has three related subfields: lithostratigraphy, biostratigraphy, and chronostratigraphy.

<span class="mw-page-title-main">Geochronology</span> Science of determining the age of rocks, sediments and fossils

Geochronology is the science of determining the age of rocks, fossils, and sediments using signatures inherent in the rocks themselves. Absolute geochronology can be accomplished through radioactive isotopes, whereas relative geochronology is provided by tools such as paleomagnetism and stable isotope ratios. By combining multiple geochronological indicators the precision of the recovered age can be improved.

A stratigraphic unit is a volume of rock of identifiable origin and relative age range that is defined by the distinctive and dominant, easily mapped and recognizable petrographic, lithologic or paleontologic features (facies) that characterize it.

<span class="mw-page-title-main">Turonian</span> Second age of the Late Cretaceous epoch

The Turonian is, in the ICS' geologic timescale, the second age in the Late Cretaceous Epoch, or a stage in the Upper Cretaceous Series. It spans the time between 93.9 ± 0.8 Ma and 89.8 ± 1 Ma. The Turonian is preceded by the Cenomanian Stage and underlies the Coniacian Stage.

<span class="mw-page-title-main">Biozone</span>

In biostratigraphy, biostratigraphic units or biozones are intervals of geological strata that are defined on the basis of their characteristic fossil taxa, as opposed to a lithostratigraphic unit which is defined by the lithological properties of the surrounding rock.

<i>Tapinocephalus</i> Assemblage Zone

The Tapinocephalus Assemblage Zone is a tetrapod assemblage zone or biozone which correlates to the middle Abrahamskraal Formation, Adelaide Subgroup of the Beaufort Group, a fossiliferous and geologically important geological Group of the Karoo Supergroup in South Africa. The thickest outcrops, reaching approximately 2,000 metres (6,600 ft), occur from Merweville and Leeu-Gamka in its southernmost exposures, from Sutherland through to Beaufort West where outcrops start to only be found in the south-east, north of Oudshoorn and Willowmore, reaching up to areas south of Graaff-Reinet. Its northernmost exposures occur around the towns Fraserburg and Victoria West. The Tapinocephalus Assemblage Zone is the second biozone of the Beaufort Group.

<i>Cistecephalus</i> Assemblage Zone

The Cistecephalus Assemblage Zone is a tetrapod assemblage zone or biozone found in the Adelaide Subgroup of the Beaufort Group, a majorly fossiliferous and geologically important geological group of the Karoo Supergroup in South Africa. This biozone has outcrops located in the Teekloof Formation north-west of Beaufort West in the Western Cape, in the upper Middleton and lower Balfour Formations respectively from Colesberg of the Northern Cape to east of Graaff-Reinet in the Eastern Cape. The Cistecephalus Assemblage Zone is one of eight biozones found in the Beaufort Group, and is considered to be Late Permian in age.

<i>Daptocephalus</i> Assemblage Zone

The Daptocephalus Assemblage Zone is a tetrapod assemblage zone or biozone found in the Adelaide Subgroup of the Beaufort Group, a majorly fossiliferous and geologically important geological Group of the Karoo Supergroup in South Africa. This biozone has outcrops located in the upper Teekloof Formation west of 24°E, the majority of the Balfour Formation east of 24°E, and the Normandien Formation in the north. It has numerous localities which are spread out from Colesberg in the Northern Cape, Graaff-Reniet to Mthatha in the Eastern Cape, and from Bloemfontein to Harrismith in the Free State. The Daptocephalus Assemblage Zone is one of eight biozones found in the Beaufort Group and is considered Late Permian (Lopingian) in age. Its contact with the overlying Lystrosaurus Assemblage Zone marks the Permian-Triassic boundary.

<i>Pristerognathus</i> Assemblage Zone

The Pristerognathus Assemblage Zone is a tetrapod assemblage zone or biozone which correlates to the upper Abrahamskraal Formation and lowermost Teekloof Formation, Adelaide Subgroup of the Beaufort Group, a fossiliferous and geologically important geological Group of the Karoo Supergroup in South Africa. The thickest outcrops, reaching not more than 300 metres (980 ft), occur just east of Sutherland through to Beaufort West in the south and Victoria West in the north. Exposures are also found west of Colesberg and south of Graaff-Reinet. The Pristerognathus Assemblage Zone is the third biozone of the Beaufort Group.

<i>Tropidostoma</i> Assemblage Zone

The Tropidostoma Assemblage Zone is a tetrapod assemblage zone or biozone which correlates to the lower Teekloof Formation, Adelaide Subgroup of the Beaufort Group, a fossiliferous and geologically important geological Group of the Karoo Supergroup in South Africa. The thickest outcrops, reaching approximately 240 metres (790 ft), occur from east of Sutherland through to Beaufort West and Victoria West, to areas south of Graaff-Reinet. Its northernmost exposures occur west/north-west of Colesberg. The Tropidostoma Assemblage Zone is the fourth biozone of the Beaufort Group.

<i>Eodicynodon</i> Assemblage Zone

The Eodicynodon Assemblage Zone is a tetrapod assemblage zone or biozone which correlates to the Abrahamskraal Formation, Adelaide Subgroup of the Beaufort Group, a fossiliferous and geologically important geological Group of the Karoo Supergroup in South Africa. The thickest outcrops, reaching approximately 620 metres (2,030 ft), occur south-east of Sutherland, north of Prince Albert, and south-east of Beaufort West. The Eodicynodon Assemblage Zone is the lowermost biozone of the Beaufort Group.

In archaeology and paleontology a faunal assemblage is a group of associated animal fossils found together in a given stratum.

