Part of a series on |
The Burgess Shale |
---|
The Mount Stephen trilobite beds (UNSM locality 14s) [1] are a series of fossil strata on Mount Stephen, British Columbia that contain exceptionally preserved fossil material. Part of the same stratigraphic unit as the Burgess Shale, many non-mineralized parts (such as anomalocarid claws, sponges, and trilobite legs) are preserved; in addition, a high density of trilobite fossils is present.
The trilobite beds were the first Burgess shale locality to be discovered. [2]
The richness of fossils in the Field area was first identified by workers associated with the construction of the Trans-Canada railway, which had (somewhat controversially) been routed through the Kicking Horse valley. [3] Richard McConnell, of the Geological Survey of Canada, was pointed to the beds by a railway worker whilst mapping the geology around the railway line in September 1886. [4] Several unusual fossils were subsequently described from this site, including sponges, worms, [5] and the appendages of the unusual Anomalocaris, identified at that time as the bodies of crabs. [6] These fossils prompted Charles Doolittle Walcott to make forays into the area, and led to his discovery of the Walcott Quarry on Fossil Ridge.
The trilobite beds are in the Campsite Cliff member, and contain the marker trilobite Ogygopsis . [7]
The Burgess Shale is a fossil-bearing deposit exposed in the Canadian Rockies of British Columbia, Canada. It is famous for the exceptional preservation of the soft parts of its fossils. At 508 million years old, it is one of the earliest fossil beds containing soft-part imprints.
Marrella is an extinct genus of arthropod known from the middle Cambrian Burgess Shale of British Columbia. It is the most common animal represented in the Burgess Shale.
Aysheaia was a genus of Cambrian-aged soft-bodied, caterpillar-shaped fossil organisms with average body lengths of 1–6 cm.
Charles Doolittle Walcott was an American paleontologist, administrator of the Smithsonian Institution from 1907 to 1927, and geologist. He is famous for his discovery in 1909 of well-preserved fossils, including some of the oldest soft-part imprints, in the Burgess Shale of British Columbia, Canada.
Field is an unincorporated community of approximately 169 people located in the Kicking Horse River valley of southeastern British Columbia, Canada, within the confines of Yoho National Park. At an elevation of 1,256 m (4,121 ft), it is 27 km (17 mi) west of Lake Louise along the Trans-Canada Highway, which provides the only road access to the town. The community is named for Cyrus West Field of Transatlantic telegraph cable fame, who visited the area in 1884.
Ottoia is a stem-group archaeopriapulid worm known from Cambrian fossils. Although priapulid-like worms from various Cambrian deposits are often referred to Ottoia on spurious grounds, the only clear Ottoia macrofossils come from the Burgess Shale of British Columbia, which was deposited 508 million years ago. Microfossils extend the record of Ottoia throughout the Western Canada Sedimentary Basin, from the mid- to late- Cambrian. A few fossil finds are also known from China.
Anomalocaris is an extinct genus of radiodont (anomalocaridid), an order of animals thought to be closely related to ancestral arthropods. The first fossils of Anomalocaris were discovered in the Ogygopsis Shale by Joseph Frederick Whiteaves, with more examples found by Charles Doolittle Walcott in the Burgess Shale. Originally several fossilized parts discovered separately were thought to be three separate creatures, a misapprehension corrected by Harry B. Whittington and Derek Briggs in a 1985 journal article. Anomalocaris is thought to be one of the earliest examples of an apex predator, though others have been found in older Cambrian lagerstätten deposits.
Odontogriphus is a genus of soft-bodied animals known from middle Cambrian Lagerstätte. Reaching as much as 12.5 centimetres (4.9 in) in length, Odontogriphus is a flat, oval bilaterian which apparently had a single muscular foot, and a "shell" on its back that was moderately rigid but of a material unsuited to fossilization.
Choia is a genus of extinct demosponge ranging from the Cambrian until the Lower Ordovician periods. Fossils of Choia have been found in the Burgess Shale in British Columbia; the Maotianshan shales of China; the Wheeler Shale in Utah; and the Lower Ordovician Fezouata formation. It was first described in 1920 by Charles Doolittle Walcott.
A number of assemblages bear fossil assemblages similar in character to that of the Burgess Shale. While many are also preserved in a similar fashion to the Burgess Shale, the term "Burgess Shale type fauna" covers assemblages based on taxonomic criteria only.
