The pes (Latin for foot) is the zoological term for the distal portion of the hind limb of tetrapod animals. [1] It is the part of the pentadactyl limb that includes the metatarsals and digits (phalanges). During evolution, it has taken many forms and served a variety of functions. It can be represented by the foot of primates, the lower hind limb of hoofed animals, [2] the lower portion of the leg of dinosaurs including birds or the rear paw. It is also represented in the rear 'paddle' of extinct marine reptiles, such as plesiosaurs. The oldest types of tetrapods had seven or eight digits. [1]
The foot is an anatomical structure found in many vertebrates. It is the terminal portion of a limb which bears weight and allows locomotion. In many animals with feet, the foot is a separate organ at the terminal part of the leg made up of one or more segments or bones, generally including claws or nails.
Tetrapods are four-limbed animals constituting the superclass Tetrapoda. It includes extant and extinct amphibians, reptiles, and synapsids. Tetrapods evolved from a group of animals known as the Tetrapodomorpha which, in turn, evolved from ancient sarcopterygians around 390 million years ago in the middle Devonian period; their forms were transitional between lobe-finned fishes and the four-limbed tetrapods. The first tetrapods appeared by the late Devonian, 367.5 million years ago. The specific aquatic ancestors of the tetrapods and the process by which they colonized Earth's land after emerging from water remains unclear. The change from a body plan for breathing and navigating in water to a body plan enabling the animal to move on land is one of the most profound evolutionary changes known. The first tetrapods were primarily aquatic. Modern amphibians, which evolved from earlier groups, are generally semiaquatic; the first stage of their lives is as fish-like tadpoles, and later stages are partly terrestrial and partly aquatic. However, most tetrapod species today are amniotes, most of those are terrestrial tetrapods whose branch evolved from earlier tetrapods about 340 million years ago. The key innovation in amniotes over amphibians is laying of eggs on land or having further evolved to retain the fertilized egg(s) within the mother.
In biology, dactyly is the arrangement of digits on the hands, feet, or sometimes wings of a tetrapod animal. It comes from the Greek word δακτυλος (dáktylos) = "finger".
Ichthyostega is an early genus of tetrapodomorphs that lived at the end of the Late Devonian Period. It was one of the first four-limbed vertebrates in the fossil record. These tetrapods lived in the shallow part of the oceans of the Northern Hemisphere. Ichthyostega possessed lungs and limbs that helped it navigate through shallow water in swamps. Although Ichthyostega is often labelled a "tetrapod" due to the possession of limbs and fingers, it was more basal ("primitive") than true crown-tetrapods, and could more accurately be referred to as a stegocephalian or stem tetrapod. Likewise, while undoubtedly of amphibian build and habit, it is not considered a true member of the group in the narrow sense, as the first modern amphibians appeared in the Triassic Period. Until finds of other early stegocephalians and closely related fishes in the late 20th century, Ichthyostega stood alone as a transitional fossil between fish and tetrapods, combining fish- and tetrapod-like features. Newer research has shown that it had an unusual anatomy, functioning more akin to a seal than a salamander, as previously assumed. They spent about 70-90% of their time underwater because they were so heavy it was a bit hard for them to keep their body on land.
Ophidia is a group of squamate reptiles including modern snakes and all reptiles more closely related to snakes than to other living groups of lizards.
Pes or the acronym PES may refer to:
A forelimb is an anterior limb on a terrestrial vertebrate's body. With reference to quadrupeds, the term foreleg is often used instead.
A polydactyl cat is a cat with a congenital physical anomaly called polydactyly, which causes the cat to be born with more than the usual number of toes on one or more of its paws. Cats with this genetically inherited trait are most commonly found along the East Coast of North America and in South West England and Wales.
Hynerpeton is an extinct genus of early four-limbed vertebrate that lived in the rivers and ponds of Pennsylvania during the Late Devonian period, around 365 to 363 million years ago. The only known species of Hynerpeton is H. bassetti, named after the describer's grandfather, city planner Edward Bassett. Hynerpeton is known for being the first Devonian four-limbed vertebrate discovered in the United States, as well as possibly being one of the first to have lost internal (fish-like) gills.
