Homininae, also called "African hominids" or "African apes", is a subfamily of Hominidae. It includes two tribes, with their extant as well as extinct species: 1) the tribe Hominini ―and 2) the tribe Gorillini (gorillas). Alternatively, the genus Pan is sometimes considered to belong to its own third tribe, Panini. Homininae comprises all hominids that arose after orangutans split from the line of great apes. The Homininae cladogram has three main branches, which lead to gorillas, and to humans and chimpanzees via the tribe Hominini and subtribes Hominina and Panina. There are two living species of Panina and two living species of gorillas, but only one extant human species. Traces of extinct Homo species, including Homo floresiensis and Homo denisova, have been found with dates as recent as 40,000 years ago. Organisms in this subfamily are described as hominine or hominines.
Orrorin tugenensis is a postulated early species of Homininae, estimated at 6.1 to 5.7 million years ago and discovered in 2000. It is not confirmed how Orrorin is related to modern humans. Its discovery was used to argue against the hypothesis that australopithecines are human ancestors, although this remains the most prevalent hypothesis of human evolution as of 2012.
The parvorder Catarrhini, catarrhine monkeys, Old World anthropoids or Old World monkeys in the broader sense consist of the Old World monkeys in the stricter sense (Cercopithecoidea) and the apes (Hominoidea). In 1812, Geoffroy grouped those two groups together and established the name Catarrhini, "Old World monkeys",. There has been some resistance to directly designate apes as monkeys despite the scientific evidence, so "Old World monkey" may be taken to mean the Cercopithecoidea or the Catarrhini. That apes are monkeys was already realized by Georges-Louis Leclerc, Comte de Buffon in the 18th century. The Catarrhini are all native to Africa and Asia. Members of this parvorder are called catarrhines.
Pierolapithecus catalaunicus is an extinct species of primate which lived about 13 million years ago during the Miocene in what is now Hostalets de Pierola, Catalonia, Spain, giving it its scientific name. It is believed by some to be a common ancestor of both modern humans and the other great apes, or at least a species that is closer to a common ancestor than any previous fossil discovery.
Mesopithecus is an extinct genus of Old World monkey that lived in Europe and Asia 7 to 5 million years ago. Mesopithecus resembled a modern macaque, with a body length of about 40 centimetres (16 in). It was adapted to both walking and climbing, possessing a slender body with long, muscular limbs and flexible fingers. Its teeth suggest that it primarily ate soft leaves and fruit. It was once thought that these extinct monkeys might be an ancestor of the grey langur, but a more recent study suggests that they are more closely related to the snub-nosed monkeys and doucs.
Chororapithecus is an extinct great ape from the Afar region of Ethiopia roughly 8 million years ago during the Late Miocene, comprising one species, C. abyssinicus. It is known from 9 isolated teeth discovered in a 2005–2007 survey of the Chorora Formation. The teeth are indistinguishable from those of gorillas in terms of absolute size and relative proportions, and it has been proposed to be an early member of Gorillini. However, this is controversial given the paucity of remains, and notable anatomical differences between Chororapithecus and gorilla teeth. The Kenyan ape Nakalipithecus has been proposed to be an ancestor of Chororapithecus or at least closely related. If correct, they would be the only identified fossil members of any modern non-human great ape lineage, and would push the gorilla–human last common ancestor from 8 million years ago to 10 million years ago. The teeth are adapted for processing tough plant fibres as well as hard, brittle food, and the formation is thought to represent a forested lakeside habitat.
Nakalipithecus nakayamai is an extinct species of great ape from Nakali, Kenya, from about 9.9–9.8 million years ago during the Late Miocene. It is known from a right jawbone with 3 molars and from 11 isolated teeth, and the specimen is presumed female as the teeth are similar in size to those of female gorillas and orangutans. Compared to other great apes, the canines are short, the enamel is thin, and the molars are flatter. Nakalipithecus is one of only three Late Miocene great apes known from Africa, the others being Samburupithecus and Chororapithecus. Nakalipithecus seems to have inhabited a sclerophyllous woodland environment.
