Homo rhodesiensis Temporal range: Middle Pleistocene | |
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Kabwe skull (1922 photograph) | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Mammalia |
Order: | Primates |
Suborder: | Haplorhini |
Infraorder: | Simiiformes |
Family: | Hominidae |
Subfamily: | Homininae |
Tribe: | Hominini |
Subtribe: | Hominina |
Genus: | Homo |
Species: | †H. rhodesiensis |
Binomial name | |
†Homo rhodesiensis Woodward, 1921 | |
Homo rhodesiensis is the species name proposed by Arthur Smith Woodward (1921) to classify Kabwe 1 (the "Kabwe skull" or "Broken Hill skull", also "Rhodesian Man"), a Middle Stone Age fossil recovered from Broken Hill mine in Kabwe, Northern Rhodesia (now Zambia). [1] In 2020, the skull was dated to 324,000 to 274,000 years ago. Other similar older specimens also exist. [2]
H. rhodesiensis is now widely considered a synonym of H. heidelbergensis . [3] Other designations such as Homo sapiens arcaicus [4] and Homo sapiens rhodesiensis [5] have also been proposed.
A number of morphologically comparable fossil remains came to light in East Africa (Bodo, Ndutu, Eyasi, Ileret) and North Africa (Salé, Rabat, Dar-es-Soltane, Djbel Irhoud, Sidi Aberrahaman, Tighenif) during the 20th century. [6]
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The Bodo cranium [20] is a fossil of an extinct type of hominin species. It was found by members of an expedition led by Jon Kalb in 1976. [21] The Rift Valley Research Mission conducted a number of surveys that led to the findings of Acheulean tools and animal fossils, as well as the Bodo Cranium. [22] The initial discovery was by Alemayhew Asfaw and Charles Smart, who found a lower face. Two weeks later, Paul Whitehead and Craig Wood found the upper portion of the face. Pieces of the cranium were discovered along the surface of one of the dry branches of the Awash River in Ethiopia. [20] The cranium, artifacts, and other animal fossils were found over a relatively large area of medium sand, and only a few of the tools were found near the cranium. [23] [24] The skull is 600,000 years old. [25]
This specimen has an unusually large cranial capacity for its age that is estimated at around 1250 cc (in the range between ~1,200–1,325 cc) within the (lower) range of modern Homo sapiens. [26] The cranium includes the face, much of the frontal bone, parts of the midvault and the base anterior to the foramen magnum. The cranial length, width and height are 21 cm (8.3 in), 15.87 cm (6.2 in) and 19.05 cm (7.5 in) respectively. Researchers have suggested that Bodo butchered animals because Acheulean hand axes and cleavers, along with animal bones, were found at the site. Cuts on the Bodo cranium show the earliest evidence of removal of flesh immediately after the death of an individual using a stone tool. [23] The findings of symmetrical cut marks with specific patterns and directionality on the cranium serve as strong evidence that de-fleshing was done purposefully for mortuary practices and represents the earliest evidence of non-utilitarian mortuary practices. [23] [27] The cut marks were located "laterally among the maxilla" causing speculation among researchers that the specific reason for de-fleshing was to remove the mandible. [28]
The front of the Bodo cranium is very broad and supports large supraorbital structures. The supraorbital torus projects and is heavily constructed, especially in the central parts of the cranium. The Glabella is rounded and projects strongly. Like Homo erectus, the braincase is low and archaic in appearance. The vault bones are also thick like Homo erectus specimens. Due to the large cranial capacity, there is a wider midvault which includes signs of parietal bossing as well as a high contour of the temporal squama. The parietal length can’t be accurately determined because that section of the specimen is incomplete. Though the mastoid is missing, insights regarding the specimen can be determined using fragments from the individual collected at the scene in 1981. The cranium’s parietal walls expand relative to the bitemporal width in a way that is characteristic of modern humans. The squamosal suture has a high arch which is present in modern human craniums as well. [29]
The cranium has an unusual appearance, which has led to debates over its taxonomy. It displays both primitive and derived features, such as a cranial capacity more similar to modern humans and a projecting supraorbital torus more like Homo erectus. [20] [30] [31] Bodo and other Mid-Pleistocene hominin fossils appear to represent a lineage between Homo erectus and anatomically modern humans, although its exact location in the human evolutionary tree is still uncertain. [32] [33] Due to the similarities to both Homo erectus and modern humans, it has been postulated that the Bodo cranium, as well as other members of Homo heidelbergensis were part of a group of hominins that evolved distinct from Homo erectus early in the Middle Pleistocene. Despite the similarities, there is still a question of where exactly Homo heidelbergensis evolved. The increased encephalization seen in fossils like the Bodo cranium is thought to have been a driving force in the speciation of anatomically modern humans. [34] [35]