The fossils of the Burgess Shale, like the Burgess Shale itself, formed around 505 million years ago in the Mid Cambrian period. They were discovered in Canada in 1886, and Charles Doolittle Walcott collected over 65,000 specimens in a series of field trips up from 1909 to 1924. After a period of neglect from the 1930s to the early 1960s, new excavations and re-examinations of Walcott's collection continue to discover new species, and statistical analysis suggests discoveries will continue for the foreseeable future. Stephen Jay Gould's book Wonderful Life describes the history of discovery up to the early 1980s, although his analysis of the implications for evolution has been contested.
The Stephen Formation is a geologic formation exposed in the Canadian Rockies of British Columbia and Alberta, on the western edge of the Western Canada Sedimentary Basin. It consists of shale, thin-bedded limestone, and siltstone that was deposited during Middle Cambrian time. It is famous for the exceptional preservation of soft-bodied fossils: the Burgess Shale biota. The formation overlies the Cathedral escarpment, a submarine cliff; consequently it is divided into two quite separate parts, the 'thin' sequence deposited in the shallower waters atop the escarpment, and the 'thick' sequence deposited in the deeper waters beyond the cliff. Because the 'thick' Stephen Formation represents a distinct lithofacies, some authors suggest it warrants its own name, and dub it the Burgess Shale Formation. The stratigraphy of the Thin Stephen Formation has not been subject to extensive study, so except where explicitly mentioned this article applies mainly to the Thick Stephen Formation.
The Burgess Shale, a series of fossil beds in the Canadian Rockies, was first noticed in 1886 by Richard McConnell of the Geological Survey of Canada (GSC). His and subsequent finds, all from the Mount Stephen area, came to the attention of palaeontologist Charles Doolittle Walcott, who in 1907 found time to reconnoitre the area. He opened a quarry in 1910 and in a series of field trips brought back 65,000 specimens, which he identified as Middle Cambrian in age. Due to the quantity of fossils and the pressures of his other duties at the Smithsonian Institution, Walcott was only able to publish a series of "preliminary" papers, in which he classified the fossils within taxa that were already established. In a series of visits beginning in 1924, Harvard University professor Percy Raymond collected further fossils from Walcott's quarry and higher up on Fossil Ridge, where slightly different fossils were preserved.
The Phyllopod bed, designated by USNM locality number 35k, is the most famous fossil-bearing member of the Burgess shale fossil Lagerstätte. It was quarried by Charles Walcott from 1911–1917, and was the source of 95% of the fossils he collected during this time; tens of thousands of soft-bodied fossils representing over 150 genera have been recovered from the Phyllopod bed alone.
Cambrorhytium is an enigmatic fossil genus known from the Latham Shale (California), and the Chengjiang (China) and Burgess Shale lagerstätte. 350 specimens of Cambrorhytium are known from the Greater Phyllopod bed, where they comprise 0.7% of the community.
Hamptonia is an extinct genus of sea sponge known from the Middle Cambrian Burgess Shale and the Lower Ordovician Fezouata formation. It was first described in 1920 by Charles Doolittle Walcott. 48 specimens of Hamptonia are known from the Greater Phyllopod bed, where they comprise < 0.1% of the community.
Pagetia Walcott, 1916. is a small genus of trilobite, assigned to the Eodiscinid family Pagetiidae and which had global distribution during the Middle Cambrian. The genus contains 55 currently recognized species, each with limited spatial and temporal ranges.
Pirania is an extinct genus of sea sponge known from the Middle Cambrian Burgess Shale and the Ordovician Fezouata formation. It is named after Mount St. Piran, a mountain situated in the Bow River Valley in Banff National Park, Alberta. It was first described in 1920 by Charles Doolittle Walcott. 198 specimens of Pirania are known from the Greater Phyllopod bed, where they comprise 0.38% of the community.
Wapkia is an extinct genus of sea sponge with radial sclerites, known from the Middle Cambrian Burgess Shale. It was first described in 1920 by Charles Doolittle Walcott. 32 specimens of Wapkia are known from the Greater Phyllopod bed, where they comprise 0.06% of the community.
The Cathedral Formation is a stratigraphic unit in the southern Canadian Rockies of Alberta and British Columbia, on the western edge of the Western Canada Sedimentary Basin. It is a thick sequence of carbonate rocks of Middle Cambrian age. It was named for Cathedral Mountain in Yoho National Park by Charles Doolittle Walcott, the discoverer of the Burgess shale fossils.