Heterostraci is an extinct subclass of pteraspidomorph, ostracoderm, jawless vertebrate that lived primarily in marine and estuary environments. Heterostraci existed from the mid-Ordovician to the conclusion of the Devonian.
Psammosteidae is an extinct family of flattened, benthic heterostracan vertebrates that lived in marine and estuary environments in Europe, Russia & North America. They arose during the Early Devonian, with the first known genus, Drepanaspis from the Hunsrück lagerstätte. The Psammosteids were the only heterostracans that survived the Upper Frasnian extinction event during the Late Devonian, dying out in the extinction event at the very end of the Devonian.
Protaspididae is an extinct family of pteraspidid heterostracan agnathans. Fossils of the various genera are found in early Devonian-aged marine strata. Protaspidids were once thought to represent a transitional form between the Pteraspididae and the Psammosteida, bearing the broad head shield shape of the latter, due to a more benthic (bottom-dwelling) existence, but recent phylogenical comparisons demonstrate that the protaspidids are actually highly derived pteraspidids, and that the anchipteraspidids, the most primitive of pteraspidids, are the sister-group of the psammosteids.
Lepidaspis serrata is an extinct heterostracan jawless fish from Early Devonian Canada. Its scientific name refers to the fact that the armor is composed of hundreds of tiny scales with serrated edges.
Drepanaspis is an extinct genus of heterostracan armoured jawless fish from the Early Devonian. Drepanaspis are assumed to have lived primarily in marine environments and is most commonly characterized by their ray-like, heavily armoured bodies, along with their lack of paired fins and jaws.
Traquairaspis is a genus of extinct heterostracan agnathan fish known from the Silurian and Early Devonian periods. It is predominantly known from Late Silurian fluvial deposits from Wales and England: some species were also found in shallow water marine environment in Canada and North America.
Palanasaspis chekhivensis is an extinct species of pteraspidid heterostracan agnathan which existed during the Pragian epoch of the early Devonian period in what is now Podolia, Ukraine. It is known primarily from a wide rostral plate, which is referenced in the generic name, a compound word combining the Latin words pala, "shovel," and nasus, "nose," with the Greek suffix aspis, "a small shield." Although the rostral plate clearly marks the creature as a pteraspidoid heterostracan, that literally nothing else of its anatomy is known forces researchers to leave it as incertae sedis.
Lechriaspis is an extinct genus of poraspid heterostracan agnathan known from the Early Devonian of northern Utah, western United States.
Poraspis is an extinct genus of heterostracan. Fossils are found in Late Silurian and Early Devonian marine strata of Norway, Canada and the United States.
Panamintaspis snowi is an extinct species of pteraspidid heterostracan agnathan which existed during the early Middle Devonian period of Death Valley, California. Fossils are found in Late Emsian-aged marine strata of the Lost Burro Formation. P. snowi strongly resembles Pteraspis, though while it was originally described as a member of the same family, Pteraspididae, a recent phylogenetic reassessment of the order Pteraspidiformes places P. snowi within the paraphyletic family "Protopeteraspidae," as the sister taxon of the suborder Pteraspidoidei.
Pteraspidiformes is an extinct order of heterostracan agnathan vertebrates known from extensive fossil remains primarily from Early Devonian strata of Europe and North America, and from Upper Silurian Canada.
Cyathaspidiformes is an extinct order of heterostracan vertebrates known from extensive fossil remains primarily from Silurian to Early Devonian strata of Europe, and North America, and from Early Devonian marine strata of Siberia.
Edaphaspis bystrowi is an extinct amphiaspidid cyathaspidid heterostracan. Its fossils are restricted to early Devonian-aged marine strata of the Taimyr Peninsula, Early Devonian Siberia. E. bystrowi, as with all other amphiaspidids, is thought to have been a benthic filter feeder that lived on top of, or buried just below the surface of the substrate of hypersaline lagoon-bottoms. It is the only representative of the family Edaphaspididae.
Weigeltaspis is a genus of extinct heterostracan agnathan fish known from the Late Silurian and Early Devonian periods. Fossils are known primarily from Early Devonian-aged marine strata of Europe and Canada. Fragments and disarticulated plates of what may be of Weigeltaspis are known from Late Silurian-aged marine strata of Arctic Canada. Rare articulated fossils, plus the overall anatomy of its plates suggest the living animals were, at least superficially, similar to psammosteids, some authorities, such as Tarlo, place them within Psammosteida. Because the ornamentation on the plates and scales are very similar to the ornamentation seen on the plates and scales of Traquairaspis to the point of constant confusion, other authorities follow the lead of Obruchev, and place Weigeltaspis within Traquairaspidiformes. Still other authorities remain unconvinced of either side, and, instead, follow Denison's lead to simply treat Weigeltaspis as incertae sedis.
Cyathaspidida is a taxon of extinct cyathaspidiform heterostracan agnathans whose fossils are found in Silurian to Lower Devonian marine strata of Europe and North America. In life, they are thought to be benthic animals that lived most of their lives either mostly buried in or resting directly on top of the substrate.
Anglaspis is an extinct genus of cyathaspidiform heterostracan agnathan. Fossils are found in marine strata of Europe, from the late Silurian period until the genus' extinction during the Early Devonian. As with other cyathaspidiforms, individuals of Anglaspis had dorsal and ventral plates covering the forebody, gill pouches, and nasal openings that lay on the roof of the oral cavity.
Paraliliaspis egregia is an extinct cyathaspidiform heterostracan agnathan from early Devonian marine strata of the Ural Mountains.
Protopteraspididae is an extinct family of pteraspidid heterostracan agnathans. Fossils of the various genera are found in early Devonian-aged marine strata. Protopteraspidids were once thought to represent a taxon of basal pteraspidids but recent evaluations demonstrate that Protopteraspididae is a paraphyletic group of various transitional forms representing a gradual transition between the more advanced Pteraspoidei, and the anchipteraspidids and the Psammosteids.
Pteraspididae is an extinct family of heterostracan vertebrates. It is an Early Devonian-aged family that contains the stereotypical, torpedo-shaped pteraspidids, such as the type genus, Pteraspis, and its close relatives Errivaspis and Rhinopteraspis, that are popularly thought to have actively swam about in the water column. The elongated rostral plate is thought to help with their hydrodynamic ability. Various genera are found in various marine and estuarine strata of Early Devonian Europe and North America.
Tartuosteus is an extinct genus of jawless fish from the Devonian of Eastern Europe. It is notable for its large size in comparison to other Ostracoderms, reaching a width of 1.5 m (4.9 ft).