Both the Bodo cranium and the Kabwe cranium share a number of similarities. Both have cranial capacities similar to, but on the low end of the range of modern humans (1250cc vs 1230cc). Both craniums have a very large supraorbital torus. These two features together suggest that they are a link between Homo erectus and Homo sapiens . [36] The morphology and the taxonomy are most similar to other specimens of type Homo heidelbergensis. [37] Both the Bodo and Kabwe specimens can be described as archaic because they retain certain features in common with Homo erectus. However, both exhibit important differences from Homo erectus in their anatomy, such as the contour of their parietals, the shape of their temporal bones, the cranial base, and the morphology of their nose and palate. While there are many similarities, there are a few differences between the specimens, including the entire brow of the Bodo cranium, particularly the lateral segments, which are less thick than the Kabwe specimen. [29]
In 2021, Canadian anthropologist Mirjana Roksandic and colleagues recommended the complete dissolution of H. heidelbergensis and "H. rhodesiensis", as the name rhodesiensis honours English diamond magnate Cecil Rhodes who disenfranchised the black population in southern Africa. They classified all European H. heidelbergensis as H. neanderthalensis, and synonymised H. rhodesiensis with a new species they named "H. bodoensis" which includes all African specimens, and potentially some from the Levant and the Balkans which have no Neanderthal-derived traits (namely Ceprano, Mala Balanica, HaZore'a and Nadaouiyeh Aïn Askar). "H. bodoensis" is supposed to represent the immediate ancestor of modern humans, but does not include the LCA of modern humans and Neanderthals. They suggested the confusing morphology of the Middle Pleistocene was caused by periodic "H. bodoensis" migration events into Europe following population collapses after glacial cycles, interbreeding with surviving indigenous populations. [38] Their taxonomic recommendations were rejected by Stringer and others as they failed to explain how exactly their proposals would resolve anything, in addition to violating nomenclatural rules. [39] [40]
Homo heidelbergensis is an extinct species or subspecies of archaic human which existed during the Middle Pleistocene. It was subsumed as a subspecies of H. erectus in 1950 as H. e. heidelbergensis, but towards the end of the century, it was more widely classified as its own species. It is debated whether or not to constrain H. heidelbergensis to only Europe or to also include African and Asian specimens, and this is further confounded by the type specimen being a jawbone, because jawbones feature few diagnostic traits and are generally missing among Middle Pleistocene specimens. Thus, it is debated if some of these specimens could be split off into their own species or a subspecies of H. erectus. Because the classification is so disputed, the Middle Pleistocene is often called the "muddle in the middle".
Meganthropus is an extinct genus of non-hominin hominid ape, known from the Pleistocene of Indonesia. It is known from a series of large jaw and skull fragments found at the Sangiran site near Surakarta in Central Java, Indonesia, alongside several isolated teeth. The genus has a long and convoluted taxonomic history. The original fossils were ascribed to a new species, Meganthropus palaeojavanicus, and for a long time was considered invalid, with the genus name being used as an informal name for the fossils.
Homo is a genus of Hominidae that emerged from the genus Australopithecus and encompasses the extant species Homo sapiens and a number of extinct species classified as either ancestral or closely related to modern humans. These include Homo erectus and Homo neanderthalensis. The oldest member of the genus is Homo habilis, with records of just over 2 million years ago. Homo, together with the genus Paranthropus, is probably most closely related to the species Australopithecus africanus within Australopithecus. The closest living relatives of Homo are of the genus Pan, with the ancestors of Pan and Homo estimated to have diverged around 5.7-11 million years ago during the Late Miocene.
Ceprano Man, Argil, and Ceprano Calvarium, is a Middle Pleistocene archaic human fossil, a single skull cap (calvarium), accidentally unearthed in a highway construction project in 1994 near Ceprano in the Province of Frosinone, Italy. It was initially considered Homo cepranensis, Homo erectus, or possibly Homo antecessor; but in recent studies, most regard it either as a form of Homo heidelbergensis sharing affinities with African forms, or an early morph of Neanderthal.
Maba Man is a pre-modern hominin whose remains were discovered in 1958 in caves near the town called Maba, near Shaoguan city in the northern part of Guangdong province, China.
Human taxonomy is the classification of the human species within zoological taxonomy. The systematic genus, Homo, is designed to include both anatomically modern humans and extinct varieties of archaic humans. Current humans have been designated as subspecies Homo sapiens sapiens, differentiated, according to some, from the direct ancestor, Homo sapiens idaltu.
