Scale insects are small insects of the order Hemiptera, suborder Sternorrhyncha. Of dramatically variable appearance and extreme sexual dimorphism, they comprise the infraorder Coccomorpha which is considered a more convenient grouping than the superfamily Coccoidea due to taxonomic uncertainties. Adult females typically have soft bodies and no limbs, and are concealed underneath domed scales, extruding quantities of wax for protection. Some species are hermaphroditic, with a combined ovotestis instead of separate ovaries and testes. Males, in the species where they occur, have legs and sometimes wings, and resemble small flies. Scale insects are herbivores, piercing plant tissues with their mouthparts and remaining in one place, feeding on sap. The excess fluid they imbibe is secreted as honeydew on which sooty mold tends to grow. The insects often have a mutualistic relationship with ants, which feed on the honeydew and protect them from predators. There are about 8,000 described species.
The Sternorrhyncha suborder of the Hemiptera contains the aphids, whiteflies, and scale insects, groups which were traditionally included in the now-obsolete order "Homoptera". "Sternorrhyncha" refers to the rearward position of the mouthparts relative to the head.
The Chamaemyiidae are a small family of acalyptrate flies with less than 200 species described worldwide. The larvae of these small flies are active and predatory and are often used for biological control of aphids, scale insects, and similar pests. Chamaemyiid fossils are poorly represented in amber deposits, but a few examples are known from the Eocene epoch onwards.
The Margarodidae or ground pearls are a family of scale insects within the superfamily Coccoidea. Members of the family include the Polish cochineal and Armenian cochineal and the original ground pearl genus, Margarodes. Beginning in 1880, a number of distinct subfamilies were recognized, with the giant coccids being the first. Although Maskell proposed a new family, many continued to regard the monophlebids as a mere subfamily for many years, and the Margarodidae classification continued to be polyphyletic through the 20th Century. Since then, taking the advice of Koteja several subfamilies and tribes have been elevated into their own families such as Matsucoccidae and Xylococcidae. The pared-down family of Margarodidae is monophyletic.
Paratachardina pseudolobata, the lobate lac scale, is a polyphagous and pestiferous lac scale insect, which damages trees and woody shrubs in Cuba, Florida, the Bahamas and the Australian territory of Christmas Island. It was mistakenly identified as Paratachardina lobata (Chamberlin), an insect native to India and Sri Lanka, but was in 2007 recognized and named as a distinct species based on material from Florida; its native distribution is as yet unknown. The new lac insect was described based on all stages of the female, during the revision of the genus Paratachardina, wherein all its known species were redescribed.
The Coccidae are a family of scale insects belonging to the superfamily Coccoidea. They are commonly known as soft scales, wax scales or tortoise scales. The females are flat with elongated oval bodies and a smooth integument which may be covered with wax. In some genera they possess legs but in others, they do not, and the antennae may be shortened or missing. The males may be winged or wingless.
Conchaspis capensis is a species of scale insect from South Africa found on Metalasia muricata and Phylica species. It was originally described by Carl Linnaeus in his 1763 work Centuria Insectorum.
Coccus is a genus of scale insects in the family Coccidae. Several species, such as Coccus viridis, a major pest of coffee, are major agricultural pests. The type species is Coccus hesperidumLinnaeus.
Beesoniidae is a family of scale insects commonly known as beesoniids. They typically cause galls on their plant hosts. Members of this family mostly come from southern Asia. The family name comes from the type genus Beesonia which is named after the entomologist C.F.C. Beeson who obtained the specimens from which they were described and named.
Monophlebidae is a family of scale insects commonly known as the giant scales or monophlebids. They occur in most parts of the world but more genera are found in the tropics than elsewhere.
Putoidae is a family of scale insects commonly known as giant mealybugs or putoids. There is probably a single genus, Puto, containing about sixty species. The genus name Macrocerococcus has also been used but it is now considered to be a synonym of Puto. The genus Puto was formerly classified as a member of the Pseudococcidae; however, it so significantly differed from the rest of the Pseudococcidae that it was accorded its own family Putoidae.
Stictococcidae is a family of scale insects commonly known as stictococcids. There are three genera containing about seventeen species and members of this family are found only in Afrotropical regions.
The Galleriinae are a subfamily of snout moths and occur essentially worldwide, in some cases aided by involuntary introduction by humans. This subfamily includes the wax moths, whose caterpillars (waxworms) are bred on a commercial scale as food for pets and as fishing bait; in the wild, these and other species of Galleriinae may also be harmful to humans as pests.
Pulvinaria regalis is a species of scale insect in the family Coccidae. Although it is commonly known as the horse chestnut scale, it affects other trees besides horse chestnuts as well as many species of woody shrubs. Adults are normally all female and produce eggs by parthenogenesis. The insects are thought to have originated in Asia but arrived in Europe in the second half of the twentieth century.
Ceroplastes cirripediformis, known generally as the barnacle scale or barnacle wax scale, is a species of soft scale insect in the family Coccidae.
Ceroplastes is a genus of wax scales in the family Coccidae. There are more than 130 described species in Ceroplastes.
Ceroplastes rubens, known generally as the red wax scale or pink wax scale, is a species of soft scale insect in the family Coccidae.
Saissetia oleae is a scale insect in the family Coccidae. It is considered one of the three main phytophagous parasites of the olive tree, together with the olive fruit fly and the olive moth. Although it is a common parasite which occurs most often in olive trees, it is a polyphagous species, also attacking citrus trees as well as various ornamental shrubs such as oleanders, pittosporums and euonymus.
Acrometopia is a genus of flies in the family Chamaemyiidae.
Ceroplastes ceriferus, the Indian wax scale, is a species of scale insect in the family Coccidae. It is native to southern Asia and has spread to many other parts of the world.