Ardipithecus ramidus is a species of australopithecine from the Afar region of Early Pliocene Ethiopia 4.4 million years ago (mya). A. ramidus, unlike modern hominids, has adaptations for both walking on two legs (bipedality) and life in the trees (arboreality). However, it would not have been as efficient at bipedality as humans, nor at arboreality as non-human great apes. Its discovery, along with Miocene apes, has reworked academic understanding of the chimpanzee-human last common ancestor from appearing much like modern day chimpanzees, orangutans and gorillas to being a creature without a modern anatomical cognate.
The Hominidae, whose members are known as great apes or hominids, are a taxonomic family of primates that includes eight extant species in four genera: Pongo ; Gorilla ; Pan ; and Homo, of which only modern humans remain.
Paracolobus is an extinct genus of primate closely related to the living colobus monkeys. It lived in eastern Africa in the Pliocene and Early Pleistocene. Fossils have been found in Kenya and Ethiopia, in places such as the Omo valley.
Equatorius is an extinct genus of kenyapithecine primate found in central Kenya at the Tugen Hills. Thirty-eight large teeth belonging to this middle Miocene hominid in addition to a mandibular and partially complete skeleton dated 15.58 Ma and 15.36 Ma. were later found.
Simiolus is an extinct genus of dendropithecid primates. It was described by Mary Leakey and Richard Leakey in 1987, and the type species is S. enjiessi, which existed during the Miocene of Kenya. The species epithet is a phonetic pun on the acronym NGS. A new species, S. andrewsi, also from the middle Miocene of Kenya, was described by Terry Harrison in 2010. In November 2018, scientists reported the discovery of the smallest known ape, Simiolus minutus, which weighed approximately eight pounds, and lived about 12.5 million years ago in Kenya in East Africa.
Cercopithecoides is an extinct genus of colobine monkey from Africa which lived during the latest Miocene to the Pleistocene period. There are several recognized species, with the smallest close in size to some of the larger extant colobines, and males of the largest species weighed over 50 kilograms (110 lb).
Epipliopithecus vindobonensis is an extinct species of pliopithecoid primate recovered from the Middle Miocene deposits of Devínska Nová Ves fissure in western Slovakia. Epipliopithecus is one of the few pliopithecoids for which both cranial and post-cranial fossil material has been recovered. Most pliopithecoids are known only from fossilized teeth, whereas Epipliopithecus is known from three nearly complete skeletons. As such, Epipliopithecus has greatly informed the modern understanding of pliopithecoid anatomy, locomotion, and phylogeny.
Enhydriodon dikikae is an extinct species in the family Mustelidae that existed during the Miocene and Pliocene epoch. Fossils from this species were discovered in Dikika, Lower Awash Valley in Ethiopia. These fossils were found in the lower basal member of the Hadar Formation, which means they existed more than 3.42 million years ago but the estimated age of the fossils is greater than 3.4 million years old and likely to be closer to 4 million years old. In comparison to the other species in the genus Enhydriodon, E. dikikae is one of the youngest.
Pliopapio is an extinct genus of Old World monkey known from the latest part of the Miocene to the early Pliocene Epochs from the Afar Region of Ethiopia. It was first described based on a very large series of fossils from the site of Aramis in the Middle Awash, which has been dated by 40Ar/39Ar to 4.4 million years old. It has since been found from similarly aged sediments at Gona, approximately 75 km to the North. Additional fossils from the Middle Awash extend its known time range back to at least 5.3 million years ago. There is only one known species, Pliopapio alemui.
The Nyanzapithecinae or Nyanzapithecines are a subfamily of extinct Dendropithecidae as sister of Simiolus. The group contains Rangwapithecus, Turkanapithecus, Rukwapithecus, Oreopithecus, and Nyanzapithecus. In the following tree the internal structure of Nyanzapithecinae of Nengo et al. is followed.
Micropithecus is an extinct genus of primates that lived in East Africa about 19 to 15 million years ago, during the early Miocene. The genus and its type species, Micropithecus clarki, were first scientifically described in 1978.
Rhinocolobus is an extinct genus of monkey closely related to modern colobus monkeys. It lived in eastern Africa during the Plio-Pleistocene, existing as recently as 1.5 million years ago.
Dolichopithecus is an extinct genus of Old World monkey that lived in Europe during the Late Miocene and Pliocene.