A chronozone or chron is a unit in chronostratigraphy, defined by events such as geomagnetic reversals (magnetozones), or based on the presence of specific fossils . According to the International Commission on Stratigraphy, the term "chronozone" refers to the rocks formed during a particular time period, while "chron" refers to that time period.

In paleontology, biochronology is the correlation in time of biological events using fossils. In its strict sense, it refers to the use of assemblages of fossils that are not tied to stratigraphic sections. Collections of land mammal ages have been defined for every continent except Antarctica, and most are correlated with each other indirectly through known evolutionary lineages. A combination of argon–argon dating and magnetic stratigraphy allows a direct temporal comparison of terrestrial events with climate variations and mass extinctions.

<i>Xystridura</i>

Xystridura Whitehouse, 1936 is a genus of Redlichiid trilobite described originally from the Middle Cambrian (Miaolingian) strata of Queensland, Australia. The genus ranges upwards from Ordian into Florian Stages in terms of Australian Cambrian stratigraphic nomenclature.

The European Land Mammal Mega Zones are zones in rock layers that have a specific assemblage of fossils (biozones) based on occurrences of fossil assemblages of European land mammals. These biozones cover most of the Cenozoic, with particular focus having been paid to the Neogene and Paleogene systems, the Quaternary has several competing systems. In cases when fossils of mammals are abundant, stratigraphers and paleontologists can use these biozones as a more practical regional alternative to the stages of the official ICS geologic timescale. European Land Mammal Mega Zones are often also confusingly referred to as ages, stages, or intervals.

Conodonts are an extinct class of animals whose feeding apparatuses called teeth or elements are common microfossils found in strata dating from the Stage 10 of the Furongian, the fourth and final series of the Cambrian, to the Rhaetian stage of the Late Triassic. These elements can be used alternatively to or in correlation with other types of fossils in the subfield of the stratigraphy named biostratigraphy.

<i>Bailiaspis</i> Genus of trilobites

Bailiaspis Resser, 1936, is a Middle Cambrian (Miaolingian) trilobite genus belonging to the Family Conocoryphidae Angelin, 1854. Within the Acado-Baltic region, the genus ranges from Wuliuan into Guzhangian age strata.

Land vertebrate faunachrons (LVFs) are biochronological units used to correlate and date terrestrial sediments and fossils based on their tetrapod faunas. First formulated on a global scale by Spencer G. Lucas in 1998, LVFs are primarily used within the Triassic Period, though Lucas later designated LVFs for other periods as well. Eight worldwide LVFs are defined for the Triassic. The first two of these LVFs, the Lootsbergian and Nonesian, are based on South African synapsids and faunal assemblage zones estimated to correspond to the Early Triassic. These are followed by the Perovkan and Berdyankian, based on temnospondyl amphibians and Russian assemblages estimated to be from the Middle Triassic. The last four LVFs, the Otischalkian, Adamanian, Revueltian, and Apachean, are based on aetosaur and phytosaur reptiles common in the Late Triassic of the southwestern United States.

References

  1. ↑ Hine, Robert. "Biostratigraphy." Oxford Reference: Dictionary of Biology, 8th ed., Oxford University Press, 2019.
  2. 1 2 Gon, S. M. "Trilobite Biostratigraphy." Edited by Nicolas Tormo, Trilobite Biostratigraphy, 4 Sept. 2018, www.trilobites.info/biostratigraphy.htm
  3. ↑ Gluyas, J. & Swarbrick, R. (2004) Petroleum Geoscience. Publ. Blackwell Publishing. pp. 80-82
  4. ↑ Young, Keith (March 1960). "Biostratigraphy and the New Paleontology". Journal of Paleontology. 34: 347–348 – via JSTOR.
  5. ↑ Simmons, Mike. (2019). ResearchGate, Biostratigraphy in Exploration. Retrieved March 5, 2020. URL: https://www.researchgate.net/publication/332188386_Biostratigraphy_in_Exploration
  6. 1 2 Subcommission on Quaternary Stratigraphy: Stratigraphic guide – Biostratigraphy". Quaternary Stratigraphy. International Union of Geological Sciences (IGUS); International Commission on Stratigraphy (ICS). Retrieved 2020-03-21.
  7. ↑ Van Helmond, Niels A. G. M.; Sluijs, Appy; Papadomanolaki, Nina M.; Plint, A. Guy; Gröcke, Darren R.; Pearce, Martin A.; Eldrett, James S.; Trabucho-Alexandre, João; Walaszczyk, Ireneusz; Van De Schootbrugge, Bas; Brinkhuis, Henk (2018). "Equatorward phytoplankton migration during a cold spell within the Late Cretaceous super-greenhouse". Biogeosciences. 13 (9): 2859. Bibcode:2016BGeo...13.2859V. doi: 10.5194/bg-13-2859-2016 . Retrieved 2020-12-27.
  8. ↑ Ireneusz Walaszczyk; William James Kennedy; Amruta R Paranjape. "Inoceramids and associated ammonite faunas from the uppermost Turonian−lower Coniacian (Upper Cretaceous) of the Anaipadi-Saradamangalam region of the Cauvery Basin, south-east India". Acta Geologica Polonica. OCLC   1151203029 . Retrieved 2020-12-27.
  9. ↑ Gehling, James; Jensen, Sören; Droser, Mary; Myrow, Paul; Narbonne, Guy (March 2001). "Burrowing below the basal Cambrian GSSP, Fortune Head, Newfoundland". Geological Magazine. 138 (2): 213–218. doi : 10.1017/S001675680100509X. 1.
  10. ↑ Strata Smith: The Man & The Map , retrieved 2022-09-26
  11. ↑ Scott, Michon (May 8, 2008). "William Smith: Discovering Faunal Succession & Faunal Succession's Legacy". Earth Observatory.[ page needed ][ ISBN missing ]
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