The Ditrysia are a natural group or clade of insects in the lepidopteran order containing both butterflies and moths. They are so named because the female has two distinct sexual openings: one for mating, and the other for laying eggs.
Macrolepidoptera is a group within the insect order Lepidoptera. Traditionally used for the larger butterflies and moths as opposed to the "microlepidoptera", this group is artificial. However, it seems that by moving some taxa about, a monophyletic macrolepidoptera can be easily achieved. The two superfamilies Geometroidea and Noctuoidea account for roughly one-quarter of all known Lepidoptera.
Riodinidae is the family of metalmark butterflies. The common name "metalmarks" refers to the small, metallic-looking spots commonly found on their wings. The 1532 species are placed in 146 genera. Although mostly Neotropical in distribution, the family is also represented both in the Nearctic, Palearctic, Australasian (Dicallaneura), Afrotropic, and Indomalayan realms.
Schreckensteinioidea is a superfamily in the insect order Lepidoptera containing a single family, Schreckensteiniidae, or "bristle-legged moths", because of the stout spines on the hindlegs. The superfamily and family were both described by Thomas Bainbrigge Fletcher in 1929. The relationships of this family within the group apoditrysia are currently uncertain. One of the species, the blackberry skeletoniser, is widespread and common across Europe and has been introduced as a biological control to Hawaii, whilst three species of Corsocasis occur in South East Asia.
Immoidea is a superfamily of pantropical moths containing only the family Immidae comprising ten genera with around 250 species, over half of them in the genus Imma. Many are brightly coloured and diurnal. The position of this group is currently uncertain within the group Obtectomera. The larvae feed on the leaves of dicotyledons and conifers including Podocarpus.
Coelolepida is a clade of insects in the lepidopteran order, containing the infraorders Acanthoctesia and Lophocoronina.
Callidulidae, the only known family of the superfamily Calliduloidea, is the family of Old World butterfly-moths, containing eight genera. They have a peculiar distribution, restricted to the Old World tropics of Southeast Asia to Australasia and Madagascar. The three subfamilies exhibit both day- and night-flying behaviour.
Carposinidae, the "fruitworm moths", is a family of insects in the order Lepidoptera. These moths are narrower winged than Copromorphidae, with less rounded forewing tips. Males often have conspicuous patches of scales on either surface. The mouthparts are quite diagnostic, usually with prominent, upcurved "labial palps", the third segment long, and the second segment covered in large scales. Unlike Copromorphidae, the "M2" and sometimes "M1" vein on the hindwings is absent. The relationship of Carposinidae relative to Copromorphidae needs further investigation. It is considered possible that the family is artificial, being nested within Copromorphidae. The Palearctic species have been revised by Alexey Diakonoff (1989).
Galacticidae is a recently recognised and enigmatic family of insects in the lepidopteran order. These moderate sized moths are 8–17 mm in wingspan and have previously been embedded within several lepidopteran superfamilies, but Galacticidae is currently placed in its own superfamily at the base of the natural group Apoditrysia.
Copromorphoidea, the "fruitworm moths", is a superfamily of insects in the lepidopteran order. These moths are small to medium-sized and are broad-winged bearing some resemblance to the superfamilies Tortricoidea and Immoidea. The antennae are often "pectinate" especially in males, and many species of these well camouflaged moths bear raised tufts of scales on the wings and a specialised fringe of scales at the base of the hindwing sometimes in females only; there are a number of other structural characteristics. The position of this superfamily is not certain, but it has been placed in the natural group of "Apoditrysia" "Obtectomera", rather than with the superfamilies Alucitoidea or Epermenioidea within which it has sometimes previously been placed, on the grounds that shared larval and pupal characteristics of these groups have probably evolved independently. It has been suggested that the division into two families should be abandoned.
A family of primitive monotrysian moths in the order Lepidoptera, Heliozelidae are small, metallic day-flying moths with shiny smooth heads. In Europe the small adult moths are seldom noticed as they fly quite early in the spring. The larvae are leaf miners and the vacated leaf mines are distinctive because the larva leaves a large hole at the end.
Mnesarchaeoidea is a superfamily of "New Zealand primitive moths" containing one family, Mnesarchaeidae and a two genera, Mnesarchaea, and Mnesarchella, both of which are endemic to New Zealand.
The Palaeosetidae or miniature ghost moths are a family of insects in the order Lepidoptera contained within the superfamily Hepialoidea.
The Exoporia are a group of primitive Lepidoptera comprising the superfamilies Mnesarchaeoidea and Hepialoidea. They are a natural group or clade. Exoporia is the sister group of the lepidopteran infraorder Heteroneura. They are characterised by their unique female reproductive system which has an external groove between the ostium bursae and the ovipore by which the sperm is transferred to the egg rather than having the mating and egg-laying parts of the abdomen with a common opening (cloaca) as in other nonditrysian moths, or with separate openings linked internally by a "ductus seminalis" as in the Ditrysia. See Kristensen for other exoporian characteristics.
Acanthopteroctetidae is a small family of primitive moths with two described genera, Acanthopteroctetes and Catapterix, and a total of seven described species. They are known as the archaic sun moths.
The Monotrysia are a group of moths in the lepidopteran order, not currently considered to be a natural group or clade. Apart from the recently discovered family Andesianidae, most of the group consists of small, relatively understudied species. The group is so named because the female has a single genital opening for mating and laying eggs, in contrast to the rest of the Lepidoptera (Ditrysia), which have two female reproductive openings. They comprise all of the group Heteroneura apart from the Ditrysia.
Heteroneura is a natural group in the insect order Lepidoptera that comprises over 99% of all butterflies and moths. This is the sister group of the infraorder Exoporia, and is characterised by wing venation which is not similar or homoneurous in both pairs of wings. Though basal groups within the Heteroneura cannot be identified with much confidence, one major subgroup is the leaf-mining Nepticuloidea. Species in this subgroup include some of the smallest lepidoterans identified.
Neolepidoptera is a clade within Myoglossata in suborder Glossata of order Lepidoptera, the butterflies and moths. They differ from other Myoglossata in the larval stage abdominal prolegs, pupal morphology, and the mandibles are reduced in area. They also differ in their reproductive systems. The prolegs have muscles and apical hooklets. The reproductive organs have two openings. There are also differences in the wing structure. The pupae are "incomplete or obtect."
Micropterigoidea is the superfamily of "mandibulate archaic moths", all placed in the single family Micropterigidae, containing currently about twenty living genera. They are considered the most primitive extant lineage of lepidoptera. The name comes from the Greek for mikros, little and pterux, a wing. The fossil record of the group goes back to the middle-late Jurassic with the earliest known species being Auliepterix from the Karabastau Formation in Kazakhstan.
The Macroheterocera are a well supported clade of moths that are closely related to butterflies and other macro-moths.
