In evolutionary biology, megatrajectories are the major evolutionary milestones and directions in the evolution of life.
Posited by A. H. Knoll and Richard K. Bambach in their 2000 collaboration, "Directionality in the History of Life," Knoll and Bamback argue that, in consideration of the problem of progress in evolutionary history, a middle road that encompasses both contingent and convergent features of biological evolution may be attainable through the idea of the megatrajectory:
We believe that six broad megatrajectories capture the essence of vectoral change in the history of life. The megatrajectories for a logical sequence dictated by the necessity for complexity level N to exist before N+1 can evolve...In the view offered here, each megatrajectory adds new and qualitatively distinct dimensions to the way life utilizes ecospace. [1]
According to Knoll and Bambach, the six megatrajectories outlined by biological evolution thus far are:
Milan M. Ćirković and Robert Bradbury, [2] [3] have taken the megatrajectory concept one step further by theorizing that a seventh megatrajectory exists: postbiological evolution triggered by the emergence of artificial intelligence at least equivalent to the biologically-evolved one, as well as the invention of several key technologies of the similar level of complexity and environmental impact, such as molecular nanoassembling or stellar uplifting.
An extinction event is a widespread and rapid decrease in the biodiversity on Earth. Such an event is identified by a sharp fall in the diversity and abundance of multicellular organisms. It occurs when the rate of extinction increases with respect to the background extinction rate and the rate of speciation. Estimates of the number of major mass extinctions in the last 540 million years range from as few as five to more than twenty. These differences stem from disagreement as to what constitutes a "major" extinction event, and the data chosen to measure past diversity.
The timeline of the evolutionary history of life represents the current scientific theory outlining the major events during the development of life on planet Earth. Dates in this article are consensus estimates based on scientific evidence, mainly fossils.
Macroevolution comprises the evolutionary processes and patterns which occur at and above the species level. In contrast, microevolution is evolution occurring within the population(s) of a single species. In other words, microevolution is the scale of evolution that is limited to intraspecific (within-species) variation, while macroevolution extends to interspecific (between-species) variation. The evolution of new species (speciation) is an example of macroevolution. This is the common definition for 'macroevolution' used by contemporary scientists. Although, the exact usage of the term has varied throughout history.
Paleontology, also spelled palaeontology or palæontology, is the scientific study of life that existed prior to the start of the Holocene epoch. It includes the study of fossils to classify organisms and study their interactions with each other and their environments. Paleontological observations have been documented as far back as the 5th century BC. The science became established in the 18th century as a result of Georges Cuvier's work on comparative anatomy, and developed rapidly in the 19th century. The term has been used since 1822 formed from Greek παλαιός, ὄν, and λόγος.
Cope's rule, named after American paleontologist Edward Drinker Cope, postulates that population lineages tend to increase in body size over evolutionary time. It was never actually stated by Cope, although he favoured the occurrence of linear evolutionary trends. It is sometimes also known as the Cope–Depéret rule, because Charles Depéret explicitly advocated the idea. Theodor Eimer had also done so earlier. The term "Cope's rule" was apparently coined by Bernhard Rensch, based on the fact that Depéret had "lionized Cope" in his book. While the rule has been demonstrated in many instances, it does not hold true at all taxonomic levels, or in all clades. Larger body size is associated with increased fitness for a number of reasons, although there are also some disadvantages both on an individual and on a clade level: clades comprising larger individuals are more prone to extinction, which may act to limit the maximum size of organisms.
A grade is a taxon united by a level of morphological or physiological complexity. The term was coined by British biologist Julian Huxley, to contrast with clade, a strictly phylogenetic unit.
The Serpukhovian is in the ICS geologic timescale the uppermost stage or youngest age of the Mississippian, the lower subsystem of the Carboniferous. The Serpukhovian age lasted from 330.9 Ma to 323.2 Ma. It is preceded by the Visean and is followed by the Bashkirian. The Serpukhovian correlates with the lower part of the Namurian Stage of European stratigraphy and the middle and upper parts of the Chesterian Stage of North American stratigraphy.
Andrew Herbert Knoll is the Fisher Research Professor of Natural History and a Research Professor of Earth and Planetary Sciences at Harvard University. Born in West Reading, Pennsylvania, in 1951, Andrew Knoll graduated from Lehigh University with a Bachelor of Arts in 1973 and received his Ph.D. from Harvard University in 1977 for a dissertation titled "Studies in Archean and Early Proterozoic Paleontology." Knoll taught at Oberlin College for five years before returning to Harvard as a professor in 1982. At Harvard, he serves in the departments of Organismic and Evolutionary Biology and Earth and Planetary Sciences.
Namapoikia rietoogensis is among the earliest known animals to produce a calcareous skeleton. Known from the Ediacaran period, before the Cambrian explosion of calcifying animals, the long-lived organism grew up to a metre in diameter and resembles a colonial sponge. It was an encruster, filling vertical fissures in the reefs in which it originally grew.
The Ediacaranbiota is a taxonomic period classification that consists of all life forms that were present on Earth during the Ediacaran Period. These were enigmatic tubular and frond-shaped, mostly sessile, organisms. Trace fossils of these organisms have been found worldwide, and represent the earliest known complex multicellular organisms. The term "Ediacara biota" has received criticism from some scientists due to its alleged inconsistency, arbitrary exclusion of certain fossils, and inability to be precisely defined.
The evolution of biological complexity is one important outcome of the process of evolution. Evolution has produced some remarkably complex organisms – although the actual level of complexity is very hard to define or measure accurately in biology, with properties such as gene content, the number of cell types or morphology all proposed as possible metrics.
The history of life on Earth traces the processes by which living and extinct organisms evolved, from the earliest emergence of life to the present day. Earth formed about 4.5 billion years ago and evidence suggests that life emerged prior to 3.7 Ga. The similarities among all known present-day species indicate that they have diverged through the process of evolution from a common ancestor.
The siphonal canal is an anatomical feature of the shells of certain groups of sea snails within the clade Neogastropoda. Some sea marine gastropods have a soft tubular anterior extension of the mantle called a siphon through which water is drawn into the mantle cavity and over the gill and which serves as a chemoreceptor to locate food. Siphonal canals allow for active transport of water to sensory organs inside the shell. Organisms without siphonal canals in their shells rely on passive or diffuse transport or water into their shell. Those with siphonal canals have a direct inhalant stream of water that interacts with sensory organs to detect concentration and direction of a stimulus, such as food or mates. In certain groups of carnivorous snails, where the siphon is particularly long, the structure of the shell has been modified in order to house and protect the soft structure of the siphon. Thus the siphonal canal is a semi-tubular extension of the aperture of the shell through which the siphon is extended when the animal is active.
Polysporangiophytes, also called polysporangiates or formally Polysporangiophyta, are plants in which the spore-bearing generation (sporophyte) has branching stems (axes) that bear sporangia. The name literally means 'many sporangia plant'. The clade includes all land plants (embryophytes) except for the bryophytes whose sporophytes are normally unbranched, even if a few exceptional cases occur. While the definition is independent of the presence of vascular tissue, all living polysporangiophytes also have vascular tissue, i.e., are vascular plants or tracheophytes. Extinct polysporangiophytes are known that have no vascular tissue and so are not tracheophytes.

Funisia is a genus of animal containing the single species F. dorothea. It is an extinct animal from the Ediacaran biota, discovered in South Australia in 2008 by Mary L. Droser and James G. Gehling.
The Mesozoic marine revolution (MMR) refers to the increase in shell-crushing (durophagous) and boring predation in benthic organisms throughout the Mesozoic era, along with bulldozing and sediment remodelling in marine habitats. The term was first coined by Geerat J. Vermeij, who based his work on that of Steven M. Stanley. While the MMR was initially restricted to the Cretaceous, more recent studies have suggested that the beginning of this ecological/evolutionary arms race extends as far back as the Triassic, with the MMR now being considered to have started in the Anisian or the Aalenian. It is an important transition between the Palaeozoic evolutionary fauna and the Modern evolutionary fauna that occurred throughout the Mesozoic.
The Khatyspyt Formation is a Neoproterozoic formation exposed in the Olenëk Uplift of north central Siberia, which contains the only known instance of the Ediacara biota preserved in a limestone bed. The Khatyspyt Formation forms one of the major parts of the Khorbusuonka Group; underlying the Khatyspyt are dolomites of the Mastakh Formation and their overlying red beds; the Turkut Formation overlies the Khatyspyt. The Khatyspyt and part of the overlying Turkut comprise a major shallowing upward marine carbonate sequence. Khatyspytia is named after this formation.
The Cambrian explosion is an interval of time approximately 538.8 million years ago in the Cambrian period of the early Paleozoic when a sudden radiation of complex life occurred, and practically all major animal phyla started appearing in the fossil record. It lasted for about 13 to 25 million years and resulted in the divergence of most modern metazoan phyla. The event was accompanied by major diversification in other groups of organisms as well.
A ghost lineage is a hypothesized ancestor in a species lineage that has left no fossil evidence, but can still be inferred to exist or have existed because of gaps in the fossil record or genomic evidence. The process of determining a ghost lineage relies on fossilized evidence before and after the hypothetical existence of the lineage and extrapolating relationships between organisms based on phylogenetic analysis. Ghost lineages assume unseen diversity in the fossil record and serve as predictions for what the fossil record could eventually yield; these hypotheses can be tested by unearthing new fossils or running phylogenetic analyses.
The Mesozoic–Cenozoic Radiation is the third major extended increase of biodiversity in the Phanerozoic, after the Cambrian Explosion and the Great Ordovician Biodiversification Event, which appeared to exceeded the equilibrium reached after the Ordovician radiation. Made known by its identification in marine invertebrates, this evolutionary radiation began in the Mesozoic, after the Permian extinctions, and continues to this date. This spectacular radiation affected both terrestrial and marine flora and fauna, during which the "modern" fauna came to replace much of the Paleozoic fauna. Notably, this radiation event was marked by the rise of angiosperms during the mid-Cretaceous, and the K-Pg extinction, which initiated the rapid increase in mammalian biodiversity.