Tiktaalik is a monospecific genus of extinct sarcopterygian from the Late Devonian Period, about 375 Mya, having many features akin to those of tetrapods.
Orthosuchus is an extinct genus of protosuchian crocodyliform that lived during the Early Jurassic, about 196 million years ago. It was first discovered in 1963 in the Red Beds Formation in the Qacha's Nek Province of Lesotho, southern Africa. The characteristics showed on its postcranial skeleton and the skull indicated that it is a Crocodyliform. The finding is significant since some of the characteristics found on this specimen were believed to be absent until Jurassic.
The manus is the zoological term for the distal portion of the fore limb of an animal. In tetrapods, it is the part of the pentadactyl limb that includes the metacarpals and digits (phalanges). During evolution, it has taken many forms and served a variety of functions. It can be represented by the hand of primates, the lower front limb of hoofed animals or the fore paw and is represented in the wing of birds, bats and prehistoric flying reptiles (pterosaurs), the flipper of marine mammals and the 'paddle' of extinct marine reptiles, such as plesiosaurs and ichthyosaurs.
A limb, or extremity, is a jointed, or prehensile, appendage of the human or other animal body. In the human body, the arms and the legs are commonly called the upper limbs and lower limbs respectively, to include part of the shoulder and hip girdles. Arms and legs are connected to torso or trunk.
A digit is one of several most distal parts of a limb, such as fingers or toes, present in many vertebrates.
A leg is a weight-bearing and locomotive anatomical structure, usually having a columnar shape. During locomotion, legs function as "extensible struts". The combination of movements at all joints can be modeled as a single, linear element capable of changing length and rotating about an omnidirectional "hip" joint.
Polydactyly in stem-tetrapods should here be understood as having more than five digits to the finger or foot, a condition that was the natural state of affairs in the very first stegocephalians during the evolution of terrestriality. The polydactyly in these largely aquatic animals is not to be confused with polydactyly in the medical sense, i.e. it was not an anomaly in the sense it was not a congenital condition of having more than the typical number of digits for a given taxon. Rather, it appears to be a result of the early evolution from a limb with a fin rather than digits.
Comparative foot morphology involves comparing the form of distal limb structures of a variety of terrestrial vertebrates. Understanding the role that the foot plays for each type of organism must take account of the differences in body type, foot shape, arrangement of structures, loading conditions and other variables. However, similarities also exist among the feet of many different terrestrial vertebrates. The paw of the dog, the hoof of the horse, the manus (foot) and pes (foot) of the elephant, and the foot of the human all share some common features of structure, organization and function. Their foot structures function as the load-transmission platform which is essential to balance, standing and types of locomotion.
The limbs of the horse are structures made of dozens of bones, joints, muscles, tendons, and ligaments that support the weight of the equine body. They include two apparatuses: the suspensory apparatus, which carries much of the weight, prevents overextension of the joint and absorbs shock, and the stay apparatus, which locks major joints in the limbs, allowing horses to remain standing while relaxed or asleep. The limbs play a major part in the movement of the horse, with the legs performing the functions of absorbing impact, bearing weight, and providing thrust. In general, the majority of the weight is borne by the front legs, while the rear legs provide propulsion. The hooves are also important structures, providing support, traction and shock absorption, and containing structures that provide blood flow through the lower leg. As the horse developed as a cursorial animal, with a primary defense mechanism of running over hard ground, its legs evolved to the long, sturdy, light-weight, one-toed form seen today.
This glossary explains technical terms commonly employed in the description of dinosaur body fossils. Besides dinosaur-specific terms, it covers terms with wider usage, when these are of central importance in the study of dinosaurs or when their discussion in the context of dinosaurs is beneficial. The glossary does not cover ichnological and bone histological terms, nor does it cover measurements.
Innovations conventionally associated with terrestrially first appeared in aquatic elpistostegalians such as Panderichthys rhombolepis, Elpistostege watsoni, and Tiktaalik roseae. Phylogenetic analyses distribute the features that developed along the tetrapod stem and display a stepwise process of character acquisition, rather than abrupt. The complete transition occurred over a period of 25 million years beginning with the tetrapodomorph diversification in the Middle Devonian.
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