Archaic humans is a broad category denoting all species of the genus Homo that are not Homo sapiens. Among the earliest modern human remains are those from Jebel Irhoud in Morocco, Florisbad in South Africa (259 ka),, Omo-Kibish I in southern Ethiopia ., and Apidima Cave in Southern Greece. Some examples of archaic humans include H. antecessor (1200–770 ka), H. bodoensis (1200–300 ka), H. heidelbergensis (600–200 ka), Neanderthals, H. rhodesiensis (300–125 ka) and Denisovans.
Dali man is the remains of a late Homo erectus or archaic Homo sapiens who lived in the late-mid Pleistocene epoch. The remains comprise a complete fossilized skull, which was discovered by Liu Shuntang in 1978 in Dali County, Shaanxi Province, China.
Nanjing Man is a specimen of Homo erectus found in China. Large fragments of one male and one female skull and a molar tooth of were discovered in 1993 in Hulu Cave on the Tangshan (汤山) hills in Jiangning District, Nanjing. The specimens were found in the Hulu limestone cave at a depth of 60–97 cm by Liu Luhong, a local worker. Dating the fossils yielded an estimated age of 580,000 to 620,000 years old.
Homo erectus is an extinct species of archaic human from the Pleistocene, with its earliest occurrence about 2 million years ago. Its specimens are among the first recognizable members of the genus Homo.
Saldanha man also known as Saldanha cranium or Elandsfontein cranium are fossilized remains of an archaic human. It is one of the key specimens for Homo heidelbergensis. It has not been dated directly, and is estimated to be roughly 0.5 million years old. The remains, which included a fragment of lower jaw, were found on an exposed surface between shifting sand dunes on the farm Elandsfontein, which is located near Hopefield, South Africa.
Samu is the nickname given to a fragmentary Middle Pleistocene human occipital, also known as Vertesszolos Man or Vertesszolos occipital, discovered in Vértesszőlős, Central Transdanubia, Hungary.
The Florisbad Skull is an important human fossil of the early Middle Stone Age, representing either late Homo heidelbergensis or early Homo sapiens. It was discovered in 1932 by T. F. Dreyer at the Florisbad site, Free State Province, South Africa.
The Daka calvaria, otherwise known as the Dakaskull, or specimen number BOU-VP-2/66, is a Homo erectus specimen from the Daka Member of the Bouri Formation in the Middle Awash Study Area of the Ethiopian Rift Valley.
The Ndutu skull is the partial cranium of a hominin that has been assigned variously to late Homo erectus, Homo rhodesiensis, and early Homo sapiens, from the Middle Pleistocene, found at Lake Ndutu in northern Tanzania.
Olduvai Hominid number 9, known as the Chellean Man, is a fossilized skull cap of an early hominin, found in LLK II, Olduvai Gorge by Louis S. B. Leakey in 1960. It is believed to be ca. 1.4 million years old. Its cranial capacity is estimated at than 1067 cm3, the largest value among all known African Homo erectus specimens. OH 9 is significant because of the features it carried and its correlation to the species classification it resides in.
Kabwe 1, also known as the Broken Hill skull and Rhodesian Man, is a Middle Paleolithic fossil assigned by Arthur Smith Woodward in 1921 as the type specimen for Homo rhodesiensis, now mostly considered a synonym of Homo heidelbergensis.
The Dmanisi hominins, Dmanisi people, or Dmanisi man were a population of Early Pleistocene hominins whose fossils have been recovered at Dmanisi, Georgia. The fossils and stone tools recovered at Dmanisi range in age from 1.85 to 1.77 million years old, making the Dmanisi hominins the earliest well-dated hominin fossils in Eurasia and the best preserved fossils of early Homo from a single site so early in time, though earlier fossils and artifacts have been found in Asia. Though their precise classification is controversial and disputed, the Dmanisi fossils are highly significant within research on early hominin migrations out of Africa. The Dmanisi hominins are known from over a hundred postcranial fossils and five famous well-preserved skulls, referred to as Dmanisi Skulls 1–5.
The Salé cranium is a pathological specimen of enigmatic Middle Pleistocene hominin discovered from Salé, Morocco by quarrymen in 1971. Since its discovery, the specimen has variously been classified as Homo sapiens, Homo erectus, Homo rhodesiensis/bodoensis, or Homo heidelbergensis. Its pathological condition and mosaic anatomy has proved difficult to classify. It was discovered with few faunal fossils and no lithics, tentatively dated to 400 ka by some sources.
The Narmada Human, originally the Narmada Man, is a species of extinct human that lived in central India during the Middle and Late Pleistocene. From a skull cup discovered from the bank of the Narmada River in Madhya Pradesh in 1982, the discoverer, Arun Sonakia classified it was an archaic human and gave the name Narmada Man, with the scientific name H. erectus narmadensis. Analysis of additional fossils from the same location in 1997 indicated that the individual could be a female, hence, a revised name, Narmada Human, was introduced. It remains the oldest human species in India.